Aistopoda

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Aïstopods
Temporal range: Early Carboniferous - Early Permian
Ophiderpeton BW.jpg
Restoration of Ophiderpeton
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: "Amphibia" (wide sense)
Subclass: Lepospondyli
Node: Holospondyli
Order: Aistopoda
Miall, 1875

Aïstopoda (Greek for "[having] not-visible feet") is an order of highly specialised snake-like amphibians known from the Carboniferous and Early Permian of Europe and North America, ranging from tiny forms only 5 centimetres (2 in), to nearly 1 metre (3.3 ft) in length. The first appear in the fossil record in the Mississippian period and continue through to the Early Permian.

The skull is small but very specialised, with large orbits, and large fenestrae. The bones at the back of the skull were reduced or absent. The primitive form Ophiderpeton has a pattern of dermal bones in the skull similar in respects to the temnospondyls. But in the advanced genus Phlegethontia the skull is very light and open, reduced to a series of struts supporting the braincase against the lower jaw, just as in snakes, and it is possible that the Aïstopods filled the same ecological niches in the Paleozoic that snakes do today.

They had an extremely elongated body, with up to 230 vertebrae. The vertebrae were holospondylous, having only a single ossification per segment. They lacked intercentra, even in the tail, and had are no free haemal arches. The neural arch was low and fused to the centrum. All of these features are very similar to those of the Nectridea, both representing the typical lepospndylous condition.

The ribs were slender, either single or double-headed, with the head shaped like a letter "K". There is no trace of limbs or even limb girdles in any known fossil, and the tail was short and primitive.

Relationships[edit]

Evolutionary relationships with other early tetrapods remain controversial, as even the earliest aïstopod, the Viséan species Lethiscus stocki, was already highly specialised. Aïstopods have been variously grouped with other lepospondyls, or placed at or prior to the batrachomorph-reptiliomorph divide. The group was quite diverse during the Late Carboniferous, with a few forms continuing through to the Permian.

Below is a cladogram from Anderson et al. (2003) showing the phylogenetic relationships of aistopods:[1]

Lepospondyli


 Asaphestera




 Scincosaurus



 Brachydectes




Aïstopoda

 Lethiscus




 Ophiderpeton 



Oestocephalidae

 Oestocephalus



 Coloraderpeton


 

Phlegethontioidea
Pseudophlegethontiidae

 Pseudophlegethontia



Phlegethontiidae

 Phlegethontia longissima



 Phlegethontia linearis








References[edit]

  • Benton, M. J. (2000), Vertebrate Paleontology, 2nd ed. Blackwell Science Ltd
  • Carroll, RL (1988), Vertebrate Paleontology and Evolution, WH Freeman & Co. pp. 176–7
  • Reisz, Robert Biology 356 - Major Features of Vertebrate Evolution (online)
  • von Zittel, K.A (1932), Textbook of Paleontology, CR Eastman (transl. and ed), 2nd edition, vol.2, p. 221-2, Macmillan & Co.

External links[edit]