2011 in paleontology

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Paleontology, palaeontology or palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred in the year 2011.

Lobopods[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Diania cactiformis[2]

Valid

Liu et al.

Lower Cambrian

Maotianshan shale

 China

A lobopodian.

Mureropodia apae

Valid

Gamez Vintaned, Linan, & Zhuravlev

Early Cambrian

Murero

 Spain

A lobopodian.

Siberion lenaicus

Valid

Dzik

Early Cambrian

Sinsk Formation

 Russia

A lobopodian.

Vetulicolians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Ooedigera peeli[3]

gen. nov

valid

Zinther, et al.

Early Cambrian

Sirius Passet

 Greenland

earliest known vetulicolian described

Artist's reconstruction

Molluscs[edit]

Arthropods[edit]

Arachnids[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Nephila jurassica[4][5]

sp nov

Valid

Selden, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

largest fossil spider described. Originally assigned to the genus Nephila; subsequently made the type species of a separate genus Mongolarachne.[5]

Insects[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Agulla protomaculata[6]

sp nov

valid

Engel

Lutetian

Green River Formation

 USA

Only snakefly from the Green River Formation

Anonychomyrma geinitzi[7]

Comb nov

valid

Mayr

Lutetian

Baltic amber

 Europe

Fossil Dolichoderin ant

Arostropsis[8]

Gen et sp nov

valid

Yunakov & Kirejtshuk

Eocene

Baltic amber

 Europe

Monotypic broad-nosed weevil genus, one species A. groehni

Astreptolabis[9]

Subfam, Gen et sp nov

valid

Engel

Albian

Burmese amber

 Myanmar

Monotypic earwig subfamily, one species A. ethirosomatia

Baltimartyria rasnitsyni[10]

sp nov

valid

Mey

Lutetian

Baltic amber

 Europe

the second species in Baltimartyria

Deinodryinus velteni[11]

sp nov

Valid

Guglielmino & Olmi

Lutetian

Baltic amber

 Europe

A dryinid wasp.

Diochus electrus[12]

sp nov

valid

Chatzimanolis & Engel

Lutetian

Baltic amber

 Europe

first valid Staphylininae species from the fossil record

Dryinus rasnitsyni[13]

sp nov

valid

Olmi & Guglielmino

Burdigalian

Dominican amber

 Dominican Republic

second fossil Dryinus lamellatus group species

Iberoraphidia[14]

sp nov

valid

Jepsen, Ansorge, & Jarzembowski

Lower Barremian

Serra del Montsec

 Spain

A mesoraphidiid snakefly, one species Iberoraphidia dividua

Lebanoraphidia[15]

sp nov

valid

Bechly & Wolf-Schwenninger

Upper Neocomian

Jezzine

 Lebanon

A mesoraphidiid snakefly, one species Lebanoraphidia nana

Leptolingia calonervis[16]

sp.

Valid

Shi, Liu, & Ren

Middle Jurassic

Jiulongshan Formation

 China

Leptolingia imminuta[17]

sp.

Valid

Liu, Shi, & Ren

Middle Jurassic

Daohugou Beds

 China

the smallest species known in the family Grammolingiidae

Litholingia ptesa[16]

sp.

Valid

Shi, Yang, & Ren

Middle Jurassic

Jiulongshan Formation

 China

Mickoleitia longimanus[18]

ord., fam., gen. et sp.

valid

Staniczek, Bechly, & Godunko

Lower Cretaceous

Crato Formation

 Brazil

A new genus in the new family Mickoleitiidae and a new insect order Coxoplectoptera.

Myanmymar[19]

gen et sp nov

valid

Huber

Albian

Burmese amber

 Myanmar

oldest mymarid described

Nanotermes[20]

Gen et sp. nov

vaild

Engel & Grimaldi

Ypresian

Cambay amber

 India

A Termitid termite

Ordralfabetix[21]

gen et sp.

valid

Szwedo

Ypresian

Oise amber

 France

Second fossil lophopid planthopper from Europe

Parastylotermes krishnai[20]

Sp. nov

vaild

Engel & Grimaldi

Ypresian

Cambay amber

 India

A Stylotermitid termite

Platyperla marquati[22]

Species

Valid

Gallego et al

Late Triassic

Potrerillos Formation

 Argentina

A platyperlid stonefly, a species of Platyperla.

Plumalexius[23]

fam, gen et sp.

vaild

Brothers

Turonian

New Jersey amber

 USA

sister genus to Plumariidae

Pristomyrmex rasnitsyni[24]

sp nov

valid

Dlussky & Radchenko

Late Eocene

Scandinavian amber

 Denmark

A fossil ant

Prostylotermes[20]

Gen et sp. nov

vaild

Engel & Grimaldi

Ypresian

Cambay amber

 India

A Stylotermitid termite

Renphasma[25]

Gen. et sp.

Valid

Nel & Delfosse

Early Cretaceous

Yixian Formation

 China

A stick insect

Termitaradus dominicanus[19]

Sp nov

vaild

Poinar

Burdigalian?

Dominican amber

 Dominican Republic

A termitaphidid termite bug

Titanomyrma lubei[26]

Gen. et sp.

valid

Archibald, Johnson, Mathewes, & Greenwood

Early Eocene

Green River Formation USA
Messel shales, Germany
Eckfeld Maar, Germany

 USA
 Germany

new genus for the largest known ants

Tytthodiplatys[9]

Gen et sp. nov

Valid

Engel

Albian

Burmese amber

 Myanmar

Oldest member of the family Diplatyidae

Undulopsychopsis[27]

Gen et sp. nov

Valid

Peng, Makarkin, Wang, & Ren

Lower Barremian to Upper Aptian

Yixian Formation

 China

A psychopsid silky lacewing, only species U. alexi

Zophotermes[20]

Gen et sp. nov

vaild

Engel & Singh

Ypresian

Cambay amber

 India

A Rhinotermitid termite

Fishes[edit]

Newly named jawless vertebrates[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Althaspis tarloi[28]

sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Djurinaspis secunda[28]

sp.

Valid

  • Voichyshyn

Early Devonian

Tyver Formation

A heterostracan

Lechriaspis[29]

Gen. et sp.

Valid

  • Elliott
  • Petriello

Early Devonian

Water Canyon Formation

A heterostracan

Palanasaspis[28]

Gen. et sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Podolaspis danieli[28]

sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Poraspis thomasi[29]

Species

Valid

  • Elliott
  • Petriello

Early Devonian

Lost Burro Formation

A heterostracan

Semipodolaspis[28]

Gen. et sp.

Valid

  • Voichyshyn

Early Devonian

Tyver Formation

A heterostracan

Shuyu[30]

Gen. et comb.

Valid

Late Telychian/Early Wenlock

Maoshan Formation

The basalmost known eugaleaspidiform galeaspid.

Zenaspis kasymyri[28]

sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A osteostracan

Newly named acanthodian[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Nerepisacanthus[31]

Gen. et sp.

Valid

  • Burrow

?Telychian/Ludlow

Cunningham Creek Formation

An acritolepid ischnacanthiform.

Newly named placoderms[edit]

Name Status Authors Age Unit Location Notes Images

Draconichthys elegans[32]

Valid

  • Rücklin

Frasnian

Anti-Atlas Mountains

 Morocco

A piscivorous selenosteid arthrodire

Newly named cartilaginous fishes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Cederstroemia ziaensis[33]

Species

Valid

  • Bourdon et al.

Late Cretaceous (Santonian)

Point Lookout Sandstone

A carpet shark, a species of Cederstroemia.

Dwardius sudindicus[34]

Species

Valid

Early Cenomanian

Karai Formation

An otodontid.

Eorhinoptera[35]

Gen. et sp. nov

Valid

  • Case, Cook & Wilson

Early Eocene (Ypresian)

Fishburne Formation

A stingray related to the genus Rhinoptera. The type species is Eorhinoptera grabdai.

Ertychius[36]

Gen. et sp. nov

Valid

  • Ginter, Hairapetian & Grigoryan

Devonian (early Famennian)

A ctenacanthiform. The type species is Ertychius intermedius.

Fayolia sharovi[37]

Species

Valid

Middle/Late Triassic

Madygen Formation

Egg capsules of a probable xenacanthid.

Gladioserratus[34]

Gen. et comb. et sp. nov

Valid

  • Underwood
  • Goswami
  • et al.

Cretaceous (Hauterivian) to Paleocene (Danian) or possibly even early Eocene[38]

A cow shark, a new genus for "Notidanus" aptiensis (Pictet, 1865). Genus also contains a new species Gladioserratus magnus; it might also contain Eocene species "Notorynchus" serratissimus Agassiz (1843).[38]

Karksiodus[39]

Gen. et sp. nov

Valid

  • Ivanov, Märss & Kleesment

Devonian (Givetian)

A member of Elasmobranchii of uncertain phylogenetic placement. The type species is Karksiodus mirus.

Kawichthys[40]

Gen. et sp.

Valid

Late Virgilian

Douglas Group

A symmoriiform.

Lissodus lusavorichi[36]

Sp. nov

Valid

  • Ginter, Hairapetian & Grigoryan

Devonian (late Famennian)

A lonchidiid hybodontiform, a species of Lissodus.

Lonchidion ferganensis[37]

Species

Valid

  • Fischer
  • Voigt
  • et al.

Middle/Late Triassic

Madygen Formation

A hybodontid shark.

Moerigaleus[41]

Gen. et sp. nov

Valid

  • Underwood & Ward

Late Eocene

A hemigaleid. The type species is Moerigaleus vitreodon.

Palaeoxyris alterna[37]

Species

Valid

  • Fischer
  • Voigt
  • et al.

Middle/Late Triassic

Madygen Formation

A hybodontid shark.

Ptychodus atcoensis[42]

Species

Valid

  • Hamm
  • Cicimurri

Late Cretaceous (early Coniacian)

Atco Formation

A species of Ptychodus.

Pueblocarcharias[33]

Gen. et sp. nov

Valid

  • Bourdon et al.

Late Cretaceous (Santonian)

Point Lookout Sandstone

A sand shark. The type species is Pueblocarcharias kawaikensis.

Reginaselache[43]

Gen. et sp.

Valid

  • Turner
  • Burrow

Middle Viséan

Ducabrook Formation

A xenacanthiform.

Scapanorhynchus puercoensis[33]

Sp. nov

Valid

  • Bourdon et al.

Late Cretaceous (Santonian)

Point Lookout Sandstone

A mitsukurinid, a species of Scapanorhynchus.

Scindocorax[33]

Gen. et sp. nov

Valid

  • Bourdon et al.

Late Cretaceous (Santonian)

Point Lookout Sandstone

An anacoracid. The type species is Scindocorax novimexicanus.

Tassiliodus[44]

Gen. et sp.

Valid

  • Derycke
  • Goujet

Emsian, Devonian

An euselachian.

Texatrygon stouti[33]

Sp. nov

Valid

  • Bourdon et al.

Late Cretaceous (Santonian)

Point Lookout Sandstone

A relative of Ptychotrygon and sclerorhynchids, a species of Texatrygon.

Newly named bony fishes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Araripichthys weberi[45]

Species

Valid

  • Alvarado-Ortega
  • Brito

Albian

Tlayúa Formation

An elopocephalan, a species of Araripichthys.

Astroscopus countermani[46]

Species

Valid

  • Carnevale, Godfrey & Pietsch

Miocene (Tortonian)

St. Marys Formation

A stargazer, a species of Astroscopus.

Beishanichthys[47]

Gen. et sp.

Valid

  • Xu
  • Gao

Early Triassic

A scanilepiform.

Bourbonnella jocelynae[48]

Species

Valid

  • Mickle

Carboniferous

Manning Canyon Shale Formation

An aeduellid.

Caucasichthys[49]

Gen. et sp.

Valid

  • Bannikov
  • Carnevale
  • Parin

Bartonian

A perciform.

Dapedium stollorum[50]

Species

Valid

  • Thies
  • Hauff

Early Toarcian

A species of Dapedium.

Guntherichthys[48]

Gen. et sp.

Valid

  • Mickle

Carboniferous

Manning Canyon Shale Formation

A ’palaeoniscoid’.

Gwawinapterus

Gen. et sp.

Valid

  • Arbour
  • Currie

Campanian

Northumberland Formation

Initially thought to be an istiodactylid pterosaur,[51] but subsequently reinterpreted as an indeterminate saurodontid fish.[52]

Laccognathus embryi[53]

Species

Valid

Frasnian

Fram Formation

A porolepiform sarcopterygian.

Lepidotes pankowskii[54]

Species

Valid

  • Forey
  • López-Arbarello
  • MacLeod

Cenomanian

Kem Kem Beds

A species of Lepidotes.

Leufuichthys[55]

Gen. et sp.

Valid

  • Gallo
  • Calvo
  • Kellner

Turonian/Coniacian

Portezuelo Formation

A clupeomorph.

Mimipiscis[56]

Nom. nov.

Valid

  • Choo

Upper Devonian

Gogo Formation

A new generic name for Mimia Gardiner & Bartram (1977). The type species is Mimipiscis toombsi (Gardiner & Bartram 1977).

Mimipiscis bartrami[56]

Species

Valid

  • Choo

Upper Devonian

Gogo Formation

A second species of Mimipiscis.

Pietschellus[57]

Gen. et sp.

Valid

  • Bannikov
  • Carnevale

Ypresian

Monte Bolca

A percomorph.

Scheenstia[58]

Gen. et sp.

Valid

  • López-Arbarello
  • Sferco

Late Jurassic

A semionotiform.

Sinosaurichthys[59]

Gen. et 3 sp.

Valid

Anisian

Guanling Formation

A saurichthyid.

Sorbinichthys africanus[60]

Species

Valid

  • Murray
  • Wilson

Cenomanian or Turonian

Akrabou Formation

A species of Sorbinichthys.

Spinofacia[48]

Gen. et sp.

Valid

  • Mickle

Carboniferous

Manning Canyon Shale Formation

A ’palaeoniscoid’.

Amphibians[edit]

Newly named lepospondyls[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Kirktonecta[61]

Gen. et sp.

Valid

  • Clack

Viséan

East Kirkton Quarry

A microsaur.

Tambaroter[62]

Gen. et sp.

Valid

Early Permian

Tambach Formation

An ostodolepid.

Newly named temnospondyls[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Acheloma dunni[63]

Species

Valid

  • Polley
  • Reisz

Early Permian

A trematopid.

Calmasuchus[64]

Gen. et sp.

Valid

  • Fortuny
  • Galobart
  • De Santisteban

early-middle Anisian

Catalonian basin

A capitosaurian.

Rotaryus[65]

Gen. et sp.

Valid

Early Permian

Tambach Formation

A trematopid.

Rubeostratilia[66]

Gen. et sp.

Valid

  • Bourget
  • Anderson

Early Permian

An amphibamid.

Rubeostratilia texensis.

Newly named lissamphibians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Urupia[67]

Gen. et sp.

Valid

  • Skutschas
  • Krasnolutskii

Bathonian

Itat Formation

A basal stem salamander.

Anapsids[edit]

Newly named basal anapsids[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Gansurhinus[68]

Gen. et sp.

Valid

Middle Permian

A moradisaurine captorinid.

Newly named turtles[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Albertwoodemys[69]

Gen. et sp.

Valid

Early Oligocene

Jebel Qatrani Formation

A podocnemidid.

Brontochelys[69]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

A podocnemidid, a new genus for "Shweboemys" gaffneyi.

Cordichelys[69]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Eocene

Qasr el-Sagha beds

A podocnemidid, a new genus for "Podocnemis" antiqua.

Gilmoremys[70]

Gen. et comb.

Valid

  • Joyce
  • Lyson

Maastrichtian

Lance Formation
Hell Creek Formation

A softshell turtle, new genus for "Aspideretes" lancensis (Gilmore, 1928).

Graptemys kerneri[71]

Sp. nov

Valid

  • Ehret & Bourque

Late Pleistocene

An emydid, a species of Graptemys.

Hispaniachelys[72]

Gen. et sp.

Valid

Oxfordian

Lorente Formation

A paracryptodire or a basal testudine.

Hoplochelys clark[73]

Species

Valid

Maastrichtian

Hell Creek Formation

A kinosternoid, a species of Hoplochelys.

Lapparentemys[69]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Paleocene

Santa Lucía Formation

A podocnemidid, a new genus for "Roxochelys" vilavilensis.

Latentemys[69]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Uncertain, possibly Miocene.

Uncertain, possibly Moghara Formation.

A podocnemidid.

Lemurchelys[69]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

Moghara Formation

A podocnemidid.

Mogharemys[69]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Miocene

A podocnemidid, a new genus for "Sternothaerus" blanckenhorni.

Oliveremys[74]

Gen. et comb. nov.

Valid

  • Vitek

Eocene

Washakie Formation

A trionychid, a new genus for "Trionyx" uintaensis.

Patagoniaemys[75]

Gen. et sp.

Valid

  • Sterli
  • De la Fuente

Campanian - Maastrichtian

La Colonia Formation

A stem turtle.

Peiropemys[69]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

A podocnemidid.

Pricemys[69]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

A podocnemidid.

Prochelidella cerrobarcinae[76]

Species

Valid

Aptian/Albian

Cerro Barcino Formation

The oldest known chelid, a species of Prochelidella.

Selenemys[77]

Gen. et sp.

Valid

  • Pérez-García
  • Ortega

Upper Kimmeridgian

Lusitanian Basin

A pleurosternid.

Sokatra[78]

Gen. et sp.

Valid

  • Gaffney
  • Krause

Maastrichtian

Maevarano Formation

A pleurodiran.

Trionyx ikoviensis[79]

Species

Valid

Middle Eocene

A species of Trionyx sensu lato.

Wilburemys[80]

Gen. et sp. nov

Valid

  • Martin
  • Mallory

Late Miocene (Hemphillian)

A relative of emydids. The type species is Wilburemys yakimensis.

Diapsids[edit]

Newly named basal diapsids[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Orovenator[81]

Gen. et sp.

Valid

Sakmarian

The oldest and most basal neodiapsid.

Newly named ichthyosaurs[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Sveltonectes[82]

Gen. et sp.

Valid

Barremian

An ophthalmosaurid.

Lepidosauromorphs[edit]

Newly named saurosphargids[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Sinosaurosphargis[83]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

An incertae sedis.

Newly named sauropterygians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Abyssosaurus[84]

Gen. et sp.

Valid

  • Berezin

Late Hauterivian

An aristonectid.

Diandongosaurus[85]

Gen. et sp.

Valid

  • Shang
  • Wu
  • Li

Middle Anisian

Guanling Formation

A relative of nothosauroids.[86]

Dianopachysaurus[87]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

A keichousaurid pachypleurosaur.

Hauffiosaurus tomistomimus[88]

Species

Valid

Early Toarcian

Whitby Limestone Formation

A second species of Hauffiosaurus.

Marmornectes[89]

Gen. et sp.

Valid

  • Ketchum
  • Benson

Callovian

Oxford Clay Formation

A pliosaurid.

Westphaliasaurus[90]

Gen. et sp.

Valid

  • Schwermann
  • Sander

Pliensbachian

Höxter district

A plesiosaurid plesiosaur.

Zarafasaura[91]

Gen. et sp.

Valid

Maastrichtian

An elasmosaurid plesiosaur.

Newly named lizards[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Brasiliguana[92]

Gen. et sp.

Valid

  • Nava
  • Martinelli

Turonian/Santonian

Adamantina Formation

An iguanian lizard.

Cryptolacerta[93]

Gen. et sp.

Valid

Lutetian

Messel pit

An amphisbaenian lizard.

Latoplatecarpus[94]

Gen. et sp.

Valid

  • Konishi
  • Caldwell

Campanian

Pierre Shale Formation
Demopolis Chalk Formation

A plioplatecarpine mosasaur. Its type species is Latoplatecarpus willistoni; "Plioplatecarpus" nichollsae (Cuthbertson et al., 2007) is a second species of Latoplatecarpus.

Plesioplatecarpus[94]

Gen. et comb.

Valid

  • Konishi
  • Caldwell

Middle Coniacian to middle Santonian.

Niobrara Chalk
Eutaw Formation

A plioplatecarpine mosasaur, a new genus for "Clidastes" planifrons (Cope, 1874).

Newly named snakes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Madtsoia pisdurensis[95]

Species

Valid

  • Mohabey
  • Head
  • Wilson

Late Cretaceous

A madtsoiid snake.

Archosauromorphs[edit]

Newly named basal archosauromorphs[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Archeopelta[96]

Gen. et sp.

Valid

  • Desojo
  • Ezcurra
  • Schultz

Late Ladinian/Early Carnian

Santa Maria Formation

A doswelliid.

Augustaburiania[97]

Gen. et sp.

Valid

  • Sennikov

Early Triassic

Don River Basin

The earliest tanystropheid.

Kyrgyzsaurus[98]

Gen. et sp.

Valid

  • Alifanov
  • Kurochkin

Late Ladinian/Early Carnian

Madygen Formation

The earliest drepanosaurid.

Protanystropheus[97]

Gen. et comb.

Valid

  • Sennikov

Middle - Late Triassic

A tanystropheid, a new genus for "Tanystropheus" antiquus (von Huene, 1905).

Newly named crurotarsans[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Acherontisuchus[99]

Gen. et sp.

Valid

  • Hastings
  • Bloch
  • Jaramillo

Middle/Late Paleocene

Cerrejón Formation

A dyrosaurid.

Anteophthalmosuchus[100]

Gen. et sp.

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A goniopholidid.

Arenysuchus[101]

Gen. et sp.

Valid

  • Puértolas
  • Canudo
  • Cruzado-Caballero

Maastrichtian

Tremp Formation

A stem crocodyloid.

Caipirasuchus[102]

Gen. et sp.

Valid

  • Iori
  • Carvalho

Turonian/Santonian

Adamantina Formation

A sphagesaurid.

Campinasuchus[103]

Gen. et sp.

Valid

Turonian/Santonian

Adamantina Formation

A baurusuchid.

Caryonosuchus[104]

Gen. et sp.

Valid

Campanian/Maastrichtian

Adamantina Formation

A sphagesaurid.

Decuriasuchus[105]

Gen. et sp.

Valid

  • França
  • Ferigolo
  • Langer

Ladinian

Santa Maria Formation

Skeletons indicate possible group behavior.

Fruitachampsa[106]

Gen. et sp.

Valid

  • Clark

Late Jurassic

Morrison Formation

A shartegosuchid.

Goniopholis kiplingi[107]

Species

Valid

Berriasian

Purbeck Limestone Group

A goniopholidid.

Goniopholis willetti[100]

Species

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A goniopholidid. Originally described as a species of Goniopholis; it was subsequently made the type species of a separate genus Hulkepholis.[108]

Labidiosuchus[109]

Gen. et sp.

Valid

Maastrichtian

Marília Formation

A notosuchian.

Leiokarinosuchus[100]

Gen. et sp.

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A neosuchian.

Lorosuchus[110]

Gen. et sp.

Valid

  • Pol
  • Powell

Paleocene

Rio Loro Formation

A sebecid.

Neptunidraco[111]

Gen. et sp.

Valid

  • Cau
  • Fanti

late Bajocian/early Bathonian

Rosso Ammonitico Veronese Formation

The oldest known metriorhynchid.

Pepesuchus[112]

Gen. et sp.

Valid

Campanian/Maastrichtian

Presidente Prudente Formation

A peirosaurid.

Pietraroiasuchus[113]

Gen. et sp.

Valid

Early Albian

Pietraroia Plattenkalk

A hylaeochampsid.

Pissarrachampsa[114]

Gen. et sp.

Valid

  • Montefeltro
  • Larsson
  • Langer

Campanian/Maastrichtian

Vale do Rio do Peixe Formation

A baurusuchid. The type species is Pissarrachampsa sera.

Theriosuchus grandinaris[115]

Species

Valid

Berriasian/Barremian

Khua Formation

A fifth species of Theriosuchus.

Newly named birds[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Alca carolinensis [116]

Sp. nov

Valid

  • Smith & Clarke

Early Pliocene

Yorktown Formation

An auk related to the razorbill.

Alca minor [116]

Sp. nov

Valid

  • Smith & Clarke

Early Pliocene

Yorktown Formation

An auk related to the razorbill.

Alca olsoni [116]

Sp. nov

Valid

  • Smith & Clarke

Early Pliocene

Yorktown Formation

An auk related to the razorbill.

Alectroenas payandeei [117]

Sp. nov

Valid

  • Julian Pender Hume

Holocene-Recent

An pigeon Columbidae from Rodrigues Island.

?Aptornis proasciarostratus [118]

Sp. nov.

Valid

  • Worthy
  • Tennyson
  • Scofield

Early Miocene, Altonian

Manuherikia Group, St Bathans Fauna

The earliest known Aptornithidae Bonaparte, 1856.

Ardea sytchevskayae [119]

Sp. nov.

Valid

  • Zelenkov

Middle Miocene

Earliest member of Ardea

Australlus gagensis [120]

Gen. nov. et Sp. nov.

Valid

  • Worthy
  • Boles

Oligocene-Miocene

Riverleigh

Rallidae. The type species is Gallinula disneyi Boles, 2005.

Bohaiornis guoi [121]

Gen. et sp.

Valid

Barremian/Aptian

Yixian Formation

An enantiornithine.

Buteo praebuteo [122]

Sp. nov

Valid

Sobolev in Sobolev & Marisova

Miocene

A buzzard, a species of Buteo.

Buteo sarmathicus [122]

Sp. nov

Valid

Sobolev in Sobolev & Marisova

Miocene

A buzzard, a species of Buteo.

Columba thiriouxi [117]

Sp. nov

Valid

  • Julian Pender Hume

Holocene-Recent

An pigeon Columbidae from Mauritius.

Colymbiculus [123]

Gen. et sp. nov

Valid

Middle Eocene

A gaviiform. The type species is Colymbiculus udovinchenkoi.

Cyrilavis colburnorum [124]

Species

Valid

  • Ksepka
  • Clarke
  • Grande

Eocene

Green River Formation

A halcyornithid stem parrot, a species of Cyrilavis.

Eostrix tsaganica [125]

Sp. nov

Valid

  • Kurochkin & Dyke

Early Eocene

A protostrigid owl, a species of Eostrix.

Falco bulgaricus [126]

Sp. nov

Valid

  • Zlatozar N. Boev

Late Miocene

A Falconidae owl, a species of Falco.

Feducciavis loftini [127]

Gen. et sp. nov.

Valid

  • Storrs L. Olson

Middle Miocene

Calvert Formation, Virginia

A Sternidae close to Anous and Procelsterna.

Gallirallus steadmani [128]

Sp. nov.

Valid

  • Trevor H. Worthy
  • Robert Bollt

Quaternary

A Rallidae as big as Gallirallus philippensis from Tubuai, Austral Islands.

Gracilornis jiufotangensis [129]

Gen. et sp.

Valid

  • Li
  • Hou

Aptian

Jiufotang Formation

An enantiornithine.

Heterostrix [125]

Gen. et sp. nov

Valid

  • Kurochkin & Dyke

Early Oligocene

An owl. The type species is Heterostrix tatsinensis.

Hoazinavis palustris [130]

Gen. et sp.

Valid

  • Mayr
  • Alvarenga
  • Mourer-Chauviré

Late Oligocene/Early Miocene

Tremembé Formation

An early hoatzin.

Itaboravis [131]

Gen. et sp.

Valid

  • Mayr
  • Alvarenga
  • Clarke

Early late Paleocene (Itaboraian).

A possible member of Cariamae. The type species is Itaboravis elaphrocnemoides.

Jamna szybiaki [132]

Gen. et sp.

Valid

Oligocene

Carpathian flysch

A passerine.

Kaiika maxwelli [133]

Gen. et sp.

Valid

  • Fordyce
  • Thomas

Early Eocene

Kauru Formation

A basal penguin.

Laricola intermedia [134]

Sp. nov.

Valid

  • Vanessa L. De Pietri
  • Loďc Costeur
  • Marcel Güntert
  • Gerald Mayr

Early Miocene

MN 2a

A Charadriiformes, Laricolidae De Pietri, Costeur, Güntert et G. Mayr, 2011.

Laricola robusta [134]

Sp. nov.

Valid

  • Vanessa L. De Pietri
  • Loďc Costeur
  • Marcel Güntert
  • Gerald Mayr

Early Miocene

MN 2a

A Charadriiformes, Laricolidae De Pietri, Costeur, Güntert et G. Mayr, 2011.

Lavocatavis africana [135]

Gen. et sp.

Valid

  • Mourer-Chauviré
  • Tabuce
  • et al.[CAL 26]

Eocene

Glib Zegdou Formation

A phorusrhacoid.

Mancalla lucasi [136]

Sp. nov

Valid

  • Smith

Pliocene or Early Pleistocene

Niguel Formation
San Diego Formation

A charadriiform related to auks, a species of Mancalla.

Mancalla vegrandis [136]

Sp. nov

Valid

  • Smith

Pliocene or Early Pleistocene

San Diego Formation
San Mateo Formation

A charadriiform related to auks, a species of Mancalla.

Miomancalla [136]

Gen. et sp. et comb. nov

Valid

  • Smith

Late Miocene to early Pliocene

Capistrano Formation
San Mateo Formation

A charadriiform related to auks, a member of Mancallinae. The type species is Miomancalla howardi; genus also contains "Praemancalla" wetmorei Howard (1966).

Mystiornis cyrili [137]

Gen. et sp.

Valid

Barremian/Aptian

Shestakovo-1 locality

Namaortyx [138]

Gen. et sp. nov

Valid

  • Mourer-Chauviré, Pickford & Senut

Middle Eocene (probably early Lutetian)

A galliform of uncertain phylogenetic placement. The type species is Namaortyx sperrgebietensis.

Nambashag [139]

Gen. et 2 sp.

Valid

  • Worthy

Late Oligocene/Early Miocene

Etadunna Formation
Namba Formation

A cormorant, Nambashag billerooensis is the type species and Nambashag microglaucus is the second species.

Nelepsittacus minimus [140]

Gen. nov. et sp. nov.

Valid

  • Trevor H. Worthy
  • Alan J. D. Tennyson
  • R. Paul Scofield

Early Miocene

St. Bathans Fauna, Manuherikia Section

A Psittaciformes, Nestoridae, smallest species of the genus.

Nelepsittacus donmertoni [140]

Sp. nov.

Valid

  • Trevor H. Worthy
  • Alan J. D. Tennyson
  • R. Paul Scofield

Early Miocene

St. Bathans Fauna, Manuherikia Section

A Psittaciformes, Nestoridae.

Nelepsittacus daphneleeae [140]

Sp. nov.

Valid

  • Trevor H. Worthy
  • Alan J. D. Tennyson
  • R. Paul Scofield

Early Miocene

St. Bathans Fauna, Manuherikia Section

A Psittaciformes, Nestoridae.

Nesoenas cicur [117]

Sp. nov

Valid

  • Julian Pender Hume

Holocene-Recent

An pigeon Columbidae from Mauritius.

Nogusunna conflictoides [141]

Gen. nov et sp. nov

Valid

  • Nikita V. Zelenkov

Middle Miocene

Oshin Formation

A diving duck, Anatidae.

Parahongshanornis chaoyangensis [142]

Gen. et sp.

Valid

  • Li
  • Wang
  • Hou

Early Cretaceous

Jiufotang Formation

A hongshanornithid.

Pellornis [143]

Gen. et sp. nov

Valid

  • Bertelli, Chiappe & Mayr

Earliest Eocene

Fur Formation

A messelornithid. The type species is Pellornis mikkelseni.

Piculoides saulcetensis [144]

Gem. nov. et sp. nov.

Valid

  • Vanessa L. De Pietri
  • Albrecht Manegold
  • Loďc Costeur
  • Gerald Mayr

Early Miocene

MN 2

A Charadriiformes, Laricolidae De Pietri, Costeur, Güntert et G. Mayr, 2011.

Protomelanitta gracilis [141]

Gen. nov et sp. nov

Valid

  • Nikita V. Zelenkov

Middle Miocene

Oshin Formation

A diving duck, Anatidae, a second species of the genus is Protomelanitta shihuibas (Hou, 1985), formaly known as Aythya shihuibas Hou, 1985.

Qiliania graffini [145]

Gen. et sp.

Valid

Aptian

Xiagou Formation

An enantiornithine.

Sharganetta mongolica [141]

Gen. nov et sp. nov

Valid

  • Nikita V. Zelenkov

Middle Miocene

Oshin Formation

A diving duck, Anatidae.

Sternalara milneedwardsi [134]

Sp. nov.

Valid

  • Vanessa L. De Pietri
  • Loďc Costeur
  • Marcel Güntert
  • Gerald Mayr

Early Miocene

MN 2a

A Charadriiformes, family Incertae Sedis.

Sternalara minuta [134]

Gen. nov. et sp. nov.

Valid

  • Vanessa L. De Pietri
  • Loďc Costeur
  • Marcel Güntert
  • Gerald Mayr

Early Miocene

MN 2a

A Charadriiformes, family Incertae Sedis this is the typespecies of the genus.

Tonsala buchanani [146]

Sp. nov

Valid

  • Dyke, Wang & Habib

Early Oligocene, probably also late Eocene

Makah Formation
Pysht Formation

A plotopterid, a species of Tonsala.

Newly named non-avian dinosaurs[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Acristavus[147]

Gen. et sp.

Valid

Campanian

Two Medicine Formation
Wahweap Formation

A brachylophosaurin hadrosaurine.

Ahshislepelta[148]

Gen. et sp.

Valid

  • Burns
  • Sullivan

Late Campanian

Kirtland Formation

An ankylosaurine ankylosaurid.

Albinykus[149]

Gen. et sp.

Valid

Late Cretaceous

An alvarezsaurid theropod.

Amtocephale[150]

Gen. et sp.

Valid

  • Watabe
  • Tsogtbaatar
  • Sullivan

Turonian-Santonian

Baynshire Formation

A pachycephalosaurid.

Angolatitan[151]

Gen. et sp.

Valid

Late Turonian

Itombe Formation

A somphospondylian sauropod.

Arcusaurus[152]

Gen. et sp.

Valid

  • Yates
  • Bonnan
  • Neveling

Lower Jurassic

Elliot Formation

Basal Sauropodomorph

Atacamatitan[153]

Gen. et sp.

Valid

Late Cretaceous

Tolar Formation

A titanosaur.

Brontomerus[154]

Gen. et sp.

Valid

  • Taylor
  • Wedel
  • Cifelli

Aptian/Albian

Cedar Mountain Formation

A camarasauromorph sauropod.

Daemonosaurus[155]

Gen. et sp.

Valid

Late Triassic

Chinle Formation

A primitive theropod.

Delapparentia[156]

Gen. et sp.

Valid

  • Ruiz-Omeñaca

Early Barremian

Camarillas Formation

An iguanodontoid.

Demandasaurus[157]

Gen. et sp.

Valid

  • Fernández-Baldor
  • Canudo
  • et al.[CAL 34]

Late Barremian-early Aptian

Castrillo la Reina Formation

A rebbachisaurid sauropod.

Drusilasaura[158]

Gen. et sp.

Valid

  • Navarrete
  • Casal
  • Martínez

Cenomanian/Turonian

Bajo Barreal Formation

A possible lognkosaurian titanosaur.

Eodromaeus[159]

Gen. et sp.

Valid

Middle Carnian

Ischigualasto Formation

A basal theropod.

Epichirostenotes[160]

Gen. et sp.

Valid

  • Sullivan
  • Jasinski
  • Van Tomme

Late Campanian

Horseshoe Canyon Formation

A caenagnathid.

Haya[161]

Gen. et sp.

Valid

Late Cretaceous

Javkhlant Formation

Basal ornithopod with many gastroliths.

Jaklapallisaurus[162]

Gen. et. sp.

Valid

Late Norian-earliest Rhaetian

Upper Maleri Formation
Lower Dharmaram Formation

A plateosaurid.

Koreaceratops[163]

Gen. et sp.

Valid

  • Lee
  • Ryan
  • Kobayashi

Albian

"Tando beds"

A basal ceratopsian.

Koreanosaurus[164]

Gen. et sp.

Valid

?Santonian/Campanian

Seonso Conglomerate

A hypsilophodontid ornithopod.

Leonerasaurus[165]

Gen. et sp.

Valid

  • Pol
  • Garrido
  • Cerda

Pliensbachian/Toarcian

Las Leoneras Formation

An anchisaurian sauropodomorph.

Leyesaurus[166]

Gen. et sp.

Valid

Lower Jurassic

Quebrada del Barro Formation

A massospondylid sauropodomorph.

Linhenykus[167]

Gen. et sp.

Valid

Campanian

Wulansuhai Formation

An alvarezsaurid.

Linhevenator[168]

Gen. et sp.

Valid

Campanian

Bayan Mandahu Formation

An short-armed troodontid.

Manidens[169]

Gen. et sp.

Vald

  • Pol
  • Rauhut
  • Becerra

Aalenian/Early Bathonian

Cañadón Asfalto Formation

A heterodontosaurid.

Nambalia[162]

Gen. et. sp.

Valid

Late Norian-earliest Rhaetian

Upper Maleri Formation

A basal sauropodomorph.

Narambuenatitan[170]

Gen. et sp.

Valid

  • Filippi
  • García
  • Garrido

lower-middle Campanian

Anacleto Formation

A lithostrotian titanosaur.

Ojoraptorsaurus[160]

Gen. et sp.

Valid

  • Sullivan
  • Jasinski
  • Van Tomme

Lower Maastrichtian

Ojo Alamo Formation

A caenagnathid.

Omeisaurus jiaoi[171]

Species

Valid

Middle Jurassic

Xiashaximiao Formation

A species of Omeisaurus.

Osmakasaurus[172]

Gen. et comb.

Valid

  • McDonald

Valanginian

Lakota Formation

A new genus for "Camptosaurus" depressus (Gilmore, 1909).

Oxalaia[173]

Gen. et sp.

Valid

Cenomanian

Alcântara Formation

Huge spinosaur 12–14 m long.

Pampadromaeus[174]

Gen. et sp.

Valid

Carnian

Santa Maria Formation

A very basal sauropodomorph.

Pamparaptor[175]

Gen. et sp.

Valid

  • Porfiri
  • Calvo
  • Santos

Turonian/Coniacian

Portezuelo Formation

A deinonychosaur.

Petrobrasaurus[176]

Gen. et sp.

Valid

Santonian

Plottier Formation

A titanosaur.

Propanoplosaurus[177]

Gen. et sp.

Valid

  • Stanford
  • Weishampel
  • Deleon

Early Aptian

Patuxent Formation

A nodosaurid.

Qiupalong[178]

Gen. et sp.

Valid

Late Cretaceous

Qiupa Formation

An ornithomimid.

Ratchasimasaurus[179]

Gen. et sp.

Valid

  • Shibata
  • Jintasakul
  • Azuma

Aptian

Khok Kruat Formation

A non-hadrosaurid iguanodontian.

Sarahsaurus[180]

Gen. et sp.

Valid

Sinemurian/Pliensbachian

Kayenta Formation

A basal sauropodomorph

Siamodon[181]

Gen. et sp.

Valid

  • Buffetaut
  • Suteethorn

Aptian

Khok Kruat Formation

An advanced iguanodontian.

Spinops[182]

Gen. et sp.

Valid

Campanian

Oldman or Dinosaur Park Formation

A centrosaurine ceratopsian.

Stegoceras novomexicanum[183]

Species

Valid

  • Jasinski
  • Sullivan

Late Campanian

Fruitland Formation
Kirtland Formation

Second known species of Stegoceras.

Talos[184]

Gen. et sp.

Valid

Late Campanian

Kaiparowits Formation

A troodontid theropod.

Tapuiasaurus[185]

Gen. et sp.

Valid

Aptian

Quiricó Formation

A titanosaur.

Teratophoneus[186]

Gen. et sp.

Valid

Late Campanian

Kaiparowits Formation

A tyrannosaurine tyrannosaurid.

Thescelosaurus assiniboiensis[187]

Species

Valid

  • Brown
  • Boyd
  • Russell

Late Maastrichtian

Frenchman Formation

Third known species of Thescelosaurus.

Titanoceratops[188]

Gen. et sp.

Valid

  • Longrich

Late Campanian

Fruitland or Kirtland Formation

A giant chasmosaurine ceratopsian from New Mexico.

Traukutitan[189]

Gen. et sp.

Valid

  • Juárez Valieri
  • Calvo

Santonian

Bajo de la Carpa Formation

A possible lognkosaurian titanosaur.

Uteodon[172]

Gen. et comb.

Valid

  • McDonald

Late Jurassic

Morrison Formation

A new genus for "Camptosaurus" aphanoecetes (Carpenter & Wilson, 2008).

Veterupristisaurus[190]

Gen. et sp.

Valid

  • Rauhut

Late Kimmeridgian/earliest Tithonian

Tendaguru Formation

A carcharodontosaurid.

Xiaotingia[191]

Gen. et sp.

Valid

early Late Jurassic

Tiaojishan Formation

An Archaeopteryx-like paravian.

Xuwulong[192]

Gen. et sp.

Valid

  • You
  • Li
  • Liu

Early Cretaceous

Xinminpu Group

A hadrosauriform.

Zhuchengtyrannus[193]

Gen. et sp.

Valid

Late Cretaceous

Wangshi Group

A tyrannosaurine tyrannosaurid.

Newly named pterosaurs[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aurorazhdarcho[194]

Gen. et sp.

Valid

  • Frey
  • Meyer
  • Tischlinger

Tithonian

Solnhofen limestone

An azhdarchoid[194] or a ctenochasmatid.[195] The type species, Aurorazhdarcho primordius, was subsequently found to be synonymous with "Pterodactylus" micronyx von Meyer (1856); however, Aurorazhdarcho was retained as a distinct genus, creating a new combination Aurorazhdarcho micronyx.[195]

Aussiedraco[196]

Gen. et sp.

Valid

  • Kellner
  • Rodrigues
  • Costa

Albian

Toolebuc Formation

An ornithocheiroid.

Barbosania[197]

Gen. et sp.

Valid

  • Elgin
  • Frey

Cenomanian

Santana Formation

An ornithocheirid.

Darwinopterus robustodens[198]

Species

Valid

Middle Jurassic

Tiaojishan Formation

A darwinopterid.

Jianchangopterus[199]

Gen. et sp.

Valid

  • Bo

Middle Jurassic

Tiaojishan Formation

A rhamphorhynchid.

Navajodactylus[200]

Gen. et sp.

Valid

  • Sullivan
  • Fowler

Late Campanian

Kirtland Formation
Dinosaur Park Formation

A possible azhdarchid.

Pterofiltrus[201]

Gen. et sp.

Valid

  • Jiang
  • Wang

Early Cretaceous

Yixian Formation

A ctenochasmatid.

Unwindia[202]

Gen. et sp.

Valid

  • Martill

Cenomanian

Santana Formation

A basal ctenochasmatoid.

Synapsids[edit]

Newly named non-mammalian synapsids[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Annatherapsidus postum[203]

Sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Basilodon[204]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" woodwardi.

Bridetherium[205]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

A morganucodontan.

Candelariodon[206]

Gen. et sp.

Valid

Middle Triassic

Santa Maria Formation

A carnivorous cynodont.

Cryptovenator[207]

Gen. et sp.

Valid

Late Gzhelian

Remigiusberg Formation

A sphenacodontid pelycosaur. The type species is Cryptovenator hirschbergeri.

Euptychognathus[204]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" bathyrhynchus.

Euromycter[208]

Gen. et comb.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid. A new genus for "Casea" rutena.

Gorochovetzia[203]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Keyseria[204]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" benjamini.

Muchia[203]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Paceyodon[205]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

A morganucodontan.

Peramodon[204]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" amalitzkii

Perplexisaurus lepusculus[203]

Species

Valid

  • Ivakhnenko

Middle Permian

A therocephalian.

Purlovia[203]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian with a short skull.

Ruthenosaurus[208]

Gen. et sp.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid. The type species is Ruthenosaurus russellorum

Syops[204]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" vanhoepeni.

Tiarajudens[209]

Gen. et sp.

Valid

Capitanian

Rio do Rasto Formation

A basal, saber-toothed herbivorous anomodont. The type species is Tiarajudens eccentricus.

Newly named non-eutherian mammals[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Cronopio[210]

Gen. et sp.

Valid

  • Rougier
  • Apesteguía
  • Gaetano

Cenomanian

Candeleros Formation

A "saber-toothed" dryolestoid.

Crusafontia amoae[211]

Species

Valid

Hauterivian to Barremian

A dryolestidan mammal.

Guggenheimia crocheti[212]

Sp. nov

Valid

  • Oliveira & Goin

Itaboraian

Itaboraí Basin

A protodidelphid "ameridelphian", a species of Guggenheimia.

Iberica[213]

Gen. et sp.

Valid

  • Badiola
  • Canudo
  • Cuenca-Bescós

Hauterivian/Barremian

Galve

A eobaatarid or a possible plagiaulacid multituberculate.

Invictokoala[214]

Gen. et sp.

Valid

  • Price
  • Hocknull

Middle Pleistocene

A Koala.

Liaoconodon[215]

Gen. et sp.

Valid

  • Meng
  • Wang
  • Li

Aptian

Jiufotang Formation

An "eutriconodont".

Lutreolina materdei[216]

Species

Valid

  • Goin & de los Reyes

Late Miocene (Huayquerian)

A relative of lutrine opossum.

Naraboryctes[217]

Gen. et sp.

Valid

Early Miocene

A marsupial mole.

Oklatheridium minax[218]

Species

Valid

  • Davis
  • Cifelli

Aptian-Albian

Antlers Formation

A deltatheroidan mammal.

Peradectes coprexeches[219]

Species

Valid

  • Williamson
  • Taylor

Early Paleocene

Nacimiento Formation

A peradectid metatherian, a species of Peradectes.

Periprotodidelphis[212]

Gen. et sp. nov

Valid

  • Oliveira & Goin

Itaboraian

Itaboraí Basin

A protodidelphid "ameridelphian". The type species is Periprotodidelphis bergqvistae.

Sairadelphys[220]

Gen. et sp. nov

Valid

  • Oliveira et al.

Pleistocene

A relative of Kalinowski's mouse opossum. The type species is Sairadelphys tocantinensis.

Sineleutherus issedonicus[221]

Species

Valid

  • Averianov
  • Lopatin
  • Krasnolutskii

Bathonian

Itat Formation

A haramiyidan allotherian.

Swaindelphys encinensis[219]

Species

Valid

  • Williamson
  • Taylor

Early Paleocene

Nacimiento Formation

A herpetotheriid metatherian, a species of Swaindelphys.

Swaindelphys johansoni[219]

Species

Valid

  • Williamson
  • Taylor

Early Paleocene

Nacimiento Formation

A herpetotheriid metatherian, a species of Swaindelphys.

Newly named eutherians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Adelpharctos ginsburgi[222]

Sp. nov

Valid

  • de Bonis

Late Oligocene

A hemicyonine bear, a species of Adelpharctos.

Aegyptocetus[223]

Gen. et sp.

Valid

  • Bianucci
  • Gingerich

Middle Eocene

Gebel Hof Formation

A protocetid cetacean.

Agriarctos beatrix[224]

Sp. nov

Valid

  • Abella, Montoya & Morales

Middle Miocene

Calatayud-Daroca Basin

An ailuropodine bear, originally described as a species of Agriarctos. It was later made the type species of the genus Kretzoiarctos.[225]

Alilepus elongatus[226]

Sp. nov

Valid

  • Winkler
  • Flynn
  • Tomida

Late Miocene

Dhok Pathan Formation

A leporid lagomorph, a species of Alilepus.

Antarctodon[227]

Gen. et sp.

Valid

Late Ypresian

La Meseta Formation

An astrapotherian.

Arazamys[228]

Gen. et sp. nov

Valid

  • Rinderknecht, Bostelmann & Ubilla

Late Miocene (Huayquerian)

Camacho Formation

A dinomyid rodent. The type species is Arazamys castiglionii.

Asilifelis[229]

Gen. et sp. nov

Valid

  • Werdelin

Lower Miocene

Hiwegi Formation

A felid. The type species is Asilifelis coteae.

Basirepomys romensis[230]

Sp. nov

Valid

  • Korth

Late Miocene (Hemphillian)

A cricetid rodent, a species of Basirepomys.

Betonnia[231]

Gen. et sp.

Valid

  • Williamson
  • Weil
  • Standhardt

Middle/Late Puercan

Nacimiento Formation

A cimolestid cimolesta.

Balochititanops[232]

Gen. et sp.

Valid

  • Missiaen
  • Gunnell
  • Gingerich

Ypresian

Ghazij Formation

A brontotheriid.

Brachygaulus[233]

Gen. et 3 sp. nov

Valid

  • Korth
  • Tabrum

Early Oligocene (Orellan)

A rodent related to mylagaulids and the mountain beaver. The type species is Brachygaulus nicholsi; genus also contains Brachygaulus leistneri and Brachygaulus xerobothrus.

Caviodon cuyano[234]

Species

Valid

Late Miocene or Pliocene.

Aisol Formation

A cardiomyine rodent, a species of Caviodon.

Cernictis repenningi[235]

Sp. nov

Valid

  • Baskin

Miocene (middle Hemphillian)

Bidahochi Formation

A galictine mustelid, a species of Cernictis.

Chacomylus[236]

Gen. et sp.

Valid

  • Williamson
  • Weil

Paleocene

Nacimiento Formation

A hyopsodontid condylarth.

Chacopterygus[231]

Gen. et sp.

Valid

  • Williamson
  • Weil
  • Standhardt

Middle Puercan

Nacimiento Formation

A cimolestid cimolesta.

Coelodonta thibetana[237]

Species

Valid

Middle Pliocene

Woolly rhinoceros.

Comahuetherium[238]

Gen. et sp.

Valid

  • Kramarz
  • Bond

Colhuehuapian

Cerro Bandera Formation

An astrapotheriid.

Cynthiacetus peruvianus[239]

Species

Valid

  • Martínez-Cáceres
  • Muizon

Late Eocene-Early Oligocene

Otuma Formation

A basilosaurid cetacean.

Delotrochanter[240]

Gen. et 3 sp. nov.

Valid

  • Hunt

Arikareean

A temnocyonine. The type species is Delotrochanter oryktes; genus also contains D. petersoni and D. major

Diplacodon gigan[241]

Species

Valid

  • Mihlbachler

Middle Eocene

Wiggins Formation

A brontotheriid.

Eoconodon hutchisoni[242]

Sp. nov

Valid

  • Clemens

Early Paleocene (Puercan)

Tullock Formation

A triisodontid, a species of Eoconodon.

Eotitanops pakistanensis[232]

Species

Valid

  • Missiaen
  • Gunnell
  • Gingerich

Ypresian

Ghazij Formation

A brontotheriid.

Eumys euryodus[243]

Sp. nov

Valid

  • Korth

Oligocene (Whitneyan)

A cricetid rodent, a species of Eumys.

Griphotherion[244]

Gen. et sp.

Valid

  • López
  • Powell

Eocene

A rodent-like notoungulate.

Hiskatherium[245]

Gen. et sp.

Valid

  • Pujos
  • Iuliis
  • Quispe

Santacrucian

A small ground sloth.

Hystrix paukensis[246]

Species

Valid

  • Yuichiro Nishioka
  • Zin-Maung-Maung-Thein
  • et al.[CAL 63]
late Miocene/early Pliocene new species in the genus Hystrix (Old World porcupine).

Jacobsomys dailyi[247]

Sp. nov

Valid

  • May et al.

Miocene (late Hemphillian)

Horned Toad Formation

A sigmodontine rodent, a species of Jacobsomys.

Juramaia[248]

Gen. et sp.

Valid

Callovian to Bathonian

Tiaojishan Formation

A basal eutherian. The type species is Juramaia sinensis.

Karydomys debruijni[249]

Species

Valid

  • Maridet et al.

Early Miocene

A cricetid, a species of Karydomys.

Khoratpithecus ayeyarwadyensis[250]

Species

Valid

Late Miocene

Irrawaddy Formation

A hominoid.

Kuntinaru[251]

Gen. et sp.

Valid

Deseadan

A dasypodid armadillo.

Lycophocyon[252]

Gen. et sp.

Valid

  • Tomiya

?Uintan-Duchesnean

Santiago Formation

A caniformian carnivoran.

Mammacyon ferocior[240]

Sp. nov.

Valid

  • Hunt

Mid- or early late Arikareean

Arikaree Group

A temnocyonine, a species of Mammacyon

Martes ginsburgi[253]

Sp. nov

Valid

  • Montoya, Morales & Abella

Upper Miocene

A marten.

Megacricetodon beijiangensis[249]

Species

Valid

  • Maridet et al.

Early Miocene

A cricetid, a species of Megacricetodon.

Megapeomys repenningi[254]

Sp. nov

Valid

  • Tomida

Early Miocene

Nakamura Formation

An apeomyine eomyid rodent, a species of Megapeomys.

Mescalerolemur[255]

Gen. et sp.

Valid

  • Kirk
  • Williams

Uintan

Devil’s Graveyard Formation

An adapiform strepsirrhini.

Mesolambdolophus[256]

Gen. et sp.

Valid

  • Holbrook
  • Lapergola

Bridgerian North American Stage

Bridger Formation

A small odd-toed ungulate of uncertain phylogenetic position, possibly related to Tapiromorpha. The type species is Mesolambdolophus setoni.

Mesoprocta[257]

Gen. et sp. nov

Valid

Croft, Chick & Anaya

Middle Miocene

A dasyproctid rodent. The type species is Mesoprocta hypsodus.

Microcardiodon[258]

Gen. et sp. nov

Valid

Pérez & Vucetich

Middle Miocene

A member of Hystricognathi, a cavioid rodent. The type species is Microcardiodon williensis.

Microtus (Pedomys) parmaleei[259]

Sp. nov

Valid

  • Martin
  • Peláez-Campomanes
  • Honey

Latest Pleistocene

An arvicoline rodent, a species of Microtus.

Mionothropus[260]

Gen. et sp.

Valid

  • De Iuliis
  • Gaudin
  • Vicars

Huayquerian

A megatheriid sloth.

Miotomodon[230]

Gen. et sp. nov

Valid

  • Korth

Late Miocene (Hemphillian)

A cricetid rodent. The type species is Miotomodon mayi.

Mondegodon[261]

Gen. et sp.

Valid

  • Tabuce
  • Clavel
  • Antunes

Earliest Eocene

A mesonychian.

Nevadomys[262]

Gen. et 3 sp. nov

Valid

  • Mou

Pliocene (Blancan)

Panaca Formation

A relative of the long-clawed mole vole. The type species is Nevadomys fejfari; genus also contains Nevadomys lindsayi and Nevadomys downsi.

Nuralagus[263]

Gen. et sp.

Valid

  • Quintana
  • Köhler
  • Moyà-Solà

Early Pliocene

A leporid rabbit.

Ocucajea[264]

Gen. et sp.

Valid

Bartonian

Paracas Formation

A basilosaurid cetacean.

Oldrichpedetes[265]

Gen. et comb. et 3 sp. nov

Valid

  • Pickford & Mein

Early Miocene to early Pliocene

A pedetid rodent, a new genus for "Megapedetes" pickfordi (Mein & Senut, 2003). Genus also contains 3 new species: Oldrichpedetes fejfari, O. brigitteae and O. praecursor.

Paranotiosorex[266]

Gen. et sp. nov

Valid

  • Mou

Pliocene

Panaca Formation

A shrew. The type species is Paranotiosorex panacaensis.

Paronychomys shotwelli[230]

Sp. nov

Valid

  • Korth

Late Miocene (Hemphillian)

A cricetid rodent, a species of Paronychomys.

Phoxomylus[267]

Gen. et sp.

Valid

  • Fox

Early Tiffanian

Paskapoo Formation

A palaechthonid plesiadapiform.

Pliosaccomys prowitteorum[80]

Sp. nov

Valid

  • Martin
  • Mallory

Late Miocene (Hemphillian)

A pocket gopher, a species of Pliosaccomys.

Pliospalax complicatus[268]

Sp. nov.

Valid

  • Şen
  • Sarica

Miocene

A spalacid rodent, a species of Pliospalax.

Postcopemys[269]

Gen. et 2 sp. et comb. nov

Valid

  • Lindsay
  • Czaplewski

Pliocene (late Hemphillian and early Blancan)

A cricetid rodent. The type species is Postcopemys repenningi; genus also contains Postcopemys maxumensis, as well as "Peromyscus" valensis Shotwell (1967) and "Copemys" vasquezi Jacobs (1977).

Prothomomys[247]

Gen. et sp. nov

Valid

  • May et al.

Miocene (late Hemphillian)

Horned Toad Formation

A gopher. The type species is Prothomomys warrenensis.

Protoryx tuvaensis[270]

Species

Valid

  • Dmitrieva
  • Serdyuk

Late Miocene

A bovid.

Protungulatum coombsi[271]

Species

Valid

Maastrichtian

Hell Creek Formation

A species of Protungulatum.

Quebradahondomys[257]

Gen. et sp. nov

Valid

Croft, Chick & Anaya

Middle Miocene

An adelphomyine spiny rat. The type species is Quebradahondomys potosiensis.

Repomys minor[262]

Sp. nov

Valid

  • Mou

Pliocene (early Blancan)

Panaca Formation

A cricetid rodent, a species of Repomys.

Rudiocyon[240]

Gen. et sp. nov.

Valid

  • Hunt

Arikareean

John Day Formation

A temnocyonine. The type species is Rudiocyon amplidens

Rusingapedetes[265]

Gen. et sp. nov

Valid

  • Pickford & Mein

Early Miocene

A pedetid rodent. The type species is Rusingapedetes tsujikawai.

Rytiodus heali[272]

Species

Valid

  • Domning
  • Sorbi

Miocene

A dugongid sirenian, a species of Rytiodus.

Sicista primus[273]

Sp. nov

Valid

  • Kimura

Early Miocene (~17 Ma)

A birch mouse.

Sinapospalax berdikensis[268]

Sp. nov.

Valid

  • Şen
  • Sarica

Miocene

A spalacid rodent, a species of Sinapospalax.

Supayacetus[264]

Gen. et sp.

Valid

Bartonian

Paracas Formation

A basilosaurid cetacean.

Tapirus rondoniensis[274]

Sp. nov

Valid

  • Holanda, Ferigolo & Ribeiro

Late Pleistocene

Rio Madeira Formation

A tapir.

Temnocyon fingeruti[240]

Sp. nov.

Valid

  • Hunt

Latest Oligocene, late mid-Arikareean

John Day Formation

A temnocyonine, a species of Temnocyon

Temnocyon macrogenys[240]

Sp. nov.

Valid

  • Hunt

Latest Arikareean

Anderson Ranch Formation

A temnocyonine, a species of Temnocyon

Temnocyon subferox[240]

Sp. nov.

Valid

  • Hunt

Early Arikareean

John Day Formation

A temnocyonine, a species of Temnocyon

Turiasorex[275]

Gen. et sp.

Valid

  • van Dam
  • van den Hoek Ostende
  • Reumer

Middle - Late Miocene

A short-snouted shrew.

Umayodus[276]

Gen. et sp.

Valid

  • Gelfo
  • Sigé

Late Paleocene/Earliest Eocene

Muñami Formation

A didolodontid "condylarth".

Plants[edit]

Angiosperms[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Palmoxylon lametaei[277]

Species

Valid

  • Dutta
  • Ambwani
  • Estrada-Ruiz

Maastrichtian

Lameta Formation

Leefructus[278]

Gen. et sp.

Valid

Early Cretaceous

Yixian Formation

A basal eudicot

Gymnosperms[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Qataniaria[279]

Gen. et sp.

Valid

  • Krassilov
  • Schrank

Albian

Hatira Formation

Cariria[280]

Gen. et sp.

Valid

Kunzmann, Mohr, Wilde & Bernardes-de-Oliveira

Aptian

Crato Formation

Pteridophytes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Equisetum thermale[281]

sp. nov

Valid

Channing, Zamuner, Edwards, & Guido

Callovian - Tithonian

La Matilde Formation

 Argentina

oldest crown group Equisetum species

Footnotes[edit]

Complete author list[edit]

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.

  1. ^ Gai, Donoghue, Zhu, Janvier, Stampanoni.
  2. ^ Underwood, Goswami Prasad, Verma and Flynn
  3. ^ Fischer, Voigt, Schneider, Buchwitz and Voigt
  4. ^ Pradel, Tafforeau, Maisey, Janvier.
  5. ^ Downs, Daeschler, Jenkins and Shubin.
  6. ^ Wu, Sun, Xu, Hao, Jiang, Sun.
  7. ^ Henrici, Martens, Berman, Sumida.
  8. ^ Berman, Henrici, Martens, Sumida, Anderson
  9. ^ a b Reisz, Liu, Li, Müller.
  10. ^ Gaffney, Meylan, Wood, Simons and Campos
  11. ^ Slater, Reolid, Wood, Schouten and Benton
  12. ^ Knauss, Joyce, Lyson and Pearson
  13. ^ de la Fuente, Umazano, Sterli and Carballido
  14. ^ Danilov, Zvonok, Syromyatnikova and Udovichenko
  15. ^ Fischer, Masure, Arkhangelsky, Godefroit.
  16. ^ Liu, Rieppel, Jiang, Aitchison, Motani, Zhang, Zhou, Sun.
  17. ^ a b Benson, Ketchum, Noè, Gómez-Pérez.
  18. ^ Müller, Hipsley, Head, Kardjilov, Hilger, Wuttke, Reisz.
  19. ^ a b Carvalho, Teixeira, Ferraz, Ribeiro, Martinelli, Neto, Sertich, Cunha, Cunha, Ferraz.
  20. ^ Kellner, Campos, Riff, de Andrade.
  21. ^ a b de Andrade, Edmonds, Benton, Schouten.
  22. ^ Buscalioni, Piras, Vullo, Signore, Barbera
  23. ^ Lauprasert, Laojumpon, Saenphala, Cuny, Thirakhupt, Suteethorn
  24. ^ Hu, Li, Hou, Xu.
  25. ^ Bocheński, Tomek, Bujoczek, Wertz.
  26. ^ Mourer-Chauviré, Tabuce, Mahboubi, Adaciy, Bensalah.
  27. ^ Kurochkin, Zelenkov, Averianov, Leshchinskiy.
  28. ^ Ji, Atterholt, O'Connor, Lamanna, Harrs, Li, You, Dodson.
  29. ^ Gates, Horner, Hanna, Nelson.
  30. ^ Nesbitt, Clarke, Turner, Norell.
  31. ^ Mateus, Jacobs, Schulp, Polcyn, Tavares, Neto, Morais, Antunes.
  32. ^ Kellner, Rubilar-Rogers, Vargas, Suárez
  33. ^ Sues, Nesbitt, Berman, Henrici
  34. ^ Fernández-Baldor, Canudo, Huerta, Montero, Pereda Suberbiola, Salgado.
  35. ^ Martinez, Sereno, Alcober, Colombi, Renne, Montañez, Currie.
  36. ^ Makovicky, Kilbourne, Sadleir, Norell.
  37. ^ a b Novas, Ezcurra, Chatterjee, Kutty.
  38. ^ Huh, Lee, Kim, Lim, Godefroit.
  39. ^ Apaldetti, Martinez, Alcober, Pol.
  40. ^ Xu, Sullivan, Pittman, Choiniere, Hone, Upchurch, Tan, Xiao, Tan, Han.
  41. ^ Xu, Tan, Sullivan, Han, Xiao.
  42. ^ Jiang, Li, Peng, Ye
  43. ^ Kellner, Azvedo, Machado, Carvalho, Henriques.
  44. ^ Schultz, Bittencourt, Soares, Fortier, Silva, Langer.
  45. ^ Filippi, Canudo, Salgado, Garrido, Garcia, Cerda, Otero.
  46. ^ Xu, Kobayashi, Lü, Lee, Liu, Tanaka, Zhang, Jia, Zhang.
  47. ^ Farke, Ryan, Barrett, Tanke, Braman, Loewen, Graham.
  48. ^ Zanno, Varricchio, O'Connor, Titus, Knell.
  49. ^ Zaher, Pol, Carvalho, Nascimento, Riccomini, Larson, Juarez-Valieri, Pires-Domingues, da Silva Jr., Campos.
  50. ^ Carr, Williamson, Britt, Stadtman.
  51. ^ Xu, You, Du, Han.
  52. ^ Hone, Wang, Sullivan, Zhao, Chen, Li, Ji, Ji, Xu.
  53. ^ Lü, Xu, Chang, Zhang.
  54. ^ De Oliveira, Schultz, Soares, Rodrigues.
  55. ^ Fröbisch, Schoch, Müller, Schindler, Schweiss.
  56. ^ a b Reisz, Maddin, Fröbisch, Falconnet.
  57. ^ Cisneros, Abdala, Rubidge, Dentzien-Dias, Bueno.
  58. ^ Cuenca-Bescós, Badiola, Canudo, Gasca and Moreno-Azanza.
  59. ^ Archer, Beck, Gott, Hand, Godthelp, Black.
  60. ^ Bond, Kramarz, MacPhee, Reguero.
  61. ^ Vucetich, Deschamps, Morgan and Forasiepi.
  62. ^ Deng, Wang, Fortelius, Li, Wang, Tseng, Takeuchi, Saylor, Säilä, Xie
  63. ^ Naoko Egi, Takehisa Tsubamoto, Takeshi Nishimura, Tsuyoshi Ito, Thaung-Htike and Masanaru Takai.
  64. ^ Luo, Yuan, Meng, Ji.
  65. ^ Jaeger, Soe, Chavasseau, Coster, Emonet, Guy, Lebrum, Maung, Shwe, Tun, Rugbumrung, Bocherens, Benammi, Chaivanich, Tafforeau, Chaimanee.
  66. ^ Billet, Hautier, de Muizon, Valentin.
  67. ^ a b Uhen, Pyenson, Devries, Urbina, Renne.
  68. ^ Archibald, Zhang, Harper and Cifelli
  69. ^ Sun, Dilcher, Wang, Chen.

References[edit]

  1. ^ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716. 
  2. ^ Liu, J.; Steiner, M.; Dunlop, J. A.; Keupp, H.; Shu, D.; Ou, Q.; Han, J.; Zhang, Z.; Zhang, X. (2011). "An armoured Cambrian lobopodian from China with arthropod-like appendages". Nature 470 (7335): 526–530. Bibcode:2011Natur.470..526L. doi:10.1038/nature09704. PMID 21350485.  edit
  3. ^ Vinther, J.; Smith, M. P.; Harper, D. A. T. (2011). "Vetulicolians from the Lower Cambrian Sirius Passet Lagerstätte, North Greenland, and the polarity of morphological characters in basal deuterostomes". Palaeontology 54 (3): 711–719. doi:10.1111/j.1475-4983.2011.01034.x.  edit
  4. ^ Selden, P. A.; Shih, C.K.; Ren, D. (2011). "A golden orb-weaver spider (Araneae: Nephilidae: Nephila) from the Middle Jurassic of China". Biology Letters 7 (5): 775–8. doi:10.1098/rsbl.2011.0228. PMC 3169061. PMID 21508021. 
  5. ^ a b Selden, P. A.; Shih, C.K.; Ren, D. (2013). "A giant spider from the Jurassic of China reveals greater diversity of the orbicularian stem group". Naturwissenschaften. doi:10.1007/s00114-013-1121-7. 
  6. ^ Engel, Michael S. (2011). "A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae)". Transactions of the Kansas Academy of Science 114 (1-2): 77–87. doi:10.1660/062.114.0107. 
  7. ^ Heterick, B. E.; Shattuck, S. (2011). "Revision of the ant genus Iridomyrmex (Hymenoptera: Formicidae)". Zootaxa 2845: 169. 
  8. ^ Yunakov, N.N.; Kirejtshuk, A.G. (2011). "New genus and species of broad-nosed weevils from Baltic amber and notes on fossils of the subfamily Entiminae (Coleoptera, Curculionidae)". ZooKeys 160: 73–96. doi:10.3897/zookeys.160.2108. PMC 3253632. PMID 22303121. 
  9. ^ a b Engel, M.S. (2011). "New earwigs in mid-Cretaceous amber from Myanmar (Dermaptera, Neodermaptera)". ZooKeys 130: 137–152. doi:10.3897/zookeys.130.1293. 
  10. ^ Mey, Wolfram (2011). "On the systematic position of Baltimartyria Skalski, 1995 and description of a new species from Baltic amber (Lepidoptera, Micropterigidae)". ZooKeys 130: 331–342. doi:10.3897/zookeys.130.1480. 
  11. ^ Guglielmino, A.; Olmi, M. (2011). "Revision of fossil species of Deinodryinus, with description of a new species (Hymenoptera, Dryinidae)". ZooKeys 130: 495–504. doi:10.3897/zookeys.130.1326. PMC 3260777. PMID 22259295. 
  12. ^ Chatzimanolis, S.; Engel, M. S. (2011). "A new species of Diochus from Baltic amber (Coleoptera, Staphylinidae, Diochini)". ZooKeys 138: 65–73. doi:10.3897/zookeys.138.1896. 
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