Temporal range: 1.977–1.98Ma Pleistocene
|In situ cranium of "Karabo"|
Berger et al., 2010
Australopithecus sediba is a species of Australopithecus of the early Pleistocene, identified based on fossil remains dated to about 2 million years ago. The species is known from six skeletons discovered in the Malapa Fossil Site at the Cradle of Humankind World Heritage Site in South Africa, one a juvenile male (MH1 also called "Karabo", the holotype), an adult female (MH2, the paratype), an adult male, and three infants. The fossils were found together at the bottom of the Malapa Cave, where they apparently fell to their death, and have been dated to between 1.977 and 1.980 million years ago.
Over 220 fragments from the species have been recovered to date. The partial skeletons were initially described in two papers in the journal Science by American and South African palaeo-anthropologist Lee R. Berger and colleagues as a newly discovered species of early human ancestor called Australopithecus sediba ("sediba" meaning "natural spring" or "well" in the Sotho language).
Australopithecus sediba may have lived in savannas but ate fruit and other foods from the forest—behavior similar to modern-day savanna chimpanzees. The conditions in which the individuals were buried and fossilized was extraordinary, permitting the extraction of plant phytoliths from dental plaque.
The first specimen of A. sediba was found by paleoanthropologist Lee Berger's nine-year-old son, Matthew, on August 15, 2008. While exploring near his father's dig site in the dolomitic hills north of Johannesburg, on the Malapa Nature Reserve, Matthew stumbled upon a fossilized bone. The boy alerted his father to the find, who could not believe what he saw — a hominid clavicle, and upon turning the block over: "sticking out of the back of the rock was a mandible with a tooth, a canine, sticking out. And I almost died", he later recalled. The fossil turned out to belong to a 4 ft 2 in (1.27 m) juvenile male, the skull of which was discovered in March 2009 by Berger's team. The find was announced to the public on April 8, 2010.
Also found at the Malapa archeological site were a variety of animal fossils, including saber-toothed cats, mongooses, and antelopes. Berger and geologist Paul Dirks speculated that the animals might have fallen into a deep, 100–150-foot (30–46 m) "death-trap", perhaps lured by the scent of water. The bodies may have then been swept into a pool of water rich with lime, and with sand at the bottom, making it possible for the remains to become fossilized.
Age estimates 
The fossil was dated using a combination of palaeomagnetism and uranium-lead (U-Pb) dating which showed that the fossils are not older than ~2.0 Ma. The occurrence of species of animal that became extinct at ~1.5 Ma indicate the deposit is not younger than 1.5 Ma. The sediments have a 'normal' magnetic polarity and the only major period between 2.0 and 1.5 Ma when this occurred is the Olduvai sub-Chron between 1.95 and 1.78 Ma. As such, the fossils were originally dated to ~1.95 Ma. Recent dating of a capping flowstone illustrated this was not possible and the normal magnetic polarity sediments have since been correlated to the 3000 year long Pre-Olduvai event at ~1.977 Ma.
Morphology and interpretations 
Because of the wide range of mosaic features exhibited in both cranial and post-cranial morphology, the authors suggest that A. sediba may be a transitional species between the southern African A. africanus (the Taung Child, Mrs. Ples) and either Homo habilis or even the later H. erectus (Turkana boy, Java man, Peking man).
The cranial capacity of MH1, which has been estimated to be at 95% of adult capacity (420 ml/cc), is at the higher end of the range for A. africanus and far from the lower range of early Homo (631 ml/cc), but the mandible and tooth size are quite gracile and similar to what one would expect to find in H. erectus; so similar are these features that, if found in isolation without other skeletal remains, they could be classified as Homo based on tooth and mandible size. However, the cusp spacing is more like Australopithecus.
Regardless of whether Australopithecus sediba is a direct ancestor of early Homo or not, our understanding of the range of variation in early hominins has been greatly increased with the finding of these new specimens.
A. sediba compared to its ancestor species A. africanus on the whole is described by Berger et al. as more derived towards Homo than A. garhi, especially showing a number of synapomorphies taken to anticipate the reorganization of the pelvis in H. erectus, associated with "more energetically efficient walking and running". The femur and tibia are fragmentary, but the foot combines an advanced anklebone combined with a primitive heel. Its cranial capacity is estimated at around 420–450 cm3 (26–27 cu in), about one-third that of modern humans.
A. sediba had a surprisingly modern hand, whose precision grip suggests it might have been another tool-making Australopithecus. Evidence of the precision gripping and stone tool production can be seen from Homo-like features such as having a long thumb and short fingers. The nearly complete wrist and hand of an adult female from Malapa, South Africa presents Australopithecus-like features, such as a strong flexor apparatus associated with arboreal locomotion.
A well-preserved and articulated ankle of A. sediba is mostly humanlike in form and function and possesses some evidence for a humanlike arch and Achilles tendon. However, Au. Sediba is apelike in possessing a more gracile calcaneal body and a more robust medial malleolus than expected. This shows that A. sediba may have practiced a unique combination of bipedalism and arboreality.
As opposed to the authors of the initial description, who interpreted both fossils as a possible transitional species between Australopithecus and Homo, other palaeoanthropologists are reluctant to do so. In an accompanying news article published with the initial descriptions in 2010, detractors of the idea that A. sediba might be ancestral to the genus Homo (e.g. Tim White and Ron Clarke) suggest that the fossils could be a late southern African branch of Australopithecus, co-existing with already existing members of the Homo genus.
This interpretation is based on the observation that the lower jaw, discovered by Friedemann Schrenk, of a 2.5 million year old fossil attributed to H. rudolfensis is the oldest known fossil ascribed to the Homo. This specimen is thus presumed to be older than the Australopithecus sediba fossils. These critics continue to ascribe to A. africanus the status of precursor of the genus Homo. Criticism has been raised in a news report on the find in Nature, to the effect that the authors of the initial description have failed to take into account the wealth of variation within A. africanus, in defining the finds as an independent species.
Additionally, the basing of the description of the species largely on the skeleton of a juvenile specimen has been subject to criticism, given that there is no certain way of saying to what extent adults would differ from juvenile specimens. More recent reviews appear to have accepted the specific status of A. sediba as an independent species from A. africanus and other early hominins. Additionally, the use of juvenile specimens as Type specimens has been a common practice in palaeoanthropology: even the type specimens of Homo habilis and Australopithecus africanus are juvenile specimens.
|Australopithecus sediba fossils|
See also 
- Berger, L. R.; de Ruiter, D. J.; Churchill, S. E.; Schmid, P.; Carlson, K. J.; Dirks, P. H. G. M.; Kibii, J. M. (2010). "Australopithecus sediba: a new species of Homo-like australopith from South Africa". Science 328 (5975): 195–204. doi:10.1126/science.1184944. PMID 20378811.
- Juliet King (June 4, 2010). "Australopithecus sediba fossil named by 17-year-old Johannesburg student". Origins Centre. Retrieved July 9, 2011.
- Ann Gibbons (2011). "A new ancestor for Homo?". Science 332 (6029): 534. doi:10.1126/science.332.6029.534-a. PMID 21527693.
- African fossils put new spin on human origins story - BBC News - Jonathan Amos - Retrieved 9 September 2011.
- Dirks, P. H. G. M.; Kibii, J. M.; Kuhn, B. F.; Steininger, C.; Churchill, S. E.; Kramers, J. D.; Pickering, R.; Farber, D. L. et al. (2010). "Geological setting and age of Australopithecus sediba from Southern Africa". Science 328 (5975): 205–208. doi:10.1126/science.1184950. PMID 20378812.
- Henry, Amanda G.; Ungar, Peter S.; Passey, Benjamin H.; Sponheimer, Matt; Rossouw, Lloyd; Bamford, Marion; Sandberg, Paul; de Ruiter, Darryl J. et al. (June 27, 2012). "The diet of Australopithecus sediba". Nature (journal). doi:10.1038/nature11185. Retrieved June 28, 2012.
- Boyle, Alan (June 27, 2012). "This pre-human ate like a chimp". MSNBC. Retrieved June 28., 2012.
- Wilford, John Noble (June 28, 2012). "Some Prehumans Feasted on Bark Instead of Grasses". New York Times. Retrieved June 28, 2012.
- Celia W. Dugger; John Noble Wilford (April 8, 2010). "New hominid species discovered in South Africa". New York Times. Archived from the original on 11 April 2010. Retrieved April 8, 2010.
- Kate Wong, "First of our kind: sensational fossils from South Africa spark debate over how we came to be human", Scientific American, April 2012:30-39.
- Kivell TL, Kibii JM, Churchill SE, Schmid P, Berger LR (2011). "Australopithecus sediba hand demonstrates mosaic evolution of locomotor and manipulative abilities". Science 333 (6048): 1411–1417. doi:10.1126/science.1202625. PMID 21903806.
- Part ape, part human - National Geographic - Retrieved 10 September 2011.
- Zipfel B, DeSilva JM, Kidd RS, Carison KJ, Churchill SE, Berger LR (2011). "The foot and ankle of Australopithecus sediba". Science 333 (6048): 1417–1420. doi:10.1126/science.1202703. PMID 21903807.
- Michael Balter (2010). "Candidate human ancestor from South Africa sparks praise and debate". Science 328 (5975): 154–155. doi:10.1126/science.328.5975.154. PMID 20378782.
- Michael Cherry (April 8, 2010). "Claim over 'human ancestor' sparks furore". Nature. doi:10.1038/news.2010.171.
- Fred Spoor (October 5, 2011). "Palaeoanthropology: Malapa and the genus Homo". Nature. doi:10.1038/478044a/news.2011.171.
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- Collection of papers on Australopithecus sediba from Science
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- Skeleton Analysis at Science
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