Temporal range: 29–19Ma Late Oligocene - Early Miocene
|A skull of Daeodon shoshonensis at the Carnegie Museum of Natural History.|
Daeodon (from Greek, daios "hostile", "dreadful" and odon "teeth") is an extinct genus of entelodont artiodactyl that inhabited North America between 29 to 19 million years ago during the late Oligocene and early Miocene epochs. The type species is Daeodon shoshonensis, the last and largest of the entelodonts, known adults of this species possess skulls about 90 cm (3 ft) in length. It had a broad distribution across the United States but it was never abundant.
The genus Daeodon was erected by the American anatomist and paleontologist Edward Drinker Cope in 1878, he classified it as a perissodactyl and thought it was closeley related to "Menodus". This classification persisted until the description of "Elotherium" calkinsi in 1905, a very similar and much more complete animal from the same rocks which was promptly assigned as a species of Daeodon by Peterson (1909) which lead to its reclassification as a member of the family Entelodontidae. The exact relationships between Daeodon and other entelodonts are not well understood, some authors (Lucas et al., 1998) consider the greater morphological similarity of Daeodon to Paraentelodon rather than to earlier North American entelodonts like Archaeotherium as evidence for Daeodon being descendent from a Late Oligocene immigration of large Asian entelodonts to North America, however, the existence of distinct specimens of Archoetherium showing characters reminiscent of those present in both Paraentelodon and Daeodon rises the possibility of both genera being actually descendant from a North American common ancestor.
The type species of Daeodon is D. shohonensis, based on a fragment of a lower jaw from the John Day Formation of Oregon. Several other species were assigned to the genus in the subsequent decades, like D. calkinsi, D. mento  and D. minor . Since 1945 it had been suggested that two other taxa were actually junior synonyms of Daeodon but the formalization of this referral didn't take place until the work of Lucas et al. (1998). Ammodon leidyanum, named by Cope's rival, O. C. Marsh, and Dinohyus hollandi, a complete skeleton from the Agate Springs quarry of Nebraska, were found to be indistinguishable from each other and in turn both were indistinguishable from D. shoshonensis.  The other previously recognized species of Daeodon were also synonymized to D. shoshonensis with the exception of D. calkinsi which was tentatively excluded from Daeodon. That same year, an obscure entelodont, Boochoerus humerosum was also synonymized to Daeodon by Foss and Fremd (1998) and albeit its status as a distinct species was retained they note that the differences could still be attributed to individual or population variation or sexual dimorphism.
Daeodon shoshonensis is the largest known entelodont; known adult individuals had skulls about 90 cm (3 ft) long and were around 1.8 m (6 ft) tall at the shoulders. It's differentiated from other entelodonts by a suite of unique dental characters, the shape and relatively small size of the cheekbone flanges of its skull compared to those of Archaeotherium, the small size of its chin tubercle as well as features of its carpus and tarsus and the fusion of the bones of the lower leg. Like other entelodonts its limbs were long and slender with the bones of the foreleg fused together and only had two fingers in each foot, it also had a relatively lightly constructed neck for the size of its head, whose weight was mostly supported by muscles and tendons attached to the tall spines of the thoracic vertebrae, similar to those of modern day bison and white rhinoceros.
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