Black mamba

From Wikipedia, the free encyclopedia
  (Redirected from Black Mamba)
Jump to: navigation, search
For other uses, see Black mamba (disambiguation).
Black mamba
Dendroaspis polylepis (14).jpg
Conservation status
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Suborder: Serpentes
Family: Elapidae
Subfamily: Elapinae
Genus: Dendroaspis
Species: D. polylepis
Binomial name
Dendroaspis polylepis
Günther, 1864[2]
Black mamba is located in Africa
Africa location map black mamba distribution.svg
Black mamba
In red, range of D. polylepis
In brown, D. polylepis may or may not occur here
  • Dendroaspis polylepis polylepis
    (Günther, 1864)
  • Dendraspis polylepis
    (Günther, 1864)
  • Dendraspis angusticeps
    (Boulenger 1896)
  • Dendraspis antinorii
    (Peters, 1873)
  • Dendroaspis polylepis antinorii
    (Peters, 1873)

The black mamba (Dendroaspis polylepis), also called the common black mamba or black-mouthed mamba, is a highly venomous snake of the genus Dendroaspis (Mambas), and is endemic to sub-Saharan Africa. It was first described in 1864 by Albert Günther. It is the longest species of venomous snake in Africa, and the second-longest venomous snake in the world after the king cobra. It is also the fastest moving snake in Africa, and one of the fastest moving snakes on the planet, perhaps the fastest,[4][5][6] capable of moving at 11 km/h (6.8 mph) over a distance of 43 m (141 ft).

The venom of the black mamba is highly toxic. Based on the median lethal dose (LD50) values in mice, the black mamba LD50 is 0.28/0.32 mg/kg subcutaneous.[7][8][9] Although there are other venomous snakes that exhibit higher LD50 toxicity scores, its venom is one of the most rapid-acting. In cases of severe envenomation, it is capable of killing an adult human in as few as 20 minutes. Two such cases have been documented in the medical literature. In one such case, an adult male was bitten on his right arm, just above the wrist by a black mamba which was estimated to be approximately 2.5 m (8.2 ft) in length. The victim began to show signs of prominent neurotoxicity within minutes. At ten minutes post-envenomation, respiratory paralysis set in and 20 minutes post-envenomation the victim showed no signs of life and was deceased.[10] Other cases of rapid death, within 30–60 minutes are relatively common with this species. However, depending on the nature of the bite, death time can be anywhere from 20 minutes to 6–8 hours. Without rapid and vigorous antivenom therapy, a bite from a black mamba is rapidly fatal almost 100% of the time.[11] One herpetologist called this species "death incarnate",[12] and to South African locals the black mamba bite is known as the "kiss of death".[13]

The black mamba is considered the most dangerous and feared[14] snake in Africa, though the ocellated carpet viper is responsible for more human fatalities due to snakebite than all other African species combined.[15] According to wildlife biologist Joe Wasilewski, black mambas are the most-advanced of all the snake species in the world. Their venom apparatus and method of delivering venom is also probably the most-effective and most-evolved among all venomous snakes.[16] Its combination of speed, unpredictable aggression, and potent venom make it an extremely dangerous species. Though the black mamba has a reputation for being very aggressive, like most snakes it usually attempts to flee from humans unless threatened.

Many experts regard the black mamba and the coastal taipan as the world's most dangerous snakes.


The eastern green mamba, Dendroaspis angusticeps, and the black mamba, Dendroaspis polylepis were considered a single species until 1946 when Dr. Vivian FitzSimons split them into separate species.[17]


The snake's scientific name is Dendroaspis polylepis: Dendroaspis meaning "tree asp", and polylepis meaning "many-scaled". The name "black mamba" is given to the snake not because of its body colour, but because of its ink-black mouth.[18] It displays this physical attribute when threatened.[18]


The black mamba's back skin colour is olive, brownish, grey, or sometimes khaki.[19] The adult snake's length is on average 2.5 meters (8.2 ft),[18] but some specimens have reached lengths of 4.3 to 4.5 meters (14.1 to 14.8 ft).[19] Black mambas weigh about 1.6 kilograms (3.5 lb)[18] on average. The species is the second longest venomous snake in the world, exceeded in length only by the king cobra.[19] The snake has an average life span of 11 years in the wild.[18]


The black mamba lives in Africa, occupying the following range: north east Democratic Republic of the Congo, south western Sudan to Ethiopia, Eritrea, Somalia, Kenya, eastern Uganda, Tanzania, Burundi, Rwanda, southwards to Mozambique, Swaziland, Malawi, Zambia, Zimbabwe and Botswana to KwaZulu-Natal in South Africa, and Namibia; then north easterly through Angola to south eastern Zaire.[20] The black mamba is not commonly found at altitudes above 1,000 metres (3,300 ft), although the distribution of black mamba does reach 1,800 metres (5,900 ft) in Kenya and 1,650 metres (5,410 ft) in Zambia.[20] The black mamba was also recorded in 1954 in West Africa in the Dakar region of Senegal.[20] However, this observation, and a subsequent observation that identified a second specimen in the region in 1956, has not been confirmed and thus the snake's distribution there is inconclusive.[20] The black mamba's distribution contains gaps within the Central African Republic, Chad, Nigeria and Mali. These gaps may lead physicians to misidentify the black mamba and administer an ineffective antivenom.[20]


The black mamba has adapted to a variety of climates ranging from savannah, woodlands, rocky slopes, dense forests and even humid swamps.[20] The grassland and savannah woodland/shrubs that extend through central, eastern and southern Africa are the black mamba's typical habitat.[20] The black mamba prefers more arid environments such as light woodland, rocky outcrops, and semi-arid dry bush country.[20]

Environmental encroachment[edit]

Sugarcane fields dominate the habitat of the black mamba.

The black mamba's environment is rapidly diminishing. In Swaziland alone, 75% of the people are involved in subsistence farming.[21] Because of agricultural encroachment on the black mamba's habitat, the snake is commonly found in sugarcane fields. The black mamba will climb to the top of the sugarcane to bask in the sun and possibly wait for prey. The majority of human attacks occur in the sugarcane fields of east and southern Africa which employ thousands of workers. This encroachment on the snake's territory contributes to potentially dangerous human contact with these venomous snakes.[18]


Dendroaspis polylepis by Bill Love.jpg
A black mamba climbing a branch, in London Zoo.

The black mamba uses its speed to escape threats, not to hunt prey.[18] It is shy and secretive; it always seeks to escape when a confrontation occurs.[18] If a black mamba is cornered it mimics a cobra by spreading a neck-flap, exposing its black mouth, and hissing.[18] If this endeavour to scare away the attacker fails, the black mamba may strike repeatedly.[18] The black mamba is a diurnal snake. Although its scientific name seems to be indicative of tree climbing, the black mamba is rarely arboreal.[22] These snakes retreat when threatened by predators.[23]

Hunting and prey[edit]

As stated, the black mamba is diurnal. It is an ambush predator that waits for prey to get close.[23] If the prey attempts to escape, the black mamba will follow up its initial bite with a series of strikes.[23] When hunting, the black mamba has been known to raise a large portion of its body off the ground.[23] The black mamba will release larger prey after biting it, but smaller prey, such as birds or rats, are held onto until the prey's muscles stop moving.[23] They have been known to prey on bushbabies, bats, and small chickens.[17]


It is the fastest moving snake found in Africa, and one of the fastest moving snakes in the world, perhaps the fastest.[4][5][6][24] Many exaggerated stories of the black mamba's moving speed have been spread,[25] and the elongated slender body can create the impression that it is moving faster than it really is.[26] It is only a myth that it can outrun a galloping horse or a running human. It is fast enough to catch a person walking but definitely cannot catch a person running. On 23 April 1906, on the Serengeti Plains, Kenya, an intentionally provoked and angry black mamba was recorded at a speed of 11 km/h (6.8 mph), over a distance of 43 m (141 ft).[27][28][29] A black mamba would not exceed 16 km/h (9.9 mph)[25][26] and it can only maintain such relatively high speeds for short distances.[26] Some have speculated that speeds of 16 to 19 km/h (9.9 to 11.8 mph) may be possible in short bursts over level ground.[30] Besides the relatively high speed with which it moves, the black mamba can strike accurately in any direction, even while moving fast. In striking, it throws its head upward from the ground to about two-fifths its total length.[31]


Not many predators challenge an adult black mamba. However, it does face a few threats such as birds of prey particularly snake eagles (genus Circaetus). Although all commonly prey on snakes, there are two species in particular that do so with high frequency, including preying on black mambas. The two species are the black-chested snake eagle (Circaetus pectoralis) and the brown snake eagle (Circaetus cinereus). The Cape file snake (Mehelya capensis) which is apparently immune to all African snake venoms and preys on other snakes including venomous ones, is a common predator of black mambas (up to a size it can swallow).[17][32][33] Mongooses which are also partially immune to venom, and are quick enough to evade a bite, will readily tackle a black mamba for prey.[34] Humans do not usually consume black mambas, but they often kill them out of fear.


Based on the median lethal dose (LD50) values in mice, the black mamba LD50 from all published sources is as follows:

Its bites can deliver about 100–120 mg of venom on average and the maximum dose recorded is 400 mg. It is reported that before the antivenom was widely available, the mortality rate from a bite was nearly 100%.[18] Black mamba bites can potentially kill a human within 20 minutes, but death usually occurs after 30–60 minutes, sometimes taking up to three hours. (The fatality duration and rate depend on various factors, such as the health, size, age, psychological state of the human, the penetration of one or both fangs from the snake, amount of venom injected, location of the bite, and proximity to major blood vessels.[18] The health of the snake and the interval since it last used its venom mechanism is also important.) Presently, there is a polyvalent antivenom produced by SAIMR (South African Institute for Medical Research) to treat black mamba bites from many localities.[11]

If bitten, common symptoms are rapid onset of dizziness, coughing or difficulty breathing, and erratic heartbeat.[11] In extreme cases, when the victim has received a large amount of venom, death can occur in less than an hour from respiratory or cardiac arrest.[11] Also, the black mamba's venom has been known to cause paralysis.[11] Death is usually due to suffocation resulting from paralysis of the respiratory muscles.[11]

The black mamba is regarded as one of the most dangerous and feared snakes in Africa.[14] Nevertheless, attacks on humans by black mambas are rare, as the snakes usually avoid confrontation with humans and their occurrence in highly populated areas is not very common compared with some other species.[40]


The black mamba's venom is dendrotoxin. The toxin disrupts the exogenous process of muscle contraction by means of the sodium potassium pump. In an experiment, the death time of a mouse after subcutaneous injection of some toxins studied is around 7 minutes. However, a black mamba venom can kill a mouse after 4.5 minutes.[14]

Case reports[edit]

In a reported case in which a black mamba attacked and killed nine people at the same time is explained by biologist Joe Wasilewski in a short video clip. In an African village, screams were heard coming from someone in a hut. Another person heard the screams and entered the hut but never came out. This continued until a total of nine people entered the hut. Screams were heard but after the ninth person who entered the hut, nobody else was willing to see what was going on inside. The next day, some courageous locals gathered with spears and other weapons and entered the hut. As they walked into the hut, they witnessed nine dead people and a baby who was alive but hungry. The hut was a refuge which the mamba had decided to spend the night in, but as one person after the other walked into the hut, the snake felt cornered and became very defensive. It attacked each of the people who entered the hut. Because of the disturbingly rapid death times of the nine victims, the attacks must have been extraordinarily savage in nature.[41]

A survey of snakebites in South Africa from 1957 to 1963 recorded over 900 venomous snakebites, but only seven of these were confirmed black mamba bites. From the 900 bites, only 21 ended in fatalities, including all seven black mamba bites – a 100% mortality rate.[42]

Danie Pienaar, head of South African National Parks Scientific Services, garnered much media attention after he survived the bite of a black mamba without antivenom. Mortality rates among those not receiving any kind of adequate medical attention (antivenom, mechanical ventilation, intubation, drug therapy, etc.) is 100%. This is why the case of Danie Pienaar stands out and why it garnered such media attention internationally. However, although no antivenom was administered, Pienaar was in serious condition despite the fact the hospital physicians declared it a "moderate" black mamba envenomation. At one point, he lapsed into a coma and his prognosis was declared "poor". Still, upon arrival at the hospital Pienaar was immediately intubated, given supportive drug therapy, put on mechanical ventilation and was placed on life support for three days until the toxins were flushed out of his system, and released from the hospital on the fifth day. He survived the bite and made a full recovery. Danie believes he survived for a number of reasons. “Firstly, it was not my time to go.” The fact that he stayed calmed and moved slowly definitely helped. The tourniquet was also essential. “It was not easy to stay calm,” he says. “It was almost as if I dislodged myself from my body and was talking to someone else the entire time.” “I also knew snakes and have never been scared of them,” he said. The experience has not left Danie with antagonism towards the black mamba. “Snakes do not eat people. I was on my way and cut the snake off from where it wanted to go.”[43][44] He was featured on the I'm Alive TV series (season one, episode 8 "Last Man Standing").

In 2013, in a very rare and unusual case, American professional photographer, Mark Laita, was bitten on the leg by a black mamba during a photo-shoot of a black mamba at a facility in Central America. Both the snake fangs hit an artery in his calf, and he was gushing blood profusely. Laita did not go to the doctor or the hospital, and except for the swollen fang marks giving him intense pain during the night, he was not affected and was fine physically. This led him to believe that the snake either gave him a "dry bite" (meaning without injecting venom) or that the heavy bleeding pushed the venom out. Some commenters to the story suggested that it was a venomoid snake (in which the venom glands are surgically removed). Laita responded that it was not the case. Only later, Laita found that he had captured the snake biting his leg on film. (see citation for the photo)[45][46][47]



  1. ^ Spawls, S. (2009). "Dendroaspis polylepis". IUCN Red List of Threatened Species. Version 3.1. International Union for Conservation of Nature. Retrieved 13 December 2013. 
  2. ^ "Dendroaspis polylepis". Integrated Taxonomic Information System. Retrieved 12 December 2013. 
  3. ^ Uetz, P. "Dendroaspis polylepis Günther, 1864". Reptile Database. Zoological Museum Hamburg. Retrieved 2 December 2011. 
  4. ^ a b Van Der Vlies, C. (2010). Southern Africa Wildlife and Adventure. British Columbia, Canada/Indiana, United States: Trafford Publishing. pp. 180–181. ISBN 978-1-4269-1932-9. 
  5. ^ a b Austin Stevens: Snakemaster, (2002) "Seven Deadly Strikes" documentary
  6. ^ a b World Book, Inc (1999) The World Book encyclopedia, Volume 1, Page 525
  7. ^ a b Spawls, S.; Branch, B. (1995). The dangerous snakes of Africa: natural history, species directory, venoms, and snakebite. Dubai: Oriental Press: Ralph Curtis-Books. pp. 49–51. ISBN 0-88359-029-8. 
  8. ^ Minton, Minton, SA, MR (1969). Venomous Reptiles. USA: New York Charles Scribner's Sons. 
  9. ^ a b c d Fry, Bryan, Deputy Director, Australian Venom Research Unit, University of Melbourne (March 9, 2002). "Snakes Venom LD50 – list of the available data and sorted by route of injection". (archived) Retrieved October 14, 2013.
  10. ^ Visser, Chapman, J, DS (1978). Snakes and Snakebite: Venomous snakes and management of snake bite in Southern Africa. Purnell. ISBN 0-86843-011-0. 
  11. ^ a b c d e f "Immediate first aid for bites by Black Mamba (Dendroaspis polylepis polylepis)". Toxicology. University of California, San Diego. Retrieved 3 December 2013. 
  12. ^ Lee, Douglas (November–December 1996). "Black Mamba! Meet the African Species That One Expert Calls 'Death Incarnate'". International Wildlife 26 (6): 48. Retrieved 20 December 2013. 
  13. ^ Black mamba: Kiss of death, Smithsonian Networks
  14. ^ a b c Strydom, Daniel (1971-11-12). "Snake Venom Toxins" (PDF). The Journal of Biological Chemistry. Retrieved 2010-04-24. 
  15. ^ a b Jerry G. Walls, The World's Deadliest Snakes, Reptiles
  16. ^ Wasilewski, Joe (2011). Wildlife Footage (Motion picture). Africa: Youtube. Retrieved 27 December 2013. 
  17. ^ a b c Haagner, GV; Morgan, DR (January 1993). "The maintenance and propagation of the Black mamba Dendroaspis polylepis at the Manyeleti Reptile Centre, Eastern Transvaal". International Zoo Yearbook 32 (1): 191–196. doi:10.1111/j.1748-1090.1993.tb03534.x.  Retrieved 14 December 2013.
  18. ^ a b c d e f g h i j k l "Black mamba". National Geographic Society. Retrieved 2010-03-12. 
  19. ^ a b c Mattison, Chris (1987-01-01). Snakes of the World. New York: Facts on File, Inc. p. 164. 
  20. ^ a b c d e f g h Håkansson, Thomas; Thomas Madsen (1983-01-01). "On the Distribution of the Black Mamba (Dendroaspis polylepis) in West Africa". Journal of Herpetology (Society for the Study of Amphibians and Reptiles). JSTOR 1563464. 
  21. ^ "UNDP: Human development indices – Table 3: Human and income poverty (Population living below national poverty line (2000–2007))". United Nations Development Programme. 28 November 2008. Retrieved 29 December 2009. 
  22. ^ Maina, J.N (1989-12). "The morphology of the lung of the black mamba Dendroaspis polylepis". The Journal of Anatomy 167: 31–46. PMC 1256818. PMID 2630539.  Check date values in: |date= (help)
  23. ^ a b c d e Richardson, Adele (2004). Mambas. Mankato, Minnesota: Capstone Press. p. 25. Retrieved 2010-05-19. 
  24. ^ Irwin, Steve (July 30, 2000) "Africa's Deadliest Snakes" , The Crocodile Hunter (SE3EP27)
  25. ^ a b Mikael Jolkkonen, Muscarinic Toxins from Dendroaspis (Mamba) Venom Page 15, Uppsala University
  26. ^ a b c Warren, Schmidt (2006) Reptiles & Amphibians of Southern Africa, Page 34
  27. ^ Gerald L. Wood , (1976) The Guinness book of animal facts and feats, Page 132
  28. ^ Leopard, Marten (2002) International Wildlife Encyclopedia, Page 1530
  29. ^ Harry W. Greene, (1997), Snakes: The Evolution of Mystery in Nature, Page 40
  30. ^ Medd Guinness (1993), The Guinness Book of Records, Page 35
  31. ^ Burton, R. (2002). International Wildlife Encyclopedia: Leopard – marten. USA: Marshall Cavendish. p. 3168. ISBN 0-7614-7277-0. 
  32. ^ Bauchot, R. (2006). Snakes: A Natural History. Sterling. pp. 41,76,176. ISBN 978-1-4027-3181-5. 
  33. ^ Mehelya capensis (Southern file snake, Cape file snake), Iziko South African Museum
  34. ^ Caught in the Act: Mongoose Vs. Snake (Video), "A mongoose takes on the most infamous snake of all, the black mamba.", National Geographic
  35. ^ a b Sherman A. Minton, (May 1, 1974) Venom diseases, Page 116
  36. ^ Philip Wexler, 2005, Encyclopedia of toxicology, Page 59
  37. ^ Thomas J. Haley, William O. Berndt, 2002, Toxicology, Page 446
  38. ^ Scott A Weinstein, David A. Warrell, Julian White and Daniel E Keyler (Jul 1, 2011) " Bites from Non-Venomous Snakes: A Critical Analysis of Risk and Management of "Colubrid" Snake Bites (page 246)
  39. ^ Zug, GR. (1996). Snakes in Question: The Smithsonian Answer Book. Washington D.C., US: Smithsonian Institution Scholarly Press. ISBN 1-56098-648-4. 
  40. ^ The new encyclopedia of Reptiles (Serpent). Time Book Ltd. 2002. 
  41. ^ Wasilewski, Joe (2011). Wildlife Footage (Documentary). Africa: Youtube. Event occurs at 0m55s. Retrieved 27 December 2013. 
  42. ^ O'Shea, M. (2005). Venomous Snakes of the World. United Kingdom: New Holland Publishers. p. 78. ISBN 0-691-12436-1. ... in common with other snakes they prefer to avoid contact; ... from 1957 to 1963 ... including all seven black mamba bites - a 100 per cent fatality rate 
  43. ^ "Scientists gather in Kruger National Park for Savanna Science Network Meeting". South African Tourism. 5 March 2013. Retrieved 26 April 2014. 
  44. ^ "Surviving a Black Mamba bite". Siyabona Africa - Kruger National Park. Siyabona Africa. Retrieved 26 April 2014. 
  45. ^ Rowan Hooper (January 19, 2012) Portraits of snake charm worth elephant-killing bite newscientist
  46. ^
  47. ^

Further reading[edit]

  • Thorpe, Roger S.; Wolfgang Wüster, Anita Malhotra (1996). Venomous Snakes: Ecology, Evolution, and Snakebite. Oxford, England: Oxford University Press. ISBN 978-0-19-854986-4
  • McDiarmid, Roy W.; Jonathan A. Campbell; T'Shaka A. Tourè (1999). Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1. Washington, District of Columbia: Herpetologists' League. ISBN 978-1-893777-01-9
  • Spawls, Stephen; Branch, Bill (1995). Dangerous Snakes of Africa: Natural History - Species Directory - Venoms and Snakebite. Ralph Curtis Pub; Revised edition. ISBN 978-0-88359-029-4
  • Dobiey, Maik; Vogel, Gernot (2007). Terralog: Venomous Snakes of Africa (Terralog Vol. 15). Aqualog Verlag GmbH.; 1st edition. ISBN 978-3-939759-04-1
  • Mackessy, Stephen P. (2009). Handbook of Venoms and Toxins of Reptiles. CRC Press; 1st edition. ISBN 978-0-8493-9165-1
  • Greene, Harry W.; Fogden, Michael; Fogden, Patricia (2000). Snakes: The Evolution of Mystery in Nature. University of California Press. ISBN 978-0-520-22487-2
  • Spawls, Stephen; Ashe, James; Howell, Kim; Drewes, Robert C. (2001). Field Guide to the Reptiles of East Africa: All the Reptiles of Kenya, Tanzania, Uganda, Rwanda and Burundi Princeton University Press. ISBN 978-0-12-656470-9
  • Broadley, D.G.; Doria, C.T.; Wigge, J. (2003). Snakes of Zambia: An Atlas and Field Guide. Frankfurt am Main, Germany: Edition Chimaira. ISBN 978-3-930612-42-0
  • Marais, Johan (2005). A Complete Guide to the Snakes of Southern Africa. Cape Town, South Africa: Struik Nature. ISBN 978-1-86872-932-6
  • Engelmann, Wolf-Eberhard (1981). Snakes: Biology, Behavior, and Relationship to Man. Leipzig; English version NY, US: Leipzig Publishing; English version published by Exeter Books (1982). ISBN 0-89673-110-3
  • Minton, Sherman A. (1969). Venomous Reptiles. US: New York Simon Schuster Trade. ISBN 978-0-684-71845-3
  • FitzSimons, Vivian FM (1970). A field guide to the snakes of Southern Africa. Canada: HarperCollins. ISBN 0-00-212146-8
  • Department of the Navy Bureau of Medicine and Surgery (2013). Venomous Snakes of the World: A Manual for Use by U.S. Amphibious Forces. Skyhorse Publishing. ISBN 978-1-62087-623-7
  • Branch, Bill (1988). Bill Branch's Field Guide to the Snakes and Other Reptiles of Southern Africa (More than 500 Photographs for Easy Identification). Cape Town, South Africa: Struik Publishers. ISBN 978-0-86977-641-4
  • Branch, Bill (1998). Field Guide to the Snakes and Other Reptiles of Southern Africa. Ralph Curtis Publishing. ISBN 978-0-88359-042-3
  • Branch, Bill (2005). Photographic Guide to Snakes Other Reptiles and Amphibians of East Africa. Sanibel Island, Florida: Ralph Curtis Books. ISBN 978-0-88359-059-1
  • Mara, Wil; Collins, Joseph T; Minton, SA (1993). Venomous Snakes of the World. TFH Publications Inc. ISBN 978-0-86622-522-9
  • Stocker, Kurt F. (1990). Medical Use of Snake Venom Proteins. CRC Press. ISBN 978-0-8493-5846-3
  • Mebs, Dietrich (2002). Venomous and Poisonous Animals: A Handbook for Biologists, Toxicologists and Toxinologists, Physicians and Pharmacists. Medpharm. ISBN 978-0-8493-1264-9
  • White, Julian; Meier, Jurg (1995). Handbook of Clinical Toxicology of Animal Venoms and Poisons. CRC Press. ISBN 978-0-8493-4489-3
  • Vitt, Laurie J; Caldwell, Janalee P. (2013). Herpetology, Fourth Edition: An Introductory Biology of Amphibians and Reptiles. Academic Press. ISBN 978-0-12-386919-7
  • Tu, Anthony T. (1991). Handbook of Natural Toxins, Vol. 5: Reptile Venoms and Toxins. Marcel Dekker. ISBN 978-0-8247-8376-1
  • Mattison, Chris (1995). The Encyclopedia of Snakes. Facts on File; 1st U.S. edition. ISBN 978-0-8160-3072-9
  • Coborn, John (1991). The Atlas of Snakes of the World. TFH Publications. ISBN 978-0-86622-749-0

External links[edit]

Media related to Dendroaspis polylepis at Wikimedia Commons