Alexander Archipelago wolf
|Alexander Archipelago wolf|
|Subspecies:||C. l. ligoni|
|Canis lupus ligoni
The Alexander Archipelago wolf (Canis lupus ligoni), also known as the Archipelago wolf and the Islands wolf, is a subspecies of the gray wolf, Canis lupus, and primarily resides in the areas in and around the Alexander Archipelago. This region composes a part of the Tongass National Forest, where this species makes its home. There have been multiple attempts to have the Alexander Archipelago wolf listed as threatened on the Endangered Species List, but they have all failed.
- 1 Physiology
- 2 Range, population and repopulation
- 3 Taxonomy
- 4 History – managerial & political
- 5 References
- 6 External links
Typically smaller than the other Alaskan subspecies of wolf, the Alexander Archipelago wolf averages between 30 and 50 pounds (14 and 23 kg). They are about 3 1⁄2 feet (1.1 m) long and 2 feet (0.61 m) high at the shoulder. Their coat is generally a dark gray, with varying patterns of lighter shades. Individuals from different islands in the archipelago have a propensity for different color phases, from pure black to combinations of black and white to a much brighter cinnamon color.
The primary prey of this species is the Sitka black-tailed deer, which it consumes more than 90% of the time. The next closest consumed species, less than 10%, is the North American beaver. Then average Alexander Archipelago wolf eats an estimated 26 deer per year. This habit of feeding almost entirely on a single species is peculiar to this wolf, and is not seen in other North American wolf species.
Range, population and repopulation
The range of the Alexander Archipelago wolf covers all of southeastern Alaska (the Alaskan panhandle) except the Admiralty, Baranof, and Chichagof Islands.
No population estimates have been made since the mid-1990s. A radio collar study done then produced a region-wide population estimate of 750 to 1,100, with the fall 1994 (before trapping season) population estimated to be 908. That study was conducted on Prince of Wales Island, and the region-wide estimate was made by an extrapolation based on the varying habitat capability for prey. The Prince of Wales population was estimated to be 300–350.
During field work in summer 2010, the Alaska Department of Fish and Game (ADFG) determined the Prince of Wales Island wolf population has recently declined sharply. ADFG was unable to collect enough wolf scats to make a population estimate based on DNA. Reportedly, "only a 'small fraction' of the expected number of scats" was found during this effort, in which a number of known denning sites were checked and transects were checked over an extensive part of the island. In a regulatory proposal for the Alaska Board of Game's November 2010 meeting to help protect the species, ADFG estimated the island's wolf population to be 150, down by half or more from the 300–350 for the island determined by the 1990s radio-collar study.
In southeast Alaska, pups are usually born during the last two weeks of April. Dens are usually built four to five weeks prior to the birth, between the roots of trees, in small caves or crevices in rocks, abandoned beaver lodges, or expanded mammal burrows.
Early taxonomists were able to determine that the Alexander Archipelago wolf was its own unique subspecies due to "common cranial characteristics". It has been suggested more recently by taxonomists that the species may have originated from another subspecies known as Canis lupus nubilis.
History – managerial & political
The Alexander Archipelago wolf first arrived in Alaska sometime between 7,000 and 8,000 years ago, after the end of the Wisconsin glaciation period. The species was likely following the migration of the Sitka deer as they traveled north because of geographical and climate change in the area. The first observation of concern for the possible instability of the Alexander Archipelago wolf population was by a USDA Forest Service-sponsored interagency committee. This concern came about because of the extensive logging being conducted in the region's forest, under the Tongass Land Management Plan.
Endangered Species Act petition – 1993 to 1997
A petition was presented to the United States Fish and Wildlife Service (FWS) in December 1993 by the Biodiversity Legal Foundation and an independent biologist, requesting the Alexander Archipelago wolf to be listed as a threatened species under the Endangered_Species_Act. The agency published a positive 90-day finding in the Federal Register on May 20, 1994, but near the of the year issued another finding that a "listing is not warranted at this time", but that if the logging was not reduced or reservation areas created, the "long-term viability of the Alexander Archipelago wolf is seriously imperiled."  To better assess the status of the species, the FWS ordered a conservation assessment to be made in terms of specific data of the species and its viability for the future.
After the assessment was completed, more studies were undertaken to understand exactly how the Alexander Archipelago wolf fits into the food chain and what effect extensive logging would cause. It was surmised after study that, between 1995 and 2045, the population of the Alexander Archipelago wolf would "decline as much as 25%", along with Sitka deer population declining by 28% within the same time period.
In 1994, the FWS issued a memo stating, "not protecting the wolf would be the 'least controversial option'". This was in regards to the logging companies and lobbyists that opposed restrictions on logging in the area, which protecting the Alexander Archipelago wolf would create.
Jack Ward Thomas wrote in his book, Jack Ward Thomas: the journals of a Forest Service chief, about a meeting held in 1995 in regards to a consideration by the Forest Service to list the Alexander Archipelago wolf and the Queen Charlotte goshawk as threatened. The meeting was between Thomas, Undersecretary James Lyons, Deputy Undersecretary Adela Backiel, and Alaska Regional Forester Phil Janik, all on behalf of the Forest Service, and Ted Stevens, Frank Murkowski, and Don Young. The main argument was from Stevens, Murkowski, and Young, who believed that the Forest Service was trying to purposefully limit the lumber market in Alaska. They demanded that the two species not be listed or that negative legislation would follow, likely resulting in budget and personnel cuts for the Forest Service.
The Tongass Land Management Plan (forest plan) was revised in 1997 after immense pressure from environmental groups to list the Alexander Archipelago wolf as threatened. The plan included a standard and guideline to sustain a habitat carrying capacity of least 18 Sitka deer per square mile to provide adequate prey and to limit the density of roads (i.e. miles of road per square mile). The forest plan also established a system of habitat reserves.  On the basis of the new plan, shortly afterward, FWS made a final determination that listing the wolf as threatened was unwarranted.
New Endangered Species Act petition – 2011
A new, 103-page petition to list the Alexander Archipelago wolf as a threatened or endangered species under the Endangered Species Act was filed with the US Fish and Wildlife Service by the Center for Biological Diversity and Greenpeace on August 10, 2011. The petition requests consideration for a separate listing of the Prince of Wales Island population because it is believed to be a distinct population segment, as well as a listing for the subspecies as a whole.
Litigation over wolf issues in four timber sales – filed in 2008, decided in 2011
However, Greenpeace and the Cascadia Wildlands Project later pointed out the data the Forest Service was using was known to be prone to overestimation of the carrying capacity for deer, the wolfs' primary prey. Furthermore, the conversion factor, known as the "deer multiplier", used in the calculations was incorrectly applied, causing a 30% overestimation of carrying capacity and corresponding underestimation of impacts. The two organizations determined, in total, the carrying capacity for the Sitka deer in places throughout the Tongass had been generally been overestimated by the Forest Service in its timber planning, by as much as 120% (varying geographically due to the faulty data).
In 2008, Greenpeace and the Cascadia Wildlands Project sued to stop the Forest Service from proceeding with four timber sales on the Tongass NF slated to extract around "30 million board-feet of Tongass timber", an amount close to the annual volume then being logged. The sales are on Prince of Wales, Kupreanof, Mitkof and Revillagigedo Islands. The suit challenged the agency's method of calculating the impact of logging on habitat carrying capacity for deer. In a radio story the plaintiff's spokesman explained that the dataset being used to represent habitat quality is actually uncorrelated to habitat quality, and that the deer multiplier mentioned above was misused according to the science under which it was derived. The story points to the Forest Service's underestimation of impacts not only to wolvles but to subsistence deer hunters.
In May 2010, US District Judge Ralph Beistline denied the plaintiff's motion for summary judgment, saying he "found no wrongdoing on the part of the Forest Service" and it was a "scientific disagreement". The plaintiffs have appealed the decision to the 9th US Circuit Court of Appeals (case 10-35567). A three-judge panel (Arthur Alarcon, Susan Graber, and Jay Bybee) heard oral arguments in the appeal on May 3, 2011 The panel ruled in favor of the plaintiffs on August 2, 2011, reversing in part, vacating in part and remanding the district court's decision. The ruling says, in part:
- "We do not think that USFS has adequately explained its decision to approve the four logging projects in the Tongass. ... USFS has failed to explain how it ended up with a table that identifies 100 deer per square mile as a maximum carrying capacity, but allows 130 deer per square mile as a potential carrying capacity. 'The agency is obligated to articulate a rational connection between the facts found and the choices made,' which the agency has not done here. Pac. Coast Fed’n of Fisherman’s Ass’ns v. U.S. Bureau of Reclamation, 426 F.3d 1082, 1091 (9th Cir. 2005)..."
- "We have similar questions about USFS’s use of VolStrata data, which identifies total timber volume and not forest structure, to approve the projects, where forest structure—and not total timber volume—is relevant to the habitability of a piece of land. USFS itself has recognized the limitations in the VolStrata data. ... Because we must remand to the agency to re-examine its Deer Model, we need not decide whether the use of the VolStrata data was arbitrary and capricious. We anticipate that, in reviewing the proposed projects, USFS will use the best available data ..."
In a statement to the press, a spokesman for the plaintiffs said the principles in this lawsuit apply to every significant timber sale between 1996 and 2008 before the Forest Service corrected errors in the deer model when the agency issued its revised Tongass Forest Plan. But, he said, the agency still fails to address cumulative impacts to deer, especially on Prince of Wales Island, as challenged in the Logjam timber sale lawsuit. He also said, "The purpose of the lawsuit is to make the Forest Service go back and do its analysis right on these timber sales. And we believe that when the Forest Service does that, that it can’t justify the volume of the timber that it decided to log in these projects."
Litigation over wolf issues in the Logjam timber sale – filed in 2010
In January 2010, the Forest Service was sued over its 73 million-board-foot Logjam timber sale on Prince of Wales Island, by Tongass Conservation Society, Cascadia Wildlands, and Greenpeace. At issue is the impact of excessive road density on wolf mortality and further loss of habitat for the primary prey (deer), among other issues. The plaintiff's motion for summary judgement was denied by the US District Court, Alaska, in September, and the case has been appealed to the 9th US Circuit Court of Appeals. The court scheduled oral arguments before Judges Betty Fletcher, Andrew Kleinfeld, and Consuelo Callahan for July 29, 2011, sitting in Anchorage.
- E. A. Goldman (1937). "The Wolves of North America". Journal of Mammalogy 18 (1): 37–45. doi:10.2307/1374306. JSTOR 1374306.
- "Canis lupus ligoni Goldman, 1937". Integrated Taxonomic Information System.
- Joshua Ross Ginsberg, David Whyte Macdonald & IUCN/SSC Canid Specialist Group (1990). "Sorting out the Canidae". Foxes, wolves, jackals, and dogs: an action plan for the conservation of canids. International Union for Conservation of Nature. pp. 4–8. ISBN 978-2-88032-996-9.
- "Wolves of the Rain Forest" – Wolf Song of Alaska. Alaska.net. Retrieved on 2012-12-31.
- "Suit Filed to Stop Four Timber Sales on Largest National Forest". Sitnews.us (2008-07-11). Retrieved on 2012-12-31.
- Pemberton, Mary. (2010-05-04) "Court upholds timber sales in Tongass Nat'l Forest". Business Week. Retrieved on 2012-12-31.
- Person, D.K. (Univ. Alaska); Kirchhoff, M. (ADF&G); van Ballenberghe, V. (USFS-FSL); Iverson, G.C. (USFS); Grossman, E. (USF&WS). 1996. "The Alexander Archipelago Wolf: A Conservation Assessment," Gen. Tech. Rep. PNW-GTR-384. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station.
- Natalie G. Dawson, Stephen O. MacDonald and Joseph A. Cook (2007) 2007 endemics AA.pdf "Endemic Mammals of the Alexander Archipelago". Southeast Alaska Conservation Assessment. Southeast Alaska Conservation Assessment – Chapter 6.7
- Comments of Greenpeace and others on the draft Tongass 5-Year Plan, fs.fed.us (November 5, 2010), citing a pers. comm. with ADF&G's Dave Person.
- See: Proposal 18 in the Alaska Board Of Game's Fall 2010 Proposal Book for the Southeast Region. http://www.boards.adfg.state.ak.us/gameinfo/meetinfo/2010-2011/southeast/crass-final9-7-10.pdf
- MacDonald, S.O. and J.A. Cook. 2007. Mammals and amphibians of Southeast Alaska. The Museum of Southwestern Biology, Special Publication 8:1–191.
- U.S. Forest Service. 1979. Tongass Land Management Plan. (See also later revisions.)
- Petition to List the Alexander Archipelago wolf under Provision of the Endangered Species Act, 12-Month Finding. US Fish & Wildlife Service, Dec. 8, 1994; signature date Feb. 15, 1995.
- Hanley, Thomas A.; Smith, Winston P.; Gende, Scott M. (2005). "Maintaining wildlife habitat in southeastern Alaska: Implications of new knowledge for forest management and research". Landscape and Urban Planning 72: 113. doi:10.1016/j.landurbplan.2004.09.016.
- Mother Jones (1997). Mother Jones Magazine. Mother Jones. pp. 46–. ISSN 0362-8841.
- Jack Ward Thomas; Harold K. Steen (2004). Jack Ward Thomas: The Journals of a Forest Service Chief. University of Washington Press. pp. 147–. ISBN 978-0-295-98398-1.
- 1997 Tongass Land Management Plan. USDA Forest Service.
- 12-Month Finding for a Petition to List the Alexander Archipelago wolf as Threatened and to Designate Critical Habitat. Notice in: Fed. Reg. 62(171:46709-46710. September 4, 1997.
- Petition to List the Alexander Archipelago wolf under Provisions of the Endangered Species Act, 12-Month Finding. U.S.F&WS. Signed Aug. 28, 1997.
- Jonathan Grass (August 11, 2011) Agency petitioned to protect SE wolves. Juneau Empire.
- Patti Epler (August 11, 2011) Should the Feds list an Alaska wolf as threatened or endangered? Alaska Dispatch.
- "Petition To List The Alexander Archipelagowolf (Canis lupus ligoni) As Threatened Or Endangered Under The United States Endangered Species Act." Center for Bioligical Diversity and Greenpeace (August 10, 2011).
- Caouette, J.; Kramer, M.; Nowacki, G. 2000. Deconstructing the Tongass Timber Paradigm. USDA Forest Service.
- Greenpeace v. Cole. Filed in U.S. District Court, Anchorage, Alaska.
- Joe Viechnicki (July 14, 2008) Groups challenge Tongass deer calculations. KFSK radio
- Audio recording. Oral arguments, 20 minutes per side. Greenpeace v. Cole. 9th Circuit Court of Appeals, siting in Anchorage, May 3, 2011.
- Memorandum. Issued unanimously by the 9th Circuit Court of Appeals panel of Judges Alarcon, Graber and Bybee. August 2, 2011.
- Jonathan Grass (August 3, 2011) Appeals court overturns 4 Tongass logging projects. Juneau Empire.
- Deanna Garrison (August 5, 2011) Court rules against four small timber sales, KRBD.
- Tongass Conservation Society v. USFS.
- Amy Lou Jenkins (January 12, 2010) [www.examiner.com/article/logging-tongass-costs-taxpayers-and-destroys-rare-ecosystems-suit-filed Logging in Tongass costs taxpayers and destroys rare ecosystems: Suit filed]. The Examiner
- KCAW radio, Nov. 16, 2010, Prince of Wales timber sale work resumes.
- Calendar for Anchorage, Alaska for July 25–29, 2011. 9th Circ. Court of Appeals.
- "Tongass loggers wary of wolf study protection could hurt timber cutting" – Anchorage Daily News
- "Environmentalists fail to get wolf on threatened list" – Orlando Sentinel
- "Group wants wolves protected in Alaska" – The Associated Press
- "'G' In The New 3g iPhone Will Not Also Stand For Green" – Common Dreams
- "Over Objections, U.S. Approves New Logging in Forest in Alaska" – New York Times
- "Ex-Fish and Game Officials Warn of Sealaska Bill" – ABC News