2012 in fish paleontology

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This list of fossil fish species is a list of taxa of fish that have been described during the year 2012. The list only includes taxa at the level of genus or species.

Newly named jawless vertebrates[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Diademaspis janvieri[1]

Sp. nov

Valid

Keating, Sansom & Purnell

Early Devonian (Lochkovian)

Wayne Herbert Quarry

 United Kingdom

A member of Osteostraci, a species of Diademapsis.

Dunyu[2]

Gen. et sp. et comb. nov

Valid

Zhu, Liu, Jia & Gai

Silurian

Huixingshao Formation
Kuanti Formation

 China

An eugaleaspidiform galeaspid. Its type species is Dunyu longiforus; genus also contains "Eugaleaspis" xiushanensis Liu 1983.

Janaspis[1]

Gen. et comb et 2 sp. nov

Valid

Keating, Sansom & Purnell

Early Devonian (Lochkovian)

Wayne Herbert Quarry

 United Kingdom

A new genus of Osteostraci including the type species "Cephalaspis" pagei Lankester, 1868[3] and new species J. newtonensis and J. punctata.

Kerreralepis[4]

Gen. et sp. nov

Valid

Blom

Lower Devonian

 United Kingdom

A birkeniid anaspid. The type species is Kerreralepis carinata.

Waengsjoeaspis platycornis[5]

Sp. nov

Valid

Scott & Wilson

Early Devonian (Lochkovian)

Man On The Hill Lagerstätte

 Canada

A member of Osteostraci, a species of Waengsjoeaspis.

Zenaspis waynensis[1]

Sp. nov

Valid

Keating, Sansom & Purnell

Early Devonian (Lochkovian)

Wayne Herbert Quarry

 United Kingdom

A member of Osteostraci, a species of Zenaspis.

Newly named acanthodians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Halimacanthodes[6]

Gen. et sp. nov

Valid

Burrow, Trinajstic & Long

Late Devonian (Frasnian)

Gogo Formation

 Australia

An acanthodiform acanthodian. The type species is Halimacanthodes ahlbergi.

Machaeracanthus goujeti[7]

Sp. nov

Valid

Botella, Martínez-Pérez & Soler-Gijón

Early Devonian

 France
 Spain

A species of Machaeracanthus.

Podoliacanthus[8]

Gen. et sp. nov

Valid

Voichyshyn & Szaniawski

Early Devonian, possibly also late Silurian.

 Ukraine
 Greenland

An ischnacanthiform, possibly an ischnacanthid. The type species is Podoliacanthus zychi; also contains three yet unnamed species.

Newly named cartilaginous fishes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Abdounia lata[9]

Sp. nov

Valid

Malyshkina

Late Eocene

Tavda Formation

 Russia

A requiem shark, a species of Abdounia.

Abdounia vassilyevae[9]

Sp. nov

Valid

Malyshkina

Late Eocene

Tavda Formation

 Russia

A requiem shark, a species of Abdounia.

Agaleorhynchus[10]

Gen. et sp. nov

Valid

Guinot et al.

Late Cretaceous (late Santonian to early Campanian)

 United Kingdom

A sclerorhynchid. The type species is Agaleorhynchus britannicus.

Argoubia[11]

Gen. et sp. nov

Valid

Adnet et al.

Late Eocene

 Morocco

A mobulid. The type species is Argoubia barbei.

Australopristis[12]

Gen. et sp. nov

Valid

Martill & Ibrahim

Late Cretaceous

 New Zealand

A sawfish. The type species is Australopristis wiffeni.

Borodinopristis shannoni[13]

Sp. nov

Valid

Case et al.

Late Cretaceous (Campanian)

 United States

A sclerorhynchid sawfish, a species of Borodinopristis.

Carcharodon hubbelli[14]

Sp. nov

Valid

Ehret et al.

Late Miocene

Pisco Formation

 Peru

A lamnid shark, a species of Carcharodon.

Cretaplatyrhinoidis[10]

Gen. et sp. nov

Valid

Guinot et al.

Late Cretaceous (Santonian to Campanian)

 France
 United Kingdom

A platyrhinid. The type species is Cretaplatyrhinoidis ornatus.

Cromhallia[15]

Gen. et sp. nov

Valid

Behan, Walken & Cuny

Early Carboniferous

 United Kingdom

A chondrichthyan of indeterminate phylogenetic relationships. The type species is Cromhallia parvunda.

Duffinselache[16]

Gen. et comb. nov

Valid

Andreev & Cuny

Rhaetian

 United Kingdom

A selachimorph, a new genus for "Polyacrodus" holwellensis (Duffin, 1998).

Eoplinthicus underwoodi[11]

Sp. nov

Valid

Adnet et al.

Late Eocene

 Morocco

A mobulid, a species of Eoplinthicus.

Galeocorax[17]

Nom. nov

Valid

Cappetta

Late Cretaceous

 United Kingdom
 United States

A member of Anacoracidae; a replacement name for Paracorax Cappetta (1977).

Isacrodus[18]

Gen. et sp. nov

Valid

Ivanov, Nestell & Nestell

Middle Permian (Roadian)

Cutoff Formation

 United States

A jalodontid chondrichthyan. The type species is Isacrodus marthae.

Kaibabvenator[19]

Gen. et sp. nov

Valid

Hodnett et al.

Permian (Late Leonardian)

Kaibab Formation

 United States

A ctenacanthiform. The type species is Kaibabvenator swiftae.

Nanocorax[17]

Nom. nov

Valid

Cappetta

Late Cretaceous

 France
 United Kingdom
 United States

A member of Anacoracidae; a replacement name for Microcorax Cappetta & Case (1975).

Nanoskalme[19]

Gen. et sp. nov

Valid

Hodnett et al.

Permian (Late Leonardian)

Kaibab Formation

 United States

A ctenacanthiform. The type species is Nanoskalme natans.

Neosaivodus[19]

Gen. et sp. nov

Valid

Hodnett et al.

Permian (Late Leonardian)

Kaibab Formation

 United States

A ctenacanthiform. The type species is Neosaivodus flagstaffensis.

Odontaspis watinensis[20]

Sp nov

Valid[21]

Cook et al.

Late Cretaceous (early Turonian)

Kaskapau Formation

 Canada

A species of Odontaspis.

Oromobula[11]

Gen. et sp. nov

Valid

Adnet et al.

Late Eocene

 Morocco

A mobulid. The type species is Oromobula dakhlaensis.

Parasquatina jarvisi[22]

Sp nov

Valid

Guinot et al.

Late Cretaceous

Western Europe

A squatiniform, a species of Parasquatina.

Parasquatina justinensis[22]

Sp nov

Valid

Guinot et al.

Late Cretaceous

Western Europe

A squatiniform, a species of Parasquatina.

Plesiobatis vandenboschi[23]

Sp. nov

Valid

Bor, Reinecke & Verschueren

Miocene (early Langhian)

Breda Formation

 Netherlands

A relative of the deepwater stingray.

Pseudoplatyrhina[10]

Gen. et sp. nov

Valid

Guinot et al.

Late Cretaceous (late Santonian to early Campanian)

 France
 United Kingdom

A platyrhinid. The type species is Pseudoplatyrhina crispa.

Ptychotrygonoides sabatieri[10]

Sp. nov

Valid

Guinot et al.

Late Cretaceous (middle Turonian)

 France

A ptychotrygonid, possibly a relative of sclerorhynchids; a species of Ptychotrygonoides.

"Rhinobatos" seruensis[10]

Sp. nov

Valid

Guinot et al.

Late Cretaceous (late Santonian)

 France

A guitarfish, a possible species of Rhinobatos.

Rhomaleodus[16]

Gen. et sp. nov

Valid

Andreev & Cuny

Anisian

 Bulgaria

A selachimorph. The type species is Rhomaleodus budurovi.

Sphenacanthus riorastoensis[24]

Sp. nov

Valid

Pauliv, Dias & Sedor

Permian (Wordian to Wuchiapingian)

Rio do Rasto Formation

 Brazil

A species of Sphenacanthus.

Squatirhina draytoni[10]

Sp. nov

Valid

Guinot et al.

Late Cretaceous (late Early Cenomanian to middle Turonian)

 France
 United Kingdom

A guitarfish, a species of Squatirhina.

Tantalepis[25]

Gen. et sp. nov

Valid

Sansom et al.

Middle Ordovician (Darriwilian)

Stairway Sandstone

 Australia

A gnathostome of uncertain phylogenetic placement, a possible cartilaginous fish. The type species is Tantalepis gatehousei.

Texasodus[18]

Gen. et sp. nov

Valid

Ivanov, Nestell & Nestell

Middle Permian (late Wordian or early Capitanian)

Bell Canyon Formation

 USA

A jalodontid chondrichthyan. The type species is Texasodus varidentatus.

Newly named bony fishes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aglyptorhynchus hakataramea[26]

sp nov

Valid

Gottfried, Fordyce & Rust

Late Oligocene

 New Zealand

A billfish

Apsopelix miyazakii[27]

Sp. nov

Valid

Yabumoto, Hikida & Nishino

Late Cretaceous (Turonian)

 Japan

A species of Apsopelix.

Aspidopleurus kickapoo[28]

Sp. nov

Valid

Alvarado-Ortega & Porras-Múzquiz

Late Cretaceous (Turonian)

Eagle Ford Formation

 Mexico

An aspidopleurid aulopiform, a species of Aspidopleurus.

Aspistor verumquadriscutis[29]

sp nov

Valid

Aguilera & Marceniuk

Late Miocene

Urumaco Formation

 Venezuela

An ariid, a species of Aspistor.

Bawitius[30]

Gen. et comb. nov

Valid

Grandstaff et al..

Cenomanian

Bahariya Formation

 Egypt

A polypterid actinopterygian, a new genus for "Polypterus" bartheli (Schaal, 1984).

Bruehnopteron[31]

Gen. et sp. nov

Valid

Schultze & Reed

Middle Devonian

 United States

A tristichopterid sarcopterygian. The type species is Bruehnopteron murphyi.

Caruso[32]

Gen. et comb. nov

Valid

Carnevale & Pietsch

Eocene

 Italy

A lophiid anglerfish, a new genus for "Lophius" brachysomus (Agassiz).

Diandongperleidus[33]

Gen. et sp. nov

Valid

Geng et al.

Middle Triassic (middle or late Anisian)

Guanling Formation

 China

A perleidid perleidiform. The type species is Diandongperleidus denticulatus.

Echinelops[34]

Gen. et sp. nov.

Valid

Murray & Hoşgör

Early Oligocene

 Turkey

An elopiform fish. The type species is Echinelops ozcani.

Eurolebias[35]

Gen. et comb. nov

Valid

Costa

Late Oligocene

 France

A pupfish; a new genus for "Prolebias" meridionalis Gaudant (1978).

Francolebias[35]

Gen. et comb. nov

Valid

Costa

Early Oligocene

 France

A relative of Valencia; a new genus for "Prolebias" delphinensis Gaudant (1989) and "Lebias" aymardi Sauvage (1869).

Frippia[36]

Gen. et sp. nov

Valid

Bannikov & Carnevale

Eocene

 Italy

A member of Percoidei. The type species is Frippia labroiformis.

Fuyuanperleidus[33]

Gen. et sp. nov

Valid

Geng et al.

Middle Triassic (late Ladinian)

Falang Formation

 China

A perleidid perleidiform. The type species is Fuyuanperleidus dengi.

Galaxias angustiventris[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A galaxiid, a species of Galaxias.

Galaxias brevicauda[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A galaxiid, a species of Galaxias.

Galaxias bobmcdowalli[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A galaxiid, a species of Galaxias.

Galaxias papilionis[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A galaxiid, a species of Galaxias.

Galaxias parvirostris[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A galaxiid, a species of Galaxias.

Galaxias tabidus[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

 New Zealand

A galaxiid, a species of Galaxias.

Gerpegezhus[38]

Gen. et sp. nov

Valid

Bannikov & Carnevale

Early Eocene

 Russia

A close relative of centriscids. The type species is Gerpegezhus paviai.

Hippotropiscis[39]

Gen. et sp. nov

Valid

Žalohar & Hitij

Middle Miocene

 Slovenia

A hippocampine syngnathid. The type species is Hippotropiscis frenki.

Illusionella[40]

Gen. et 2 sp. nov

Valid

Baykina

Miocene

 Russia

A clupeid fish. Genus contains two species: Illusionella tsurevica and Illusionella pshekhensis. The genus Illusionella was subsequently synonymized with the genus Sarmatella Menner (1949) by Baykina (2013); however, Baykina kept the species I. tsurevica and I. pshekhensis as valid, distinct species within the genus Sarmatella.[41]

Isurichthys breviusculus[42]

Sp nov

Valid

Bannikov

Lower Oligocene

 Russia

An ariommatid, a species of Isurichthys.

Kyphosichthys[43]

Gen. et sp. nov

Valid

Xu & Wu

Middle Anisian

Guanling Formation

 China

A ginglymodian actinopterygian.

Luopingperleidus[33]

Gen. et sp. nov

Valid

Geng et al.

Middle Triassic (middle or late Anisian)

Guanling Formation

 China

A perleidid perleidiform. The type species is Luopingperleidus sui.

Luoxiongichthys[44]

Gen. et sp. nov

Valid

Wen et al.

Middle Anisian

Guanling Formation

 China

A basal actinopterygian.

Macrosemimimus[45]

Gen. et sp. et comb. nov

Valid

Schröder, López-Arbarello & Ebert

Late Jurassic

 France
 Germany
 United Kingdom

A semionotiform. The type species is Macrosemimimus fegerti; "Lepidotes" lennieri (Sauvage, 1893) is a second species of Macrosemimimus.

Mataichthys[37]

Gen. et sp. nov

Valid

Schwarzhans, Scofield, Tennyson & T. Worthy in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

An eleotrid. Contains M. bictenatus, M. procerus, M. rhinoceros, and M. taurinus

Nanaichthys[46]

Gen. et sp. nov

Valid

Amaral & Brito

Early Cretaceous, possibly Aptian

Marizal Formation

 Brazil

A member of Chanidae. The type species is Nanaichthys longipinnus.

Palatinichthys[47]

Gen. et sp. nov

Valid

Witzmann & Schoch

Lower Permian

Saar–Nahe Basin

 Germany

A megalichthyid sarcopterygian. The type species is Palatinichthys laticeps.

Paleotetra[48]

Gen. et 2 sp.

Valid

Weiss, Malabarba & Malabarba

Eocene to Oligocene

 Brazil

A characiform, probably a characid. The genus contains two species: Paleotetra entrecorregos and Paleotetra aiuruoca.

Paranaichthys[49]

Gen. et sp. nov

Valid

Dias

Permian, Guadalupian (Wordian or Capitanian)

Rio do Rasto Formation

 Brazil

A probable relative of Platysomus and other platysomids. The type species is Paranaichthys longianalis.

Parawenzichthys[50]

Gen. et sp. nov

Valid

de Figueiredo, Gallo & Delarmelina

Turonian

Atlântida Formation

 Brazil

A protacanthopterygian.

Prototroctes modestus[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A retropinnid, a species of Prototroctes.

Prototroctes vertex[37]

Sp nov

Valid

Schwarzhans in Schwarzhans et al..

Early Miocene

Bannockburn Formation

 New Zealand

A retropinnid, a species of Prototroctes.

Rebellatrix[51]

Gen. et sp. nov

Valid

Wendruff & Wilson

Lower Triassic

Sulphur Mountain Formation

 Canada

A coelacanth. The type species is Rebellatrix divaricerca.

Reidus[52]

Gen. et sp. nov

Valid

Graf

Albian

 United States

A coelacanth. The type species is Reidus hilli.

Rhinodipterus kimberleyensis[53]

Sp nov

Valid

Clement

Late Devonian

Gogo Formation

 Australia

A lungfish, a species of Rhinodipterus.

Sangiorgioichthys valmarensis[54]

Sp. nov

Valid

Lombardo, Tintori & Tona

Middle Triassic (late Ladinian)

  Switzerland

A semionotid semionotiform, a species of Sangiorgioichthys.

Sinamia liaoningensis[55]

sp nov

Valid

Zhang

Early Cretaceous (Barremian to Aptian)

Jiufotang Formation
Yixian Formation

 China

A sinamiid amiiform, a species of Sinamia.

Thoracopterus wushaensis[56]

Sp. nov

Valid

Tintori et al.

Middle Triassic (late Ladinian)

Falang Formation

 China

A thoracopterid peltopleuriform, a species of Thoracopterus.

Tinirau[57]

Gen. et sp. nov

Valid

Swartz

Middle Devonian

Red Hill I beds

 United States

An eotetrapodiform tetrapodomorph related to the clade Elpistostegalia[57] or a tristichopterid.[58] The type species is Tinirau clackae.

Tomognathus gigeri[59]

sp nov

Valid

Cavin & Giner

Valanginian

 France

A halecomorph fish, a species of Tomognathus.

Tungsenia[60]

Gen. et sp. nov

Valid

Lu et al.

Early Devonian

 China

A basal member of Tetrapodomorpha. The type species is Tungsenia paradoxa.

Voelklichthys[61]

Gen. et sp. nov

Valid

Arratia & Schultze

Kimmeridgian

 Germany

A macrosemiiform. The type species is Voelklichthys comitatus.

Zoqueichthys[62]

Gen. et sp. nov

Valid

Alvarado-Ortega & Than-Marchese

Late Cretaceous (Cenomanian)

Sierra Madre Formation

 Mexico

A close relative of Aipichthyoides and Aspesaipichthys, more distantly related to Aipichthys. The type species is Zoqueichthys carolinae.

References[edit]

  1. ^ a b c Joseph N. Keating, Robert S. Sansom and Mark A. Purnell (2012). "A new osteostracan fauna from the Devonian of the Welsh Borderlands and observations on the taxonomy and growth of Osteostraci". Journal of Vertebrate Paleontology 32 (5): 1002–1017. doi:10.1080/02724634.2012.693555. 
  2. ^ Zhu Min, Liu Yu-Hai, Jia Lian-Tao and Gai Zhi-Kun (2012). "A new genus of eugaleaspidiforms (Agnatha: Galeaspida) from the Ludlow, Silurian of Qujing, Yunnan, Southwestern China". Vertebrata PalAsiatica 50 (1): 1–7. 
  3. ^ Lankester E. R. (1868). "Monograph of fishes of the Old Red Sandstone of Britain. Part I. The Cephalaspidae". The Palaeontographical Society,London.: 62pp. 
  4. ^ Henning Blom (2012). "New birkeniid anaspid from the Lower Devonian of Scotland and its phylogenetic implications". Palaeontology 55 (3): 641–652. doi:10.1111/j.1475-4983.2012.01142.x. 
  5. ^ Bradley R. Scott and Mark V. H. Wilson (2012). "A new species of Waengsjoeaspis (Cephalaspidomorpha, Osteostraci) from the Early Devonian of northwestern Canada, with a redescription of W. nahanniensis and implications for growth, variation, morphology, and phylogeny". Journal of Vertebrate Paleontology 32 (6): 1235–1253. doi:10.1080/02724634.2012.694514. 
  6. ^ Carole J. Burrow, Kate Trinajstic and John Long (2012). "First acanthodian from the Upper Devonian (Frasnian) Gogo Formation, Western Australia". Historical Biology 24 (4): 349–357. doi:10.1080/08912963.2012.660150. 
  7. ^ Héctor Botella, Carlos Martínez-Pérez and Rodrigo Soler-Gijón (2012). "Machaeracanthus goujeti n. sp. (Acanthodii) from the Lower Devonian of Spain and northwest France, with special reference to spine histology". Geodiversitas 34 (4): 761–783. doi:10.5252/g2012n4a3. 
  8. ^ Victor Voichyshyn and Hubert Szaniawski (2012). "Acanthodian jaw bones from Lower Devonian marine deposits of Podolia, Ukraine". Acta Palaeontologica Polonica 57 (4): 879–896. doi:10.4202/app.2011.0079. 
  9. ^ a b T. P. Malyshkina (2012). "New sharks of the genus Abdounia (Carcharhiniformes: Carcharhinidae) from the Upper Eocene of the Trans-Ural Region". Paleontological Journal 46 (4): 392–399. doi:10.1134/S0031030112040053. 
  10. ^ a b c d e f Guillaume Guinot, Henri Cappetta, Charlie J. Underwood and David J. Ward (2012). "Batoids (Elasmobranchii: Batomorphii) from the British and French Late Cretaceous". Journal of Systematic Palaeontology 10 (3): 445–474. doi:10.1080/14772019.2011.588255. 
  11. ^ a b c Sylvain Adnet, Henri Cappetta, Guillaume Guinot and Giuseppe Notarbartolo di Sciara (2012). "Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference". Zoological Journal of the Linnean Society 166 (1): 132–159. doi:10.1111/j.1096-3642.2012.00844.x. 
  12. ^ David M. Martill and Nizar Ibrahim (2012). "Aberrant rostral teeth of the sawfish Onchopristis numidus from the Kem Kem beds (?early Late Cretaceous) of Morocco and a reappraisal of Onchopristis in New Zealand". Journal of African Earth Sciences 64: 71–76. doi:10.1016/j.jafrearsci.2011.11.009. 
  13. ^ Gerard R. Case, Todd D. Cook, Mark V.H. Wilson and Paul D. Borodin (2012). "A new species of the sclerorhynchid sawfish Borodinopristis from the Campanian (Upper Cretaceous) of North Carolina, USA". Historical Biology: An International Journal of Paleobiology 24 (6): 592–597. doi:10.1080/08912963.2012.663367. 
  14. ^ Dana J. Ehret, Bruce J. MacFadden, Douglas S. Jones, Thomas J. Devries, David A. Foster and Rodolfo Salas-Gismondi (2012). "Origin of the white shark Carcharodon (Lamniformes: Lamnidae) based on recalibration of the Upper Neogene Pisco Formation of Peru". Palaeontology 55 (6): 1139–1153. doi:10.1111/j.1475-4983.2012.01201.x. 
  15. ^ Claire Behan, Gordon Walken and Gilles Cuny (2012). "A Carboniferous chondrichthyan assemblage from residues within a Triassic karst system at Cromhall quarry, Gloucestershire, England". Palaeontology 55 (6): 1245–1263. doi:10.1111/j.1475-4983.2012.01193.x. 
  16. ^ a b Plamen S. Andreev and Gilles Cuny (2012). "New Triassic stem selachimorphs (Chondrichthyes, Elasmobranchii) and their bearing on the evolution of dental enameloid in Neoselachii". Journal of Vertebrate Paleontology 32 (2): 255–266. doi:10.1080/02724634.2012.644646. 
  17. ^ a b H. Cappetta (2012). Handbook of Paleoichthyology Volume 3E. Chondrichthyes. Mesozoic and Cenozoic Elasmobranchii: Teeth. Verlag Dr. Friedrich Pfeil. pp. 512pp. ISBN 978-3-89937-148-2. 
  18. ^ a b Alexander Ivanov, Merlynd Nestell and Galina Nestell (2012). "New jalodontid chondrichthyans from the Middle Permian of West Texas, USA". Historical Biology 24 (4): 359–368. doi:10.1080/08912963.2012.662229. 
  19. ^ a b c John-Paul M. Hodnett, David K. Elliott, Tom J. Olson and James H. Wittke (2012). "Ctenacanthiform sharks from the Permian Kaibab Formation, northern Arizona". Historical Biology 24 (4): 381–395. doi:10.1080/08912963.2012.683193. 
  20. ^ Todd D. Cook, Mark V. H. Wilson, Alison M. Murray, A. Guy Plint, Michael G. Newbrey and Michael J. Everhart (2013). "A high latitude euselachian assemblage from the early Turonian of Alberta, Canada". Journal of Systematic Palaeontology 11 (5): 555–587. doi:10.1080/14772019.2012.707990. 
  21. ^ http://zoobank.org/NomenclaturalActs/0B658442-06A8-4FB0-8E0E-DFA614142285
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