2020 in arthropod paleontology

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List of years in arthropod paleontology
In paleontology
2017
2018
2019
2020
2021
2022
2023
In science
2017
2018
2019
2020
2021
2022
2023
+...

2020 in arthropod palentology is a list of new arthropod fossil taxa, including arachnids, crustaceans, insects, trilobites, and other arthropods that were announced or described, as well as other significant arthropod paleontological discoveries and events which occurred in 2020.

Arachnids

Newly named taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Aliendiguetia[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider of uncertain phylogenetic placement, possibly related to the Ochyroceratoidea. The type species is A. praecursor.

Bicornoculus granulans[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Biungulus[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family Epedanidae. Genus includes new species B. xiai.
Burmacharon[3] Gen. et sp. nov In press Hu et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A whip spider. Genus includes new species B. dunlopi.

Burmesarchaea bilongapophyses[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Archaeidae.

Burmesia[1]

Gen. et sp. nov

Junior homonym

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Nemesiidae. The type species is B. sordida. The generic name is preoccupied by Burmesia Healey (1908); Wunderlich (2021) coined a replacement name Burmesiana.[4]

?Burmesiola kachinensis[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A tree trunk spider.

Burmorchestina circular[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Oonopidae.

Burmorchestina prominens[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Oonopidae.

Burmorsolus globosus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider, probably related to trogloraptorids.

Burmorsolus longembolus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider, probably related to trogloraptorids.

Burmorsolus longibulbus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider, probably related to trogloraptorids.

Consteniusi[5]

Gen. et sp. nov

Valid

Downen & Selden

Eocene

Kishenehn Formation

 United States

An orb-weaver spider. The type species is C. leonae.

Curculioides[6]

Gen. et sp. nov

Valid

Whalen & Selden

Carboniferous (Pennsylvanian)

Energy Shale

 United States

A member of Ricinulei. Genus includes new species C. bohemondi.

Cymbioblemma fusca[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Cymbioblemma hamoembolus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Electroblemma bifurcata[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Electroblemma caula[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Electroblemma pinnae[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Ellenbergellus[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family Tithaeidae. Genus includes new species E. tuberculatus.

Eogamasomorpha rostratis[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Epieremulus sidorchukae[7] Sp. nov In press Arillo, Subías & Peñalver Cretaceous Asturian amber  Spain A mite belonging to the group Oribatida and to the family Caleremaeidae
Gigantocheles[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family Epedanidae. Genus includes new species G. nilsi.

Gigarachne[8]

Gen. et sp. nov

Valid

Jiang & Li in Jiang et al.

Late Cretaceous

Burmese amber

 Myanmar

A spider belonging to the group Araneomorphae and to the new family Gigarachnidae. The type species is G. bian.

Greenwaltarachne[5]

Gen. et sp. nov

Valid

Downen & Selden

Eocene

Kishenehn Formation

 United States

An orb-weaver spider. The type species is G. pamelae.

Hamoderces[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the superfamily Scytodoidea and the family Praepholcidae. The type species is H. opilionoides.

Magnosegestria[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Probably a tube-dwelling spider. The type species is M. tuber.

Mesokanus[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the group Laniatores and to the new family Mesokanidae. Genus includes new species M. oehmkuehnlei.
Mesozomus[9] Gen. et sp. nov Valid Müller et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Schizomida. Genus includes new species M. groehni.
Modisimus chiapanecus[10] Sp. nov In press García-Villafuerte & Valdez-Mondragón Miocene (Aquitanian) Mexican amber  Mexico A species of Modisimus

Myannemesia[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Nemesiidae. The type species is M. glaber.

Myrmecarchaea antecessor[11] Sp. nov Carbuccia et al. Eocene (Ypresian) Oise amber  France A member of the family Archaeidae.
Neophyllobius electrus[12] Sp. nov Valid Zmudzinski Eocene Baltic amber Europe (Baltic Sea region) A mite, a species of Neophyllobius.
Neophyllobius glaesus[12] Sp. nov Valid Zmudzinski Eocene Baltic amber Europe (Baltic Sea region) A mite, a species of Neophyllobius.
Odontomegops[13] Gen. et sp. nov Valid Guo et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A spider belonging to the family Lagonomegopidae. Genus includes new species O. titan.
Odontoscirus cretacico[14] Sp. nov Valid Porta et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A mite belonging to the family Bdellidae.
Palaeobeloniscus[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family Beloniscidae. Genus includes new species P. thilolebi.

Palazarqaraneus[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Zarqaraneidae. The type species is P. hamulus.

Parvocteniza[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Ctenizidae. The type species is P. parvula.

Paurospina fastigata[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Zarqaraneidae.

Pilosarachne[8]

Gen. et sp. nov

Valid

Jiang & Li in Jiang et al.

Late Cretaceous

Burmese amber

 Myanmar

A spider belonging to the group Araneomorphae and to the new family Pilosarachnidae. The type species is P. ju.

Praearaneus araneoides[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Praearaneidae.

Priscaleclercera furcata[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Psilodercidae.

Priscaleclercera hamo[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Psilodercidae.

Priscaleclercera liber[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Psilodercidae.

Procheiridium[15] Gen. et sp. nov Valid Porta et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A pseudoscorpion belonging to the family Cheiridiidae and the subfamily Pycnocheiridiinae. Genus includes new species P. judsoni.

Propterpsiloderces crassitibia[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Eopsilodercidae.

Propterpsiloderces cymbioseta[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Eopsilodercidae.

Propterpsiloderces duplex[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Eopsilodercidae.

Protertheraphosa[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A tarantula. The type species is P. spinipes.

Protopyramidops[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones, possibly belonging to the family Pyramidopidae. Genus includes new species P. nalae.

Spinicreber vacuus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Pholcochyroceridae.

Suraju[16] Gen. et sp. nov In press Martine et al. Permian (Guadalupian) Teresina Formation  Brazil A scorpion. Genus includes new species S. itayma.

Unicornutiblemma[1]

Gen. et 3 sp. et comb. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae. The type species is U. gracilicornis; genus also includes new species U. brevicornis and U. longicornis, as well as "Eogamasomorpha" unicornis Wunderlich (2017).

Crustaceans

Newly named taxa

Malacostracans

Name Novelty Status Authors Age Type locality Country Notes Images
Aliaplax[17] Gen. et sp. nov Valid Pasini et al. Early Pleistocene  Italy A member of the family Goneplacidae. The type species is A. tyrsenorum.

Ammopylocheles romankijoki[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Pylochelidae.

Amphoranina[19] Gen. et 2 sp. nov Valid Nyborg et al. Eocene to Oligocene  Canada
 United States
A crab belonging to the family Raninidae. Genus includes new species A. blandi and A. multispinata.

Ancipitecancer[20]

Gen. et sp. nov

Valid

Pasini, Luque & Garassino

Pleistocene

 Italy

A member of Xanthoidea sensu lato, of uncertain phylogenetic placement within that group. Genus includes new species A. collinsi.

Angusteryon[21]

Gen. et sp. nov

Valid

Audo & Furrer

Early Jurassic

  Switzerland

A member of Polychelida, possibly belonging to the family Coleiidae. Genus includes new species A. oberlii.

Annuntidiogenes sagittula[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Annuntidiogenidae.

Bournelyreidus shiranui[22]

Sp. nov

Valid

Ando in Ando et al.

Late Cretaceous (Cenomanian)

Enokuchi Formation

 Japan

A lyreidid raninoid crab.

Braggitrapezia[23]

Gen. et sp. nov

Valid

De Angeli & Caporiondo

Eocene

 Italy

A crab belonging to the group Trapezioidea. Genus includes new species B. alessandroi.

Caecorchestia[24]

Gen. et sp. nov

Valid

Hegna & Lazo-Wasem in Hegna et al.

Early Miocene

Mexican amber

 Mexico

A member of Amphipoda belonging to the family Talitridae. Genus includes new species C. bousfieldi.

Calappilia erwinhartei[25]

Sp. nov

Valid

Wallaard et al.

Miocene (Burdigalian)

 Portugal

A species of Calappilia.

Callianassa reinhardpalorum[26] Sp. nov Valid Hyžný Miocene  Hungary
 Poland
 Slovakia
A species of Callianassa.
Calliax nishiki[27] Sp. nov Valid Ando et al. Latest Pleistocene to Holocene Moeshima Silt Bed  Japan A member of Axiidea belonging to the family Eucalliacidae.

Callichirus santensis[28]

Sp. nov

In press

Hernáez, Buchmann & Santana

Late Pleistocene

 Brazil

A species of Callichirus.

Cancer nancei[29] Sp. nov Valid Bahman, Feldmann & Schweitzer Miocene St. Marys  United States A species of Cancer
Cancer zahrae[29] Sp. nov Valid Bahman, Feldmann & Schweitzer Miocene St. Marys  United States A species of Cancer
Cherpihomola[30] Gen. et sp. nov Valid Marangon & De Angeli Oligocene (Rupelian)  Italy A member of the family Homolidae. The type species is C. italica.

Collinscarcinus[31]

Gen. nov

Valid

Artal & van Bakel

Eocene (Ypresian)

 Spain

A crab belonging to the family Xanthidae.

Cretacalliax[32] Gen. et sp. nov Valid Pasini, Poore & Garassino Late Cretaceous (Cenomanian)  Lebanon A ghost shrimp belonging to the family Callianopsidae. Genus includes new species C. levantina.
Cuchiadromites[33] Gen. et sp. nov In press Ossó, van Bakel & Ferratges Early Cretaceous (Aptian) Patrocinio Formation  Spain A crab belonging to the family Longodromitidae. Genus includes new species C. jadeae.

Daira corallina[34]

Sp. nov

Valid

Ferratges, Zamora & Aurell

Late Eocene

 Spain

Dardanus cherpionensis[35]

Sp. nov

Valid

Marangon & De Angeli

Oligocene (Rupelian)

Ligure-Piemontese Basin

 Italy

A species of Dardanus.

Dardanus faxensis[36] Sp. nov Valid Jakobsen et al. Paleocene (Danian) Faxe Formation  Denmark A species of Dardanus.
Diogenes augustinus[37] Sp. nov Valid Ferratges, Zamora & Aurell Eocene (Bartonian) Arguis Formation  Spain A hermit crab, a species of Diogenes.
Discosalaputium[38] Gen. et sp. nov Valid Schädel et al. Late Triassic (Carnian) Reingraben  Austria A member of Isopoda belonging to the group Scutocoxifera. The type species is D. aschauerorum.
Dynomene collinsi[39] Sp. nov Valid Ossó et al. Eocene (Priabonian)  Spain A crab belonging to the family Dynomenidae.
Eodromites bernchrisdomiorum[40] Sp. nov Valid Klompmaker et al. Late Jurassic (Oxfordian)  Poland A crab belonging to the family Goniodromitidae.
Eodromites cristinarobinsae[40] Sp. nov Valid Klompmaker et al. Early Cretaceous (Albian) Eguino Formation  Spain A crab belonging to the family Goniodromitidae.

Eopolycheles[41]

Gen. et sp. nov

Valid

Gašparič et al.

Late Triassic (Carnian)

 Slovenia

A member of the family Polychelidae. Genus includes new species E. cornuaureus.

Eryma moriedaorum[22]

Sp. nov

Valid

Ando et al.

Late Cretaceous (Cenomanian)

Enokuchi Formation

 Japan

A member of the family Erymidae.

Etyus tresgalloi[42]

Sp. nov

Valid

Ossó & Moreno-Bedmar

Early Cretaceous (Aptian)

 Spain

A crab belonging to the family Etyiidae.

Eupilumnus? vokesae[43]

Sp. nov

In press

Luque in Luque et al.

Miocene

 Mexico

A crab.

Europrosopon[40] Gen. et comb. nov Valid Klompmaker et al. Late Jurassic  Austria
 Czech Republic
 Germany
 Romania
A crab belonging to the family Prosopidae. The type species is "Prosopon" aculeatum von Meyer (1857); genus also includes "Prosopon" abbreviatum Schweitzer & Feldmann (2009).
Galathea tuscia[17] Sp. nov Valid Pasini et al. Early Pleistocene  Italy A species of Galathea.

Gammaroidorum[44]

Gen. et sp. nov

In press

Jarzembowski et al.

Early Cretaceous (Hauterivian)

Weald

 United Kingdom

A member of Amphipoda. Genus includes new species G. vonki.

Gelrincola[38] Gen. et sp. nov Valid Schädel et al. Middle Triassic (Anisian) Lower Muschelkalk Vossenveld Formation  Netherlands A member of Isopoda belonging to the group Cymothoida. The type species is G. winterswijkensis.

Gemmellarocarcinus riglosensis[34]

Sp. nov

Valid

Ferratges, Zamora & Aurell

Late Eocene

 Spain

Glebocarcinus doii[45]

Sp. nov

Valid

Karasawa & Takahashi

Early Miocene

Igami

 Japan

A member of the family Cancridae

Graptocarcinus collinsi[46]

Sp. nov

Valid

Vega in Vega et al.

Late Cretaceous (Maastrichtian)

Arroyo Grande Formation

 Cuba

A crab belonging to the family Dynomenidae.

Hefriga schlechtingerae[47]

Sp. nov

Valid

Winkler

Late Jurassic (Tithonian)

Solnhofen

 Germany

A caridean shrimp

Hispanigalathea raymondcaseyi[48]

Sp. nov

Valid

Van Bakel et al.

Early Cretaceous (Albian)

Gault

 United Kingdom

A member of Galatheoidea.

Hoploparia echinata[49]

Sp. nov

Valid

Pinheiro et al.

Late Cretaceous (SantonianCampanian)

Santa Marta Formation

Antarctica

Hoploparia hinokunika[22]

Sp. nov

Valid

Ando & Shimada in Ando et al.

Late Cretaceous (Cenomanian)

Enokuchi Formation
Karakizaki Formation

 Japan

A lobster.

Hoploparia tectumque[50]

Sp. nov

In press

Fraaije et al.

Late Cretaceous (Cenomanian)

 Kazakhstan

Icriobranchiocarcinus rodas[51]

Sp. nov

In press

Vega in Vega et al.

Late Cretaceous (Maastrichtian)

Arroyo Grande Formation

 Cuba

A crab belonging to group Portunoidea and the family Lithophylacidae. Announced in 2020; the final version of the article naming it is scheduled to be published in 2021.

Joecalcinus[52]

Gen. et sp. nov

Valid

Fraaije et al.

Eocene

 Italy

A hermit crab belonging to the family Calcinidae. Genus includes new species J. bolcensis.

Juranecrocarcinus[53]

Gen. et comb. nov

In press

Van Bakel et al.

Late Jurassic (Kimmeridgian)

 Germany

A crab belonging to the family Necrocarcinidae. The type species is "Nodoprosopon" angulosum Wehner (1988).

Laeviprosopon musialiki[54]

Sp. nov

Valid

Starzyk

Late Jurassic (Oxfordian)

 Poland

A crab belonging to the family Prosopidae.

Levashidromites[55]

Gen. et sp. nov

In press

Van Bakel et al.

Early Cretaceous (Aptian)

Kubina Formation

 Russia

A crab belonging to the family Longodromitidae. The type species is L. cornutus.

Lobogalenopsis joei[34]

Sp. nov

Valid

Ferratges, Zamora & Aurell

Late Eocene

 Spain

Mariaplax ohiranorikoae [56]

Sp. nov

Valid

Karasawa and Kobayashi

Pleistocene

Atsumi Group

 Japan

A member of the family Hexapodidae.

Marmacuma[57]

Gen. et sp. nov

Valid

Artüz & Sakınç

Late Pleistocene

 Turkey

A member of Cumacea belonging to the family Nannastacidae. Genus includes new species M. samimei.

Masticacheles septemgradu[18]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

 Czech Republic

A hermit crab belonging to the family Pilgrimchelidae.

Matutites collinsi[58]

Sp. nov

Valid

Davis, Garassino & Weaver

Middle Eocene

Castle Hayne

 United States

A member of the family Matutidae.

Maurocarpilius[59] Gen. et sp. nov Valid Ossó, Gagnaison & Bailleul Eocene (Ypresian) Ait Ouarhitane  Morocco A member of the family Carpiliidae. The type species is M. binodosus.
Mesogalathea ardua[60] Sp. nov Valid Gašparič, Robins & Gale Late Jurassic (Kimmeridgian/Tithonian)  Slovenia A member of the family Galatheidae.

Mesoparapylocheles janetjacksonae[18]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

 Czech Republic

A hermit crab belonging to the family Parapylochelidae.

Mizuhocancer[45] Gen. et comb. nov Valid Karasawa in Karasawa & Takahashi EarlyMiddle Miocene  Japan A member of the family Cancridae. Monotypic with the type species, Cancer? Imamurae Imaizumi, 1962.

Montemagrecarcinus[61]

Gen. et sp. nov

Valid

De Angeli & Ceccon

Early Eocene

 Italy

A crab belonging to the family Pilumnidae. Genus includes new species M. efremi.

Moravacarcinus[53]

Gen. et sp. nov

In press

Van Bakel et al.

Early Cretaceous (Berriasian)

Štramberk Limestone

 Czech Republic

A crab belonging to the family Necrocarcinidae. The type species is M. stramberkensis.

Munidopsis kaedetatsuyai [62]

Sp. nov

Valid

Karasawa

Miocene

Higashibessho Formation

 Japan

A member of the family Munidopsidae.

Neoliomera zovoensis[63] Sp. nov Valid De Angeli & Garassino Early Eocene  Italy A species of Neoliomera.

Nitotacarcinus ambrosii[64]

Sp. nov

Valid

Beschin & Checchi

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Carpilioidea and the family Tumidocarcinidae.

Obtusotelson[38] Gen. et sp. nov Valid Schädel et al. Late Triassic (Carnian) Reingraben  Austria A member of Isopoda belonging to the group Scutocoxifera. The type species is O. summesbergeri.

Orithyia eikii[65]

Sp. nov

Valid

Karasawa

Early Miocene

Mizunami Group

 Japan

A member of the family Orithyiidae.

Paguristes collinsi[35]

Sp. nov

Valid

Marangon & De Angeli

Oligocene (Rupelian)

Ligure-Piemontese Basin

 Italy

A species of Paguristes.

Paguristes frigoscopulus[36] Sp. nov Valid Jakobsen et al. Paleocene (Danian) Faxe Formation  Denmark A species of Paguristes.
Paguristes joecollinsi[66] Sp. nov Valid Wallaard et al. Miocene (Tortonian) Pakhna Formation  Cyprus A species of Paguristes.

Paguristes laurae[35]

Sp. nov

Valid

Marangon & De Angeli

Oligocene (Rupelian)

Ligure-Piemontese Basin

 Italy

A species of Paguristes.

Palaeopalinurellus[67]

Gen. et 2 sp. et comb. nov

Valid

Fraaije et al.

Late Jurassic to Late Cretaceous (Cenomanian)[68]

 Czech Republic
 France
 Lebanon[68]

A furry lobster. Genus includes new species P. culocervus and P. jbeilensis,[68] as well as "Palinurus" strambergensis Bachmayer (1959).

Palaeopolycheles nantosueltae[69] Sp. nov Jauvion et al. Middle Jurassic (Callovian) La Voulte-sur-Rhône  France A member of Polychelida

Personadorippe[55]

Gen. et 2 sp. et comb. nov

In press

Van Bakel et al.

Cretaceous (Aptian to Cenomanian)

Hudspeth Formation
Kubina Formation
Lyaminsk Formation

 Russia
 United States

A crab belonging to the superfamily Dorippoidea and the family Telamonocarcinidae. The type species is P. kalashnikovi; genus also includes new species P. levashiensis, as well as "Eodorippe" connori Nyborg et al. (2019).

Petrolisthes landsendi[70] Sp. nov Valid Garassino & Nyborg in Garassino et al. Late Cretaceous (Santonian) Nanaimo Group  Canada A species of Petrolisthes.
?Proeryon erinaceus[71] Sp. nov Valid Audo, Schweigert & Charbonnier Late Jurassic (Oxfordian)  France
?Proeryon quiltyi[71] Sp. nov Valid Audo, Schweigert & Charbonnier Middle Jurassic (Callovian) Antarctica
Prosopon josephcollinsi[40] Sp. nov Valid Klompmaker et al. Late Jurassic (Tithonian) Ernstbrunn Limestone  Austria A crab belonging to the family Prosopidae.
Protoapseudoidus[72] Fam. et Gen. et sp. nov In press Heard et al. Early Cretaceous (Aptian) Sierra Madre  Mexico A member of Tanaidacea belonging to the group Apseudomorpha, in the new family Protoapseudoidae. Genus includes new species P. espinalensis.

Protopagurus cerebellum[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Paguridae.

Protopagurus duopupae[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Paguridae.

Pseudastacus lemovices[73] Sp. nov Valid Charbonnier & Audo Early Jurassic (Sinemurian)  France A member of Astacidea belonging to the family Stenochiridae
Sabellidromites lanae[74] Sp. nov Valid Nyborg, Garassino & Ross Late Cretaceous (late Campanian)  Canada A dromioid crab
Stenodactylina beardi[75] Sp. nov Valid Feldmann, Schweitzer & Haggart Late Cretaceous Haslam  Canada A member of the family Erymidae.
Stenodactylina devillezi[76] Sp. nov Valid Schweigert & Härer Late Jurassic (Kimmeridgian) Nusplingen  Germany A member of the family Erymidae.
Stenodactylina geigerae[76] Sp. nov Valid Schweigert & Härer Late Jurassic (Tithonian) Usseltal  Germany A member of the family Erymidae.
Tanidromites nightwishorum[40] Sp. nov Valid Klompmaker et al. Late Jurassic (Tithonian) Ernstbrunn Limestone  Austria A crab belonging to the family Tanidromitidae.
Tanidromites weinschenki[40] Sp. nov Valid Klompmaker et al. Late Jurassic (Kimmeridgian) Oberjura-Massenkalk Formation  Germany A crab belonging to the family Tanidromitidae.

Tymolus bottemilleri[77]

Sp. nov

Valid

Nyborg & Garassino

Miocene

Astoria

 United States

A crab belonging to the family Cyclodorippidae.

Tymolus collinsi[77]

Sp. nov

Valid

Nyborg & Garassino

Miocene

 United States

A crab belonging to the family Cyclodorippidae.

Upogebia azorensis[78]

Sp. nov

Valid

Hyžný in Hyžný et al.

Pliocene

 Azores

A species of Upogebia. Announced in 2020; the final version of the article naming it was published in 2021.

Vecticallichirus kawanoi[79]

Sp. nov

Valid

Ando

Late Eocene

Funazu

 Japan

A ghost shrimp belonging to the family Callichiridae.

Vectis collinsi[55]

Sp. nov

In press

Van Bakel et al.

Early Cretaceous (Aptian)

Kubina Formation

 Russia

A crab belonging to the family Viaiidae.

Ostracods

Name Novelty Status Authors Age Type locality Country Notes Images
Adelphobolbina praeclara[80] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
Ampuloides beckeri[81] Sp. nov Valid Nazik et al. Late Devonian  Mongolia A member of Podocopida belonging to the superfamily Bairdiocypridoidea and the family Pachydomellidae.
Angarallina[82] Gen. et 2 sp. nov Valid Melnikova Ordovician  Russia Genus includes new species A. demissa and A. aenigma.
Araxobairdia[83] Gen. et sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Podocopida belonging to the family Bairdiidae. The type species is A. formosa.
Bairdia andrecrasquini[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
Bairdia gambaneraensis[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
Bairdia paramolesta[85] Sp. nov Valid Cabral, Lord & Pinto in Cabral et al. Early Jurassic (Pliensbachian and Toarcian)  Portugal A member of the family Bairdiidae.
Bairdiacypris argonautaii[86] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian) Kössen Formation  Austria
 Romania
A member of Podocopida belonging to the family Bairdiidae.
Bairdiacypris kathleenae[83] Sp. nov Valid Gliwa in Gliwa et al. Late Permian and Early Triassic (Changhsingian–Griesbachian)  Iran A member of Podocopida belonging to the family Bairdiidae.
Beyrichiopsis hushootensis[81] Sp. nov Valid Nazik et al. Late Devonian  Mongolia A member of Platycopida belonging to the superfamily Kloedenelloidea and the family Beyrichiopsidae.
Bythocypris dorsogibba[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
Bythocypris? multagracilis[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
Bythocypris? pythagorasi[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Aptian) Orbata  Tunisia
Candonopsis carthaginensis[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
Capsacythere baldanzae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere fekihi[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere giovannae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere salaji[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Carinaknightina hofmanni[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Palaeocopida belonging to the family Kirkbyidae.
Cavellina fosteri[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Platycopida belonging to the family Cavellinidae.
Cavellina hairapetiani[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Platycopida belonging to the family Cavellinidae.
Ceratobairdia? akhilleus[86] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Podocopida belonging to the family Bairdiidae.
Chrysocythere arutai[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Chrysocythere gliozziae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Chrysocythere russoi[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Cimbaurila maamourii[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Costa ponticulocarinata[89] Sp. nov Valid Yasuhara et al. Early Miocene  India
Costoprimites multicostatus[90] Sp. nov Valid Gonta Ordovician  Russia
Costoprimites processus[90] Sp. nov Valid Gonta Ordovician  Russia
Cypridea punctacentralis[91] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
Cytherella ilariae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A species of Cytherella.
Cytheretta mariaantoniettae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Cytherettidae.
Dicerobairdia buseri[92] Sp. nov Valid Forel in Forel, Kolar-Jurkovšek & Jurkovšek Late Triassic (Carnian)  Slovenia A member of Podocopida belonging to the family Bairdiidae.
Direigina[82] Gen. et sp. nov Valid Melnikova Ordovician  Russia Genus includes new species D. nana.
Disulcina proxima[82] Sp. nov Valid Melnikova Ordovician  Russia A member of the family Tetradellidae.
Dolocythere amphistiela[93] Sp. nov Valid Lord, Cabral & Danielopol Early Cretaceous (Albian)  Germany A member of Podocopida belonging to the family Cytheridae
Dolocytheridea donzei[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
Dorsogibbella alium[90] Sp. nov Valid Gonta Ordovician  Russia
Dorukella crasquinae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Dorukella nevioi[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Dorukella razgallahae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Dorukella ruggeroi[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Eumiraculum mettei[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Wuchiapingian–Changhsingian)  Iran A member of Podocopida belonging to the superfamily Bairdioidea.
Fabalicypris veronicae[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Podocopida belonging to the family Bairdiidae.
Foramenella sibirica[82] Sp. nov Valid Melnikova Ordovician  Russia
Graptocythere octopus[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Hallatina oblonga[82] Sp. nov Valid Melnikova Ordovician  Russia A member of the group Leperditellocopida belonging to the family Primitiidae.
Hastacypris[91] Gen. et sp. nov Junior homonym Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil Genus includes new species H. adamantem. The generic name is preoccupied by Hastacypris Croneis & Gutke (1939).
Heterocypris aspirasensis[94] Sp. nov Valid Durak, Akkiraz & Nazik Early Miocene Aspiras Basin  Turkey A species of Heterocypris. Announced in 2020; the final version of the article naming it was published in 2021.
Histriabairdia[86] Gen. et sp. et comb. nov Valid Forel in Forel & Grădinaru Late Triassic (Carnian) to Middle Jurassic (Bathonian)  Austria
 France
 Germany
 India
 Italy
 Romania
  Switzerland
A member of Podocopida belonging to the family Bairdiidae. The type species is H. pontuseuxinusensis; genus also includes "Anchistrocheles"? spinosa Sheppard in Brand (1990), "Anchistrocheles"? tuningensis Beher (2004) and "Anchistrocheles" gemmellaroi Crasquin in Crasquin et al. (2018).
Hungarella forelae[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Healdiidae.
Hungarella siciliiensis[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Healdiidae.
Hungaroleberis striatus[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Myodocopida belonging to the family Cylindroleberididae.
Illativella triquetra[80] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
Ilyocypris? arca[91] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
Ilyocypris? latanodi[91] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
Isobythocypris atalantella[86] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Podocopida belonging to the family Bairdiidae.
Jonesites chunensis[90] Sp. nov Valid Gonta Ordovician  Russia
Kerocythere dittainoensis[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Cytheruridae.
Kiesowia (?) incerta[82] Sp. nov Valid Melnikova Ordovician  Russia A member of the family Ctenonotellidae.
Kinkelinella ventrocarinata[85] Sp. nov Valid Cabral, Pinto & Lord in Cabral et al. Early Jurassic (Toarcian)  Portugal
 United Kingdom
A member of the family Protocytheridae.
Kinnekullea gaia[95] Sp. nov Valid Hearing et al. Ordovician (Katian) Phu Ngu  Vietnam
Korolyukina[82] Gen. et 2 sp. nov Valid Melnikova Ordovician  Russia A member of the family Bolliidae. Genus includes new species K. conspicua and K. taimyrica.
Laccochilina (?) jugata[82] Sp. nov Valid Melnikova Ordovician  Russia A member of the family Eurychilinidae.
Laccochilina (Prochilina) yanpingensis[96] Sp. nov In press Zhang Ordovician Shihtzupu Formation  China
Leucocytherella dangeloi[97] Sp. nov Valid Alivernini in Alivernini et al. Miocene (Messinian) and Pliocene Zhada Basin  China A member of the family Limnocytheridae.
Liuzhinia julfensis[83] Sp. nov Valid Gliwa in Gliwa et al. Late Permian and Early Triassic (Changhsingian–Griesbachian)  Iran A member of Podocopida belonging to the superfamily Bairdioidea.
Lyumellina[90] Gen. et sp. nov Valid Gonta Ordovician  Russia Genus includes new species L. risus
Minyocythere[93] Gen. et 2 sp. et comb. nov Valid Lord, Cabral & Danielopol Middle Jurassic (Aalenian and Bajocian)  Germany

 Netherlands
 United Kingdom

A cytherid Podocopida. The type species is M. macroporosa;
genus also includes new species M. angulata,
as well as "Dolocythere" maculosa Bate (1963) and "Dolocythere" tuberculata Luppold (2012).
Mockella barbroae[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Limnocytheridae.
Mongolianella kyranbeki[98] Sp. nov In press Choi et al. Cretaceous Yanji  China
Myanmarcypris[99] Gen. et sp. nov Valid Wang et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of the family Candonidae belonging to the subfamily Paracypridinae. Genus includes new species M. hui.
Nikolina entonipteros[90] Sp. nov Valid Gonta Ordovician  Russia
Nikolina pteroventralis[90] Sp. nov Valid Gonta Ordovician  Russia
Okadaleberis (?) benzartiae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis decimai[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis guerneti[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis hendae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis khayatii[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis memmiae[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Orthobairdia capuliformis[83] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Podocopida belonging to the family Bairdiidae.
Paijenborchellina reitanoi[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Cytheridae.
Paractinocythereis[89] Gen. et comb. nov Valid Yasuhara et al. Early Miocene  India A new genus for "Actinocythereis" gujaratensis Tewari & Tandon (1960)
Paracypris chekhmai[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Aptian) Orbata  Tunisia
Paracypris ovidi[86] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Cypridoidea belonging to the family Paracyprididae.
Perissocytheridea altesulcata[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Aptian) Orbata  Tunisia
Perissocytheridea kouminiensis[87] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
Petasobairdia amazonella[86] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of the family Bairdiidae.
Petasobairdia jeandercourti[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
Pontocypris? karavankensis[92] Sp. nov Valid Forel in Forel, Kolar-Jurkovšek & Jurkovšek Late Triassic (Carnian)  Slovenia A member of Cypridoidea belonging to the family Pontocyprididae.
Postsinusella[90] Gen. et sp. nov Valid Gonta Ordovician  Russia Genus includes new species P. guttaformis.
Prasuchonella? huarpe[100] Sp. nov Valid Carignano, Echevarría & Zavattieri Middle Triassic Cuyo Basin  Argentina A member of Darwinulocopina.
Pseudoleperditia? vetusta[80] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
Pseudomacrocypris? kerabani[86] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Cypridoidea belonging to the family Pontocyprididae.
Pseudulrichia replicata[82] Sp. nov Valid Melnikova Ordovician  Russia A member of the family Richinidae.
Ptychobairdia iudicaensis[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
Ptychobairdia leonardoi[84] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
Pythagoracypris[91] Gen. et sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil Genus includes new species P. latavectis.
Rhinocypris? ericius[91] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
Ruggieria quadricarinata[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Cytherettidae.
Steusloffia rudiformis[90] Sp. nov Valid Gonta Ordovician  Russia
Sulcicuneus gemmifer[80] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
Tasmanocypris lochardi[101] Sp. nov In press Warne Late Miocene and Pliocene  Australia
Tesakovites[82] Gen. et sp. nov Valid Melnikova Ordovician  Russia A member of the family Circulinidae. Genus includes new species T. rotundatus.
Urftella triangula[80] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
Urocythere (Pokornyella) salaktaensis[88] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Velatomorpha rochacamposi[102] Sp. nov Valid Bergue, Maranhao & Ng Permian  Brazil

Other crustaceans

Name Novelty Status Authors Age Type locality Country Notes Images

Actinobalanus? sloveniensis[103]

Sp. nov

Valid

Buckeridge et al.

Oligocene and Miocene

 Slovenia

A barnacle.

Altiverruca capsa[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Altiverruca fusione[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Ambocyclus[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Ireland
 Russia
 United States?

A member of Cyclida. Genus includes A. capidulum (Chernyschev, 1933), A. simulans (Reed, 1908) and possibly A. ? minutus (Rogers, 1902).

Angulotergum kiamichensis[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Angulotergum milviformis[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Angulotergum robustum[106] Sp. nov In press Gale Cretaceous  United States A barnacle.

Arcoscalpellum s.l. isaonishikawai [107]

Sp. nov

Valid

Karasawa

Miocene

Katsuta Group

 Japan

A member of the family Scalpellidae.

Arcoscalpellum s.l. Joei [107]

Sp. nov

Valid

Karasawa

Miocene

Yotsuyaku Formation

 Japan

A member of the family Scalpellidae.

Arcuatoscalpellum wacoensis[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Ashinkailepas indica[108] Sp. nov Valid Gale in Gale et al. Late Pleistocene  India A barnacle

Aurivillialepas mascarensis[109]

Sp. nov

Valid

Gale

Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Calanticidae.

Brachylepas hantonensis[110] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
Brachylepas thieli[110] Sp. nov Valid Gale Cretaceous  Germany A barnacle

Brittaniclus[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 United Kingdom

A member of Cyclida belonging to the family Americlidae. Genus includes B. rankini (Woodward, 1868), B. scotti (Woodward, 1894) and B. testudo (Peach, 1882).

Carabicyclus[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Ireland
 United Kingdom

A member of Cyclida. Genus includes C. wrighti (Woodward, 1870).

Catherinum busselli[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Scalpellidae.

Chernyshevine[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Tajikistan

A member of Cyclida. Genus includes C. spinosus (Chernyschev, 1933).

Collinslepas[110] Gen. et 3 sp. nov Valid Gale Cretaceous  Morocco
 Tunisia
A barnacle. Genus includes new species C. aitlaminensis, C. robustus and C. tunisiensis.

Costatolepas[104]

Gen. et sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Scalpellidae. The type species is C. buckeridgei.

Costatoverruca baxteri[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Costatoverruca macropluteum[109]

Sp. nov

Valid

Gale

Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Costatoverruca tredecima[109]

Sp. nov

Valid

Gale

Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Cretiscalpellum robaszynskii[111] Sp. nov In press Gale, Jagt & Goolaerts Late Cretaceous (Maastrichtian)  Tunisia A barnacle.
Daohugounaias[112] Gen. et sp. nov Valid Luo et al. Middle Jurassic Daohugou  China A member of Anostraca. Genus includes new species D. cheni.
Diotascalpellum acies[106] Sp. nov In press Gale Cretaceous  United States A barnacle.

Dziklus[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

Mazon Creek fossil beds

 United States

A member of Cyclida belonging to the family Americlidae. Genus includes D. obesus (Schram, Vonk & Hof, 1997).

Eoverruca aubensis[110] Sp. nov Valid Gale Early Cretaceous (Albian)  France A barnacle
Eoverruca symmetrica[110] Sp. nov Valid Gale Late Cretaceous (Campanian)  United Kingdom A barnacle
Etcheslepas portlandensis[113] Sp. nov Valid Gale TithonianBerriasian Portland Group  United Kingdom A barnacle
Fallaxlepas[110] Gen. et comb. nov Valid Gale Cretaceous  United Kingdom A brachylepadid barnacle. The type species is "Pollicipes" fallax Darwin (1851)

Gibbosaverruca youngi[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Iliestheria urumqiensis[114] Sp. nov Valid Teng & Li Early Jurassic (PliensbachianToarcian) Sangonghe  China A clam shrimp. Announced in 2020; the final version of the article naming was published in 2021.
Ivolepas worthensis[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Jagtscalpellum[110] Gen. et comb. et 3 sp. nov Valid Gale Cretaceous  Tunisia
 United Kingdom
 United States
A barnacle. The type species is "Pollicipes" striatus Darwin (1851); genus also includes "Pollicipes" filosus Withers (1911), as well as new species J. africanum, J. spinosum and J. luteorum.[106]
Lahuerguinagrapta[115] Gen. et sp. nov In press Gallego et al. Early Cretaceous La Huérguina  Spain An afrograptid clam shrimp. Genus includes new species L. multicostata.

Litocyclus[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Ireland
 United Kingdom
 United States?

A member of Cyclida. Genus includes L. bilobatus (Woodward, 1870), L. jonesianus (Woodward, 1870), L. torosus (Woodward, 1870) and possibly also L. ? communis (Rogers, 1902) and L. ? permarginatus (Rogers, 1902).

Lophobalanus fresvillensis[116] Sp. nov Valid Gale Eocene (Lutetian)  France A barnacle. Announced in 2020; the final version of the article naming it was published in 2021.
Loriculina fragila[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Loriculina rubrifluvia[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Loriolepas whytei[113] Sp. nov Valid Gale TithonianBerriasian  United Kingdom A barnacle

Newmaniverruca multitabulata[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Pachylasma traceyi[109]

Sp. nov

Valid

Gale

Pliocene-Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Pachylasmatidae.

Proverruca anglica[110] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
Regioscalpellum kennedyi[110] Sp. nov Valid Gale Cretaceous  Tunisia A barnacle

Rostratoverruca darwini[104]

Sp. nov

Valid

Gale

Pliocene-early Pleistocene

Indian Ocean (Mascarene Plateau)

A barnacle belonging to the family Verrucidae.

Striascalpellum[110] Gen. et 3 sp. et comb. nov Valid Gale Cretaceous  Sweden
 United Kingdom
A barnacle. The type species is S. elegans; genus also includes new species S. bromleyi and S. barringtonensis, as well as "Scalpellum (Titanolepas)" martini Withers (1935).
Subsecolepas[110] Gen. et sp. nov Valid Gale Cretaceous  United Kingdom A barnacle. The type species is S. holtwilsoni.

Tazawacyclus[105]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Japan

A member of Cyclida. Genus includes T. tazawai (Niko & Ibaraki, 2011).

Texaslepas[106] Gen. et sp. nov In press Gale Cretaceous  United States A barnacle. Genus includes new species T. holterhoffi.
Vectibalanus[116] Gen. et comb. et sp. nov Valid Gale Eocene and Oligocene Barton
Bouldnor
Headon Hill
Selsey
 United Kingdom A barnacle. The type species is "Balanus" unguiformis Sowerby (1846); genus also includes "Balanus" erisma Sowerby (1846) and a new species V. mortoni. Announced in 2020; the final version of the article naming it was published in 2021.
Virgilepas[110] Gen. et 3 sp. nov Valid Gale Cretaceous  Morocco
 United Kingdom
A barnacle. Genus includes new species V. aboudaensis, V. peakei and V. hancocki.
Virgiscalpellum laevis[110] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
Virgiscalpellum marysensis[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Virgiscalpellum mhrilensis[110] Sp. nov Valid Gale Cretaceous  Tunisia A barnacle
Virgiscalpellum multilineatum[110] Sp. nov Valid Gale Cretaceous  Tunisia A barnacle
Virgiscalpellum rugosum[106] Sp. nov In press Gale Cretaceous  United States A barnacle.
Virgiscalpellum sussexiense[110] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
Virgiscalpellum truncatum[110] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
Witherscalpellum[110] Gen. et comb. nov Valid Gale Cretaceous  France

 United Kingdom

A barnacle. The type species is "Cretiscalpellum aptiensis Withers (1935); genus also includes "Cretiscalpellum" matrioni Gale (2019).

General research

Insects

Trilobites

New taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Acastoides biebrichensis[124]

Sp. nov

Valid

Alberti

Devonian (Emsian)

Rupbach Shale

 Germany

A member of the family Acastidae.

Acastoides requadti[124]

Sp. nov

Valid

Alberti

Devonian (Emsian)

Rupbach Shale

 Germany

A member of the family Acastidae.

Aciculolenus askewi[125]

Sp. nov

Valid

Chatterton

Cambrian (Jiangshanian)

 Canada

Arduennella hainauensis[126]

Sp. nov

Valid

Van Viersen & Alberti

Devonian (Emsian)

 Germany

Atlanticalymene[127] Gen. et sp. nov Valid Adrain, Karim & McAdams Ordovician (Darriwilian) Table Cove Formation  Canada A calymenine trilobite. The type species is A. bardensis.
Balvibole[128] Gen. et sp. nov Valid Basse & Lemke Devonian (Famennian) Dasberg Limestone  Germany A cyrtosymboline trilobite. Genus includes new species B. kaufmanni.
Blountia angelae[129] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Nolichucky Formation  United States A member of the family Kingstoniidae.
Blountia morgancreekensis[129] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Riley Formation  United States A member of the family Kingstoniidae.
Blountia nevadensis[129] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Mendha Formation  United States A member of the family Kingstoniidae.
Blountia newfoundlandensis[129] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Shallow Bay Formation  Canada A member of the family Kingstoniidae.
Blountia tennesseensis[129] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Nolichucky Formation  United States A member of the family Kingstoniidae.

Boethiusia[130]

Gen. et comb. nov

Valid

Geyer

Cambrian

 Italy

A new genus for "Dolerolenus" courtessolei Pillola (1991).

Clappaspis tupeq[131]

Sp. nov

Valid

Peel

Cambrian (Miaolingian)

Telt Bugt Formation

 Greenland

A member of the family Alokistocaridae.

Cliffia nicoleae[125]

Sp. nov

Valid

Chatterton

Cambrian (Jiangshanian)

 Canada

Cyphaspides paulospinatus[132]

Sp. nov

Valid

Alberti

Devonian

 Germany

A member of Proetida belonging to the family Aulacopleuridae.

Cyphaspides speculator[132]

Sp. nov

Valid

Alberti

Devonian

 Germany

A member of Proetida belonging to the family Aulacopleuridae.

Dicranurus webbyi[133]

Sp. nov

Valid

Holloway, Smith & Thomas

Ordovician (Katian)

Gunningbland Formation

 Australia

An odontopleurid trilobite.

Ehmaniella sermersuaqensis[131]

Sp. nov

Valid

Peel

Cambrian (Miaolingian)

Telt Bugt Formation

 Greenland

A member of the family Alokistocaridae.

Elrathia hensonensis[134]

Nom. nov

Valid

Geyer & Peel

Cambrian

Ekspedition Bræ Formation

 Greenland

A replacement name for Elrathia groenlandica Geyer & Peel (2017).

Eowinterbergia effenbergensis[128] Sp. nov Valid Basse & Lemke Devonian (Famennian) Wocklum Limestone  Germany

Epicurella[130]

Gen. et sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of the family Despujolsiidae. Genus includes new species E. pustulata.

Feistops[135]

Gen. et 2 sp. nov

Valid

Crônier et al.

Devonian (Famennian)

 Mongolia

A phacopid trilobite. Genus includes new species F. mongoliensis and F. khovdensis. Announced in 2020; the final version of the article naming was published in 2021.

Gravicalymene bakeri[136]

Sp. nov

In press

Smith & Ebach

Ordovician (Katian)

Gordon Group

 Australia

A calymenid trilobite.

?Houseops olonbulagensis[135]

Sp. nov

Valid

Crônier et al.

Devonian (Famennian)

 Mongolia

A phacopid trilobite. Announced in 2020; the final version of the article naming was published in 2021.

Kingaspidoides spinirecurvatus[137]

Sp. nov

Valid

Geyer, Pais & Wotte

Cambrian

Jbel Wawrmast Formation

 Morocco

A member of Redlichiida belonging to the family Ellipsocephalidae.

Marsaisia devoillei[130]

Sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of the family Despujolsiidae.

Marsaisia? lemdadensis[130]

Sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of the family Despujolsiidae.

Phantaspis[138]

Gen. et sp. nov

Valid

Sun, Zeng & Zhao

Cambrian (Wuliuan)

Mantou Formation

 China

A member of Ptychopariacea of uncertain phylogenetic placement. The type species is P. auritus.

Proabadiella[139]

Gen. et 2 sp. nov

Valid

Liñán et al.

Cambrian Stage 3

Pusa Formation

 Spain

Genus includes new species P. toletana and P. vidalii.

Prophalaron[133]

Gen. et sp. nov

Valid

Holloway, Smith & Thomas

Ordovician (Katian)

Gunningbland Formation

 Australia

A calymenid trilobite. Genus includes new species P. jonesi.

Prosaukia oculata[140]

Sp. nov

Valid

Wernette et al.

Cambrian (Furongian)

Ao Mo Lae Formation

 Thailand

Satunarcus[141]

Gen. et sp. nov

Valid

Wernette et al.

Late Cambrian

Ao Mo Lae Formation

 Thailand

A kaolishaniid trilobite belonging to the new subfamily Ceronocarinae. Genus includes new species S. molaensis.

Saukianda? dresnayi[142]

Sp. nov

In press

Geyer & Landing

Cambrian

MoroccanAlgerian border region

Treveropyge hellemondi[143]

Sp. nov

Valid

Van Viersen & Taghon

Devonian (Emsian)

Our Formation

 Belgium

A member of the family Acastidae.

Utia debra[144]

Sp. nov

Valid

Sundberg

Cambrian (Wuliuan)

Lakeview Limestone
Spence Shale

 United States

A member of Ptychopariida belonging to the family Utiidae.

General research

  • A study on the timing of the trilobite extinctions at the early to middle Cambrian transition is published by Sundberg et al. (2020).[145]
  • Evidence of gregarious moulting of Cambrian (Stage 4) trilobites from the "Tsinghsutung" Formation (China), representing one of the earliest records of synchronized moulting behaviours in the fossil record, is presented by Corrales-García et al. (2020).[146]
  • A study on trilobite size variations in the Ordovician Fezouata Formation (Morocco) deposited in a cold-water environment is published by Saleh et al. (2020).[147]
  • A study on the nature of variation among oryctocephalid specimens from the Combined Metals Member of the Pioche Formation (Nevada; Cambrian Stage 4) will be published by Webster & Sundberg (2020).[148]
  • A study on the ontogeny of the olenid trilobite Leptoplastides salteri is published by Månsson & Clarkson (2020).[149]
  • A study on the anatomy of the internal structures of the compound eye of Aulacopleura koninckii is published by Schoenemann & Clarkson (2020).[150]
  • A study on exuviae of Ovalocephalus tetrasulcatus from the Ordovician Linhsiang Formation (Hubei, China) is published by Zong (2020), who reports exuviae arranged with two or three together end to end or superimposed, and evaluates the implications of these findings for the knowledge of the behavior of trilobites.[151]

Other arthropods

Newly named taxa

Name Novelty Status Authors Age Type locality Country Notes Images

Adelophthalmus pyrrhae[152]

Sp. nov

Valid

Lamsdell et al.

Carboniferous (Tournaisian)

St. Nazaire Group (possibly Lydiennes Formation)

 France

A eurypterid.

Allolimulus[153]

Gen. et comb. nov

Disputed

Lamsdell

Jurassic

 United Kingdom

A horseshoe crab. The type species is "Limulus" woodwardi Watson (1909). However, Bicknell et al. (2021) transferred "L." woodwardi to the genus Mesolimulus.[154]

Andersoniella[153]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 Germany
 United States

A member of the family Belinuridae. The type species is "Euproops" longispina Packard (1885).

Batracholimulus[153]

Gen. et comb. nov

Valid

Lamsdell

Triassic

 Germany

A member of the family Austrolimulidae. The type species is "Paleolimulus" fuchsbergensis Hauschke & Wilde (1987).

Boeotiaspis[153]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 United States

A member of the family Austrolimulidae. The type species is "Paleolimulus" longispinus Schram (1979).

Bushizheia[155] Gen. et sp. nov Valid O’Flynn & Liu in O’Flynn et al. Cambrian Stage 3 Chiungchussu  China An early euarthropod of uncertain phylogenetic placement. The type species is B. yangi.

Concavicaris martinae[156]

Sp. nov

Valid

Broda, Rak & Hegna

Devonian (Famennian)

 Czech Republic

A thylacocephalan.

Concavicaris pikae[156]

Sp. nov

Valid

Broda, Rak & Hegna

Devonian (Famennian)

 Poland

A thylacocephalan.

Concavicaris submarinus[157]

Sp. nov

Jobbins, Haug & Klug

Devonian (Famennian)

 Morocco

A thylacocephalan.

Concavicaris woodruffensis[156]

Sp. nov

Valid

Broda, Rak & Hegna

Devonian (Famennian)

Woodruff Formation

 United States

A thylacocephalan.

Devonopilio[158]

Gen. et sp. nov

Valid

Tihelka, Tian & Cai

Devonian (Pragian)

Rhynie chert

 United Kingdom

Originally described as a member of Opiliones, but this interpretation was subsequently questioned by Pérez-González & Shultz (2021), who were unsure about the phylogenetic placement of this arthropod, but noted anatomical similarities with the euthycarcinoid species Heterocrania rhyniensis.[159] The type species is D. hutchinsoni.

Diplacanthocaris[160]

Gen. et sp. nov

Valid

Ji et al.

Early Triassic

 China

A thylacocephalan. Genus includes new species D. chaohuensis.

Dolesea[161]

Gen. et sp. nov

Valid

Hannibal & May

Permian (Cisuralian)

Richards Spur locality

 United States
( Oklahoma)

A millipede belonging to the group Helminthomorpha. The type species is D. subtila.

Ericixerxes[162] Gen. et sp. nov Gueriau et al. Devonian (Famennian) Evieux  Belgium A member of Euthycarcinoidea. The type species is E. potii.
Hipponicharion perforata[163] Sp. nov Valid Skovsted et al. Cambrian Mural  Canada A member of Bradoriida belonging to the family Hipponicharionidae.

Itagnostus idahoensis[144]

Sp. nov

Valid

Sundberg

Cambrian (Wuliuan)

Emigrant Formation
Lakeview Limestone

 United States

A member of Agnostida belonging to the family Peronopsidae.

Kanoshoia[164]

Gen. et sp. nov

Valid

Lerosey‐Aubril et al.

Cambrian (Drumian)

Wheeler Shale

 United States

A megacheiran. Genus includes new species K. rectifrons.

Karstiulus[161]

Gen. et sp. nov

Valid

Hannibal & May

Permian (Cisuralian)

Richards Spur locality

 United States
( Oklahoma)

A millipede belonging to the group Juliformia, possibly a member of the superfamily Xyloiuloidea. The type species is K. fortsillensis.

Keuperlimulus[153]

Gen. et comb. nov

Valid

Lamsdell

Triassic

 France

A horseshoe crab. The type species is "Limulus" vicensis Bleicher (1897).

Macrobelinurus[153]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 United Kingdom

A member of the family Belinuridae. The type species is "Steropis" arcuatus Baily (1859).

Mesolimulus tafraoutensis[165]

Sp. nov

In press

Lamsdell et al.

Late Cretaceous (CenomanianTuronian)

Gara Sbaa Lagerstätte

 Morocco

A horseshoe crab.

Microcaris rectilineatus[160]

Sp. nov

Valid

Ji et al.

Early Triassic

 China

A thylacocephalan.

Norilimulus[153]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous (Tournaisian)

Horton Bluff Formation

 Canada

A member of Xiphosura belonging to the family Paleolimulidae. The type species is "Paleolimulus" woodae Lerner, Lucas & Mansky (2016).

Oklahomasoma[161]

Gen. et sp. nov

Valid

Hannibal & May

Permian (Cisuralian)

Richards Spur locality

 United States
( Oklahoma)

A millipede belonging to the group Helminthomorpha. The type species is O. richardsspurense.

Parabelinurus[153]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 Russia
 Ukraine
 United States

A member of the family Belinuridae. The type species is "Enthomolithus" lunatus Martin (1809); genus also includes P. iswariensis (Chernyshev, 1928), P. lacoei (Packard, 1885), P. metschetensis (Chernyshev, 1928) and P. stepanovi (Chernyshev, 1928).

Paraconcavicaris[156]

Gen. et comb. nov

Valid

Broda, Rak & Hegna

Carboniferous (Tournaisian)

Líšeň Formation

 Czech Republic

A thylacocephalan; a new genus for "Concavicaris" viktoryni Rak, Broda & Kumpan (2018).

Parioscorpio[166]

Gen. et sp. nov

Wendruff et al.

Silurian (Telychian)

Brandon Bridge

 United States

An arthropod of uncertain phylogenetic placement. Originally classified as an early scorpion, but subsequently argued to be basal to crown group Mandibulata and Chelicerata.[167] The type species is P. venator.

Patesia[168]

Gen. et comb. nov

Valid

Bicknell & Smith

Late Devonian

 United States

A stem xiphosurid; a new genus for "Kasibelinurus" randalli. Announced in 2020; the final version of the article naming it was published in 2021.

Pauropsxenus extraneus[169]

Sp. nov

In press

Su, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede belonging to the family Polyxenidae.

Pauropsxenus ordinatus[169]

Sp. nov

In press

Su, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede belonging to the family Polyxenidae.

Pruemopterus[170]

Gen. et sp. nov

Valid

Poschmann

Early Devonian

 Germany

A eurypterid. Genus includes new species P. salgadoi. Announced in 2020; the final version of the article naming it was published in 2021.

Pseudobeyrichona taurata[163] Sp. nov Valid Skovsted et al. Cambrian Mural  Canada A member of Bradoriida belonging to the family Hipponicharionidae.

Shpineviolimulus[171]

Gen. et comb. nov

Bicknell, Naugolnykh & Brougham

Permian (Asselian)

Araukaritovaya Formation

 Ukraine

A member of Xiphosura belonging to the group Limulina; a new genus for "Paleolimulus" jakovlevi Glushenko & Ivanov (1961).

Sklerolibyon[172]

Gen. et sp. nov

Aria et al.

Cambrian Stage 3

Maotianshan Shale Member of the Yu’anshan Formation

 China

A megacheiran related to Jianfengia and Fortiforceps. The type species is S. maomima.

Soligorskopterus shpinevi[173]

Sp. nov

Valid

Naugolnykh & Areshin

Late Devonian

 Russia

An eurypterid. Announced in 2019; the final version of the article naming it was published in 2020.

Tariccoia tazagurtensis[174] Sp. nov Valid Pérez-Peris et al. Early Ordovician Fezouata  Morocco A nektaspid arthropod. Announced in 2020; the final version of the article naming it was published in 2021.

Thelxiope holmani[175]

Sp. nov

Valid

Lerosey-Aubril, Skabelund & Ortega-Hernández

Cambrian (Drumian)

Wheeler Shale

 United States

A possible member of Chelicerata.

Volanalimulus[153]

Gen. et sp. nov

Valid

Lamsdell

Early Triassic

Ankitokazo Basin

 Madagascar

A horseshoe crab. The type species is V. madagascarensis.

Xiaocaris[176]

Gen. et sp. nov

Liu et al.

Cambrian Stage 3

Chiungchussu Formation

 China

A member of the group Fuxianhuiida. The type species is X. luoi.

Xiazhuangocaris[177]

Gen. et sp. nov

Valid

Zeng et al.

Cambrian Stage 3

Hongjingshao Formation

 China

A member of Hymenocarina. Genus includes new species X. chenggongensis. Announced in 2020; the final version of the article naming it was published in 2021.

General research

  • Exceptionally preserved specimens of Chuandianella ovata carrying a large number of small eggs are described from the Cambrian Chengjiang biota (China) by Ou et al. (2020), who compare the reproductive strategies of C. ovata and closely related Waptia fieldensis.[178]
  • Sun, Zeng & Zhao (2020) report the discovery of a member of the genus Sidneyia belonging or related to the species S. inexpectans from the Wuliuan Mantou Formation of North China, expanding known distribution of this genus.[179]
  • A putative xiphosuran Kiaeria is reinterpreted as a member of Chasmataspidida by Lamsdell (2020a).[180]
  • A study on the anatomy of head structures of Heterocrania rhyniensis, evaluating its implications for the knowledge of the phylogenetic relationships of euthycarcinoids, is published by Edgecombe et al. (2020).[181]
  • A study on heterochrony in Xiphosura, showing that clades that occupy non-marine environments show elevated rates of peramorphosis or paedomorphosis is published by Lamsdell (2020b).[182]
  • A study on the morphology of the feeding apparatuses of Eurypterida and Trigonotarbida is published by Haug (2020).[183]
  • A study aiming to determine the range of prey sizes that could be captured by sweep-feeding eurypterids is published by Hughes & Lamsdell (2020).[184]
  • A study on the anatomy and phylogenetic relationships of Anhelkocephalon handlirschi is published by Schädel & Haug (2020), who interpret this taxon as a probable member of Cyclida rather than an isopod.[185]
  • Liu et al. (2020) report evidence of the presence of a three-dimensionally preserved labrum associated with the mouth opening in juvenile specimens of Leanchoilia illecebrosa.[186]

References

  1. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af Jörg Wunderlich; Patrick Müller (2020). "New and already described fossil spiders (Araneae) of 20 families in mid and Late Cretaceous Burmese ambers, with notes on spider phylogeny, evolution and classification" (PDF). In Jörg Wunderlich (ed.). Beiträge zur Araneologie, 13. pp. 22–164. ISBN 978-3-931473-19-8.
  2. ^ a b c d e f Christian Bartel; Jason A. Dunlop; Prashant P. Sharma; Paul A. Selden; Dong Ren; Chungkun Shih (2020). "Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber". Cretaceous Research. 119: Article 104703. doi:10.1016/j.cretres.2020.104703. S2CID 228911877.
  3. ^ Xueying Hu; Xiaojie Lei; Cihang Luo; E.A. Jarzembowski; Bo Wang; Chuantao Xiao (2020). "A new whip spider (Arachnida: Amblypygi) in mid-Cretaceous Kachin amber". Cretaceous Research. 116: Article 104596. doi:10.1016/j.cretres.2020.104596. S2CID 225379275.
  4. ^ Jörg Wunderlich; Patrick Müller (2021). "Description of new fossil spiders (Araneae) in Late (mid) Cretaceous Burmese (Kachin) amber with focus on the superfamilies Palpimanoidea and Deinopoidea and members of the RTA-clade, as well as remarks on palaeobehaviour, palaeofauna, taxonomy and phylogenetics" (PDF). In Jörg Wunderlich (ed.). Beiträge zur Araneologie, 14. pp. 25–262. OCLC 1249433671. ISBN 978-3-931473-20-1
  5. ^ a b Matthew R. Downen; Paul A. Selden (2020). "Fossil spiders (Araneae) from the Eocene Kishenehn Formation of Montana, USA". Palaeontologia Electronica. 23 (3): Article number 23(3):a56. doi:10.26879/1135. S2CID 229652618.
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