Breeding summer visitorDistributions are very approximate. Most birds winter in the eastern African range.
The European Nightjar, Eurasian Nightjar or just Nightjar (Caprimulgus europaeus) is a crepuscular and nocturnal bird in the nightjar family that breeds in most of Europe and temperate Asia. The six subspecies differ clinally, with birds becoming smaller and paler towards the east of the range. All populations are migratory, wintering in Africa south of the Sahara. Their densely patterned grey and brown plumage makes individuals difficult to see in the daytime when they rest on the ground or perch motionless along a branch, although the male shows white patches in the wings and tail as he flies at night.
This is a bird of dry, open country with some trees and small bushes, such as heaths, forest clearings or newly planted woodland. The male occupies a territory in spring and advertises his presence with a distinctive sustained churring trill from a perch. He patrols his territory with wings held in a V and tail fanned, chasing intruders while wing-clapping and calling. Wing clapping also occurs when he chases the female in a spiralling display flight. The European Nightjar does not build a nest, and its two grey and brown blotched eggs are laid directly on the ground. The phase of the moon may affect the timing of egg laying. The eggs hatch after about 17–21 days and the downy chicks fledge in another 16–17 days .
The European Nightjar feeds on a wide variety of flying insects, which it seizes in flight, often fly-catching from a perch. It hunts by sight silhouetting its prey against the night sky. It has relatively large eyes, each with a reflective layer, which improves night vision. It appears not to rely on its hearing to find insects and does not echolocate. It drinks and bathes on the wing. Although it suffers a degree of predation and parasitism, the main threats to this species are habitat loss, disturbance and a reduction of its insect prey through pesticide use. Despite population decreases, its large numbers and huge breeding range mean that this species is classified by the International Union for Conservation of Nature as being of Least Concern.
The myth that this nightjar suckled goats causing them to cease to give milk dates back to antiquity, and the view that this nocturnal bird represents the souls of unbaptised children is also very old. Modern poets have tended to depict this bird as illustrative of warm summer nights.
The nightjars, Caprimulgidae, are a large family of mostly nocturnal insect-eating birds. The largest and most widespread of the several genera is Caprimulgus, the "typical" nightjars, characterised by stiff bristles around the mouth and patterned plumage. The males, and sometimes females, often have white markings in the wing or tail. Within the genus, the European Nightjar's closest relatives are two African species with similar songs, the Rufous-cheeked Nightjar and the Sombre Nightjar. The Jungle Nightjar is sometimes considered to be the eastern equivalent of the European species. The European Nightjar was described by Linnaeus in his Systema naturae in 1758 under its current scientific name. Caprimulgus is derived from the Latin capra, "nanny goat" and mulgere "to milk", referring to an old myth that nightjars suck milk from goats, and the species name, europaeus is Latin for "European".
"Nightjar", first recorded in 1630, refers to the nocturnal habits of the bird, the second part of the name deriving from the distinctive churring song.
There are six recognised subspecies, although differences are mainly clinal; birds become smaller and paler in the east of the range and the males have larger white wing spots. Birds of intermediate appearance occur where the subspecies' ranges overlap.
|C. e. europaeus||Linnaeus, 1758||Across north and central Europe and north Central Asia||The nominate subspecies|
|C. e. meridionalis||Hartert, 1896||Northwest Africa and southern Europe east to the Caspian Sea||Smaller, paler and greyer than nominate, with larger white spots|
|C. e. sarudnyi||Hartert, 1912||Kazakhstan to Kyrgyzstan||Variable due to interbreeding with other forms, but generally pale with large white spots|
|C. e. unwini||Hume, 1871||Iraq and Iran east to Uzbekistan||Hume's Nightjar; paler, greyer and plainer than the nominate form and often has white throat patches|
|C. e. plumipes||Przewalski, 1876||Northwestern China and western Mongolia||A pale, sandy-buff form with large white spots|
|C. e. dementievi||Stegmann, 1949||Northeastern Mongolia||A poorly known paler and greyer form|
The fossil record is deficient, but it is likely that these poorly defined subspecies diverged as global temperature rose over the last 10,000 years or so. Only one record of this species possibly dates back to the late Eocene, and records of any member of the family as far back as the late Pleistocene are very rare.
The European Nightjar is 24.5–28 cm (9.6–11.0 in) long with a 52–59 cm (20–23 in) wingspan. The male weighs 51–101 g (1.8–3.6 oz) and the female 67–95 g (2.4–3.4 oz). The adult of the nominate subspecies has greyish-brown upperparts with dark streaking, a pale buff hindneck collar and a white moustachial line. The closed wing is grey with buff spotting, and the underparts are greyish-brown, with brown barring and buff spots. The bill is blackish, the iris is dark brown and the legs and feet are brown.
The flight on long pointed wings is noiseless, due to their soft plumage, and very buoyant. Flying birds can be sexed since the male has a white wing patch across three primary feathers and white tips to the two outer tail feathers, whereas females do not show any white in flight. Chicks have downy brown and buff plumage, and the fledged young are similar in appearance to the adult female. Adults moult their body feathers from June onwards after breeding, suspend the process while migrating, and replace the tail and flight feathers on the wintering grounds. Moult is completed between January and March. Immature birds follow a similar moult strategy to the adults unless they are from late broods, in which case the entire moult may take place in Africa.
Other nightjar species occur in parts of the breeding and wintering ranges. The Red-necked Nightjar breeds in Iberia and northwest Africa; it is larger, greyer and longer winged than the European Nightjar, and has a broad buff collar and more conspicuous white markings on the wings and tail. Wintering European Nightjars in Africa may overlap with the related Rufous-cheeked and Sombre Nightjars. Both have a more prominent buff hind-neck collar and more spotting on the wing coverts. The Sombre Nightjar is also much darker than its European cousin. Given their nocturnal habits, cryptic plumage and difficulty of observation, nightjar observation "is as much a matter of fortune as effort or knowledge".
The male European Nightjar's song is a sustained churring trill, given continuously for up to 10 minutes with occasional shifts of speed or pitch. It is delivered from a perch, and the male may move around its territory using different song posts. Singing is more frequent at dawn and dusk than during the night, and is reduced in poor weather. The song may end with a bubbling trill and wing-clapping, perhaps indicating the approach of a female. Migrating or wintering birds sometimes sing. Individual male Nightjars can be identified by analysing the rate and length of the pulses in their songs. Even a singing male may be hard to locate; the perched bird is difficult to spot in low light conditions, and the song has a ventriloquial quality as the singer turns his head. The song is easily audible at 200 m (660 ft), and can be heard at 600 m (2,000 ft) in good conditions; it can be confused with the very similar sound of the European mole cricket.
When on the wing, both sexes give a short cuick, cuick call, also used when chasing predators. Other calls include variations on a sharp chuck when alarmed, hisses given by adults when handled or chicks when disturbed, and an assortment of wuk, wuk, wuk, muffled oak, oak and murmurs given at the nest.
Distribution and habitat
The breeding range of the European Nightjar comprises Europe north to around latitude 64°N and Asia north to about 60°N and east to Lake Baikal and eastern Mongolia. The southern limits are northwestern Africa, Iraq, Iran and the northwestern Himalayas. This nightjar formerly bred in Syria and Lebanon.
All populations are migratory, and most birds winter in Africa south of the Sahara, with just a few records from Pakistan, Morocco and Israel. Migration is mainly at night, singly or in loose groups of up to twenty birds. European breeders cross the Mediterranean and North Africa, whereas eastern populations move through the Middle East and East Africa. Some Asian birds may therefore cross 100° of longitude on their travels. Most birds winter in eastern or southeastern Africa, although individuals of the nominate race have been recently discovered wintering in the Democratic Republic of the Congo; records elsewhere in West Africa may be wintering birds of this subspecies or C. e. meridionalis. Most autumn migration takes place from August to September, and the birds return to the breeding grounds by May. Vagrants have occurred in Iceland, the Faroe Islands, the Seychelles, the Azores, Madeira and the Canary Islands.
This is a bird of dry, open country with some trees and small bushes, such as heaths, commons, moorland, forest clearings or felled or newly planted woodland. When breeding, it avoids treeless or heavily wooded areas, cities, mountains, and farmland, but it often feeds over wetlands, cultivation or gardens. In winter it uses a wider range of open habitats including acacia steppe, sandy country and highlands. It has been recorded at altitudes of 2,800 m (9,100 ft) on the breeding grounds and 5,000 m (16,400 ft) in the wintering areas.
The European Nightjar is crepuscular and nocturnal. During the day it rests on the ground, often in a partly shaded location, or perches motionless lengthwise along an open branch or a similar low perch. The cryptic plumage makes it difficult to see this species in the daytime, and birds on the ground, if they are not already in shade, will turn occasionally to face the sun thereby minimising their shadow. A roost site is used regularly if it is undisturbed, sometimes for weeks at a time. If it feels threatened, the nightjar flattens itself to the ground with eyes almost closed, flying only when the intruder is 2–5 m (7–16 ft) away. It may call or wing clap as it goes, and lands as far as 40 m (130 ft) from where it was flushed. In the wintering area it often roosts on the ground but also uses tree branches up to 20 m (66 ft) high. It uses trees more often than the local nightjar species. Again, roost sites are used on a regular basis.
Like other nightjars, it will sit on roads or paths during the night and hover to investigate large intruders such as deer or humans. It may be mobbed by birds while there is still light, and by bats, other nightjar species or Eurasian Woodcocks during the night. Owls and other predators such as red foxes will be mobbed by both male and female European Nightjars. Like other aerial birds, such as swifts and swallows, nightjars make a quick plunge into water to wash. They have a unique serrated comb-like structure on the middle claw that is used to preen and perhaps remove parasites.
In cold or inclement weather, several nightjar species can slow their metabolism and go into torpor, notably the Common Poorwill, which will maintain this state for weeks. The European Nightjar has been observed in captivity to be able to maintain a state of torpor for at least eight days without harm, but the relevance of this to wild birds is unknown.
Breeding is normally from late May to August, but may be significantly earlier in northwest Africa or western Pakistan. Returning males arrive about two weeks before the females and establish territories from 1.5–32 ha (3.7–80 acres) in area. They patrol their territories with wings held in a V and tail fanned, chasing intruders while wing-clapping and calling. Fights make take place in flight or on the ground. The male's display flight involves a similar wing and tail position with frequent wing clapping as he follows the female in a rising spiral. If she lands, he continues to display with bobbing and fluttering until the female spreads her wings and tail for copulation. Mating occasionally takes place on a raised perch instead of the ground. In good habitat, there may be 20 pairs per square kilometre (50 per square mile).
This species is normally monogamous. There is no nest, and the eggs are laid on the ground among plants or tree roots, or beneath a bush or tree. The site may be bare ground, leaf litter or pine needles, and is used for a number of years. The clutch is usually one or two whitish eggs, rarely unmarked, but normally blotched with browns and greys. The eggs average 32 x 22 mm (1.26 x 0.87 in) and weigh 8.4 g (0.3 oz), of which 6% is shell.
Several nightjar species are known to be more likely to lay in the two weeks before the full moon than the during the waning moon, possibly because insect food may be easier to catch as the moon waxes. A study specifically looking at the European Nightjar showed that the phase of the moon is a factor for birds laying in June, but not for earlier breeders. This strategy means that a second brood in July would also have a favourable lunar aspect.
Eggs are laid 36–48 hours apart, and incubation, mainly by the female, starts with the first egg. The male may incubate for short periods, especially around dawn or dusk, but spends the day roosting, sometimes outside his territory or close to other males. Each egg hatches after about 17–21 days. The semi-precocial downy chicks are mobile when hatched, but are brooded to keep them warm. They fledge in 16–17 days and become independent of the adults around 32 days after hatching. A second brood may be raised by early nesting pairs, in which case the female leaves the first brood a few days before they fledge; the male then cares for the first brood and assists with the second. Both adults feed the young with balls of insects. If the female is disturbed while breeding, she runs or flutters along the ground feigning injury until she has drawn the intruder away. She may also move the eggs a short distance with her bill.
Broods that fail tend to do so during incubation. One English study showed that only 14.5% of eggs survived to hatching, but once that stage was reached the chances of fledging successfully were high. European Nightjars breed when aged one year, and typically live for four years. The adult annual survival rate is 70%, but that for juveniles is unknown. The maximum known age is just over 12 years.
The European Nightjar feeds on a wide variety of flying insects, including moths, beetles, mantises, dragonflies, cockroaches and flies. It will pick glowworms off vegetation. It consumes grit to aid with digesting its prey, but any plant material and non-flying invertebrates consumed is taken inadvertently while hunting other food items. Young chicks have been known to eat their own faeces.
Birds hunt over open habitats and woodland clearings and edges, and may be attracted to insects concentrating around artificial lights, near farm animals or over stagnant ponds. They usually feed at night, but occasionally venture out on overcast days. Nightjars pursue insects with a light twisting flight, or flycatch from a perch; they may rarely take prey off the ground. They drink by dipping to the water surface as they fly. Breeding European Nightjars travel on average 3.1 km (3,400 yd) from their nests to feed. Migrating birds live off their fat reserves.
European Nightjars hunt by sight, silhouetting their prey against the night sky. They tend to flycatch from a perch on moonlit nights, but fly continuously on darker nights when prey is harder to see. Hunting frequency reduces in the middle of the night. Although they have tiny bills, the mouth can be opened very wide as they catch insects. They have long sensitive bristles around the mouth, which may help to locate or funnel prey into the mouth. Indigestible parts of insects, such as chitin exoskeletons are regurgitated as pellets.
Nightjars have relatively large eyes each with a tapetum lucidum (reflective layer behind the retina), which makes the eyes shine in torchlight and improves light detection at dusk, dawn and in moonlight. The retinas of nocturnal birds, including nightjars, are adapted for sight in low-light levels with a higher density of rod cells and far fewer cone cells, when compared to those of most diurnal birds. These adaptations favouring nightjars night vision will have compromised their colour vision. In many day-flying species, light passes through coloured oil droplets within the cone cells to improve colour vision. In contrast, nightjars have a limited number of cone cells, either lacking or having only a few oil droplets. The nocturnal eyesight of nightjars is probably equivalent to that of owls. Although they have good hearing, European Nightjars appear not to rely on sound to find insects, and nightjars do not echolocate.
Predators and parasites
The eggs and chicks of this ground-nesting bird are vulnerable to predation by red foxes, pine martens, European hedgehogs, least weasels and domestic dogs, and by birds including crows, Eurasian Magpies, Eurasian Jays and owls. Snakes, such as common adders, may also rob the nest. Adults may be caught by birds of prey including Northern Goshawks, Hen Harriers, Eurasian Sparrowhawks, Common Buzzards, Peregrine and Sooty Falcons.
Parasites recorded on this nightjar include a single species of biting louse found on the wings, and a feather mite that occurs only on the white wing markings. Avian malaria has been recorded in this bird. The leucocytozoon blood parasite L. caprimulgi is rare in the European Nightjar. Its scarcity and the fact that it is the only one of its genus found in nightjars support the suggestion that it has crossed over from close relatives that normally infect owls.
The European population of the European Nightjar has been estimated from 470,000 to more than one million birds, suggesting a total global population of 2–6 million individuals. Although the population appear to be declining, the decrease is not rapid enough to trigger vulnerability criteria. The large numbers and huge breeding range mean that this species is therefore classified by the International Union for Conservation of Nature as being of Least Concern.
The largest breeding populations are in Russia (up to 500,000 pairs), Spain (112,000 pairs) and Belarus (60,000 pairs). There have been declines in much of the range, but especially in northwestern Europe. The loss of insect prey through pesticide use, coupled with disturbance, collision with vehicles and habitat loss have contributed to the falling population. As ground-nesting birds, they are adversely affected by disturbance, especially by domestic dogs, which may destroy the nest or advertise its presence to crows or predatory mammals. Breeding success is higher in areas with no public access; where access is permitted, and particularly where dog owners allow their pets to run loose, successful nests tend to be far from footpaths or human habitation.
In Britain and elsewhere, commercial forestry has created new habitat which has increased numbers, but these gains are likely to be temporary as the woodland develops and becomes unsuitable for nightjars. In the UK, it is red-listed as a cause for concern, and is close to extinction in Ireland.
Poets sometimes use the Nightjar as an indicator of warm summer nights, as in George Meredith's "Love in the Valley" Lone on the fir-branch, his rattle-notes unvaried/Brooding o'er the gloom, spins the brown eve-jar, Dylan Thomas's "Fern Hill" and all the night long I heard, blessed among stables, the nightjars/flying with the ricks, or Wordsworth's "Calm is the fragrant air", The busy dor-hawk chases the white moth/With burring note. Nightjars only sing when perched, and Thomas Hardy referenced the eerie silence of a hunting bird in "Afterwards": If it be in the dusk when, like an eyelid's soundless blink/The dewfall-hawk comes crossing the shades to alight/Upon the wind-warped upland thorn.
Caprimulgus and the old name "goatsucker" both refer to the myth, old even in Aristotle's day, that nightjars suckled from nanny goats, which subsequently ceased to give milk or went blind. This ancient belief is reflected in nightjar names in other European languages, such as German zeigenmelker and Italian succiacapri, which also mean goatsucker, but despite its antiquity, it has no equivalents in Arab, Chinese or Hindu traditions. It is likely that the birds were attracted by insects around domestic animals, and, as strange nocturnal creatures, were then blamed for any misfortune that befell the beast. Another old name, "puckeridge", was used to refer to both the bird and a disease of farm animals, the latter actually caused by botfly larvae under the skin.
Other old names refer to the song, "churn owl", habitat, "fern owl", diet, "dor hawk" and "moth hawk" or superstition, "lich fowl" (corpse bird). The last of these names, like "gabble ratchet" (corpse hound) may refer to the belief that the souls of unbaptised children were doomed to wander in nightjar form until Judgement Day.[a]
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|Wikimedia Commons has media related to Caprimulgus europaeus.|
- Ageing and sexing by Javier Blasco-Zumeta and Gerd-Michael Heinze
- Song at xeno-canto
- Stills images and videos at ARKive