The biosphere is the global sum of all ecosystems. It can also be called the zone of life on Earth, a closed (apart from solar and cosmic radiation), and self-regulating system. From the broadest biophysiological point of view, the biosphere is the global ecological system integrating all living beings and their relationships, including their interaction with the elements of the lithosphere, hydrosphere, and atmosphere. The biosphere is postulated to have evolved, beginning through a process of biogenesis or biopoesis, at least some 3.5 billion years ago.
In a broader sense; biospheres are any closed, self-regulating systems containing ecosystems; including artificial ones such as Biosphere 2 and BIOS-3; and, potentially, ones on other planets or moons.
Origin and use of the term 
"The place on Earth's surface where life dwells."
While this concept has a geological origin, it is an indication of the impact of both Charles Darwin and Matthew F. Maury on the earth sciences. The biosphere's ecological context comes from the 1920s (see Vladimir I. Vernadsky), preceding the 1935 introduction of the term "ecosystem" by Sir Arthur Tansley (see ecology history). Vernadsky defined ecology as the science of the biosphere. It is an interdisciplinary concept for integrating astronomy, geophysics, meteorology, biogeography, evolution, geology, geochemistry, hydrology and, generally speaking, all life and earth sciences.
Narrow definition 
Some life scientists and earth scientists use biosphere in a more limited sense. For example, geochemists define the biosphere as being the total sum of living organisms (the "biomass" or "biota" as referred to by biologists and ecologists). In this sense, the biosphere is but one of four separate components of the geochemical model, the other three being lithosphere, hydrosphere, and atmosphere. The narrow meaning used by geochemists is one of the consequences of specialization in modern science. Some might prefer the word ecosphere, coined in the 1960s, as all encompassing of both biological and physical components of the planet.
The Second International Conference on Closed Life Systems defined biospherics as the science and technology of analogs and models of Earth's biosphere; i.e., artificial Earth-like biospheres. Others may include the creation of artificial non-Earth biospheres—for example, human-centered biospheres or a native Martian biosphere—in the field of biospherics.
Gaia hypothesis 
In the early 1970s, the British chemist James Lovelock and Lynn Margulis, a microbiologist from the United States, added to the hypothesis, specifically noting the ties between the biosphere and other Earth systems. For example, when carbon dioxide levels increase in the atmosphere, plants grow more quickly. As their growth continues, they remove more and more carbon dioxide from the atmosphere.
Ecosystems occur when communities and their physical environment work together as a system. The difference between this and a biosphere is simple, the biosphere is everything in general terms.
Extent of Earth's biosphere 
Every part of the planet, from the polar ice caps to the Equator, supports life of some kind. Recent advances in microbiology have demonstrated that microbes live deep beneath the Earth's terrestrial surface, and that the total mass of microbial life in so-called "uninhabitable zones" may, in biomass, exceed all animal and plant life on the surface. The actual thickness of the biosphere on earth is difficult to measure. Birds typically fly at altitudes of 650 to 1,800 metres, and fish that live deep underwater can be found down to -8,372 metres in the Puerto Rico Trench.
There are more extreme examples for life on the planet: Rüppell's vulture has been found at altitudes of 11,300 metres; bar-headed geese migrate at altitudes of at least 8,300 metres; yaks live at elevations between 3,200 to 5,400 metres above sea level; mountain goats live up to 3,050 metres. Herbivorous animals at these elevations depend on lichens, grasses, and herbs.
Microscopic organisms live at such extremes that, taking them into consideration, makes the thickness of the biosphere much greater. Culturable microbes have been found in the Earth's upper atmosphere as high as 41 km (25 mi) (Wainwright et al., 2003, in FEMS Microbiology Letters). It is unlikely, however, that microbes are active at such altitudes, where temperatures and air pressure are extremely low and ultraviolet radiation very high. More likely, these microbes were brought into the upper atmosphere by winds or possibly volcanic eruptions. Barophilic marine microbes have been found at more than 10 km (6 mi) depth in the Mariana Trench (Takamia et al., 1997, in FEMS Microbiology Letters). In fact, single-celled life forms have been found in the deepest part of the Mariana Trench, Challenger Deep, at depths of 36,201 feet (11,034 meters) . Microbes are not limited to the air, water or the Earth's surface. Culturable thermophilic microbes have been extracted from cores drilled more than 5 km (3 mi) into the Earth's crust in Sweden (Gold, 1992, and Szewzyk, 1994, both in PNAS), from rocks between 65-75 °C. Temperature increases with increasing depth into the Earth's crust. The speed at which the temperature increases depends on many factors, including type of crust (continental vs. oceanic), rock type, geographic location, etc. The upper known limit of temperature at which microbial life can exist is 122 °C (Methanopyrus kandleri Strain 116), and it is likely that the limit of life in the "deep biosphere" is defined by temperature rather than absolute depth.
Our biosphere is divided into a number of biomes, inhabited by broadly similar flora and fauna. On land, biomes are separated primarily by latitude. Terrestrial biomes lying within the Arctic and Antarctic Circles are relatively barren of plant and animal life, while most of the more populous biomes lie near the equator. Terrestrial organisms in temperate and Arctic biomes have relatively small amounts of total biomass, smaller energy budgets, and display prominent adaptations to cold, including world-spanning migrations, social adaptations, homeothermy, estivation and multiple layers of insulation.
Specific biospheres 
When the word is followed by a number, it is usually referring to a specific system or number. Thus:
- Biosphere 1, the planet Earth.
- Biosphere 2, a laboratory in Arizona, United States, which contains 3.15 acres (13,000 m²) of closed ecosystem.
- BIOS-3, a closed ecosystem at the Institute of Biophysics in Krasnoyarsk, Siberia, in what was then the Soviet Union.
- Biosphere J (CEEF, Closed Ecology Experiment Facilities), an experiment in Japan.
See also 
- The Columbia Encyclopedia, Sixth Edition. Columbia University Press. 2004. Retrieved 2010-11-12.
- Campbell, Neil A.; Brad Williamson; Robin J. Heyden (2006). Biology: Exploring Life. Boston, Massachusetts: Pearson Prentice Hall. ISBN 0-13-250882-6.
- "Meaning of biosphere". WebDictionary.co.uk. WebDictionary.co.uk. Retrieved 2010-11-12.
- Seuss, E. (1875) Die Entstehung Der Alpen [The Origin of the Alps]. Vienna: W. Braunmuller.
- Nakano et al.(1998)"Dynamic Simulation of Pressure Control System for the Closed Ecology Experiment Facility", Transactions of the Japan Society of Mechanical Engineers. 64:107-114.
- "Institute for Environmental Sciences". Ies.or.jp. Retrieved 2011-11-08.
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- Biosphere Definition
- Article on the Biosphere at Encyclopedia of Earth
- GLOBIO.info, an ongoing programme to map the past, current and future impacts of human activities on the biosphere
- Paul Crutzen Interview, freeview video of Paul Crutzen Nobel Laureate for his work on decomposition of ozone talking to Harry Kroto Nobel Laureate by the Vega Science Trust.
- Atlas of the Biosphere