This subgenus was shown on the basis of DNA analysis to be polyphyletic. A revision of this subgenus on a morphological basis by Landua et al. moved several of the species in this subgenus into a new subgenus Papernaia.
Species in the subgenus Giovanolaia have the following characteristics:
Gametocytes are elongated.
Both gametocytes and schizonts are stretched along the red cell nucleus.
Exoerythrocytic schizogony occurs in the mononuclear phagocyte system.
Species in this subgenus
- Plasmodium anasum
- Plasmodium buteonis
- Plasmodium circumflexum
- Plasmodium dissanaikei
- Plasmodium fallax
- Plasmodium ghadiriani
- Plasmodium gundersi
- Plasmodium heroni
- Plasmodium lophurae
- Plasmodium octamerium
- Plasmodium tranieri
- "Plasmodium (Giovannolaia)". NCBI Taxonomy Browser. 418112.
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