Players must get their Xbox 360 avatars through increasingly difficult obstacle courses. The game consists of four fields of Amazon, Antarctic, Egypt, and Pirate Island, each with their unique style to make the player's avatar wipe out in any way possible.
In the original Doritos Crash Course, players unlocked levels by simply playing them; the case is not the same here. In Crash Course 2, players must now collect "stars" in the game, which are used to unlock levels, buy power-ups, and have avatar effects to change their Avatar's appearance in-game. Coins are another type of token which players can buy for Microsoft Points and are used for the same purposes as stars.