Fossil Hepialoidea appear to be few.Prohepialus (possibly Hepialidae) has been described from the about 35 million year old Bembridge Marls of Isle of Wight. A mid-Miocene hepialoid fossil is also known from China.
^Kristensen, N. P. and Skalski, A.W. (1999). Phylogeny and paleontology. Pages 7-25 in: Lepidoptera: Moths and Butterflies. 1. Evolution, Systematics, and Biogeography. Handbook of Zoology Vol. IV, Part 35. N. P. Kristensen, ed. De Gruyter, Berlin and New York
^Jarzembowski, E.A. (1980). Fossil, insects from the Bembridge Marls, Palaeogene of the Isle of Wight, southern England. Bulletin of the British Museum (Natural History) (Geology), 33: 237-293.
^Zhang, J. (1989). Fossil insects from Shanwung, Shandong, China. Shandong Science and Technology Publishing House.
Kristensen, N. P.. (1999). The homoneurous Glossata. pp. 51–63 In: Kristensen, N.P. (ed.), Lepidoptera, Moths and Butterflies, 1: Evolution, Systematics, and Biogeography. Handbook of Zoology, volume 4 (35). Walter de Gruyter. Berlin, New York.