Largest and heaviest animals

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The largest animal currently alive is the blue whale. The maximum recorded weight was 190 tonnes for a specimen measuring 27.6 metres (91 ft), whereas longer ones, up to 33.6 metres (110 ft), have been recorded but not weighed.[1][2][3] It is estimated, this individual could have a mass of 250 tonnes or more.[4][5][6] The longest non-colonial animal is the lion's mane jellyfish (37 m, 120 ft).[7]

In 2023, paleontologists estimated that the extinct whale Perucetus, discovered in Peru, may have outweighed the blue whale, with a mass of 85 to 340 t (94–375 short tons; 84–335 long tons).[8] However more recent studies suggest this whale was much smaller than previous estimates putting its weight at 60 to 113 tonnes.[9] While controversial, estimates for the weight of the sauropod Bruhathkayosaurus suggest it was around 110–170 tons, with the highest estimate being 240 tons, if scaled with Patagotitan, although actual fossil remains no longer exist, and that estimation is based on described dimensions in 1987.[10] The upper estimates of weight for these two prehistoric animals would have easily rivaled or exceeded the blue whale.

The African bush elephant (Loxodonta africana) is the largest living land animal. A native of various open habitats in sub-Saharan Africa, males weigh about 6.0 tonnes (13,200 lb) on average.[11] The largest elephant ever recorded was shot in Angola in 1974. It was a male measuring 10.67 metres (35.0 ft) from trunk to tail and 4.17 metres (13.7 ft) lying on its side in a projected line from the highest point of the shoulder, to the base of the forefoot, indicating a standing shoulder height of 3.96 metres (13.0 ft). This male had a computed weight of 10.4 tonnes.[1]

Heaviest living animals[edit]

The heaviest living animals are all whales. Since no scale can accommodate the whole body of a large whale, most have been weighed by parts.

Rank Animal Average mass
[tonnes] 		Maximum mass
[tonnes] 		Average total length
[m (ft)]
1 Blue whale[12] 110[13] 190[1] 24 (79)[14]
2 North Pacific right whale 60[15] 120[1] 15.5 (51)[13]
3 Southern right whale 58[13] 110[16] 15.25 (50)[13]
4 Fin whale 57[13] 120[16] 19.5 (64)[13]
5 Bowhead whale 54.5[13][17] 120[1] 15 (49)[13]
6 North Atlantic right whale 54[13][18] 110[16][19] 15 (49)[13][19]
7 Sperm whale 31.25[13][20] 57[21] 13.25 (43.5)[13][20]
8 Humpback whale 29[13][22] 48[23] 13.5 (44)[13]
9 Sei whale 22.5[13] 45[24] 14.8 (49)[13]
10 Gray whale 19.5[13] 45[25] 13.5 (44)[13]
Infographic showing the size of marine megafauna.
The blue whale is the heaviest living animal.

Heaviest terrestrial animals[edit]

The heaviest land animals are all mammals. The African elephant is now listed as two species, the African bush elephant and the African forest elephant, as they are now generally considered to be two separate species.[26]

Rank Animal Average mass
[tonnes] 		Maximum mass
[tonnes] 		Average total length
[m (ft)]
1 African bush elephant 6[27][28] 10.4[29] 7 (23)[30]
2 Asian elephant 4.5[1][31] 8.15[1] 6.8 (22.3)[31]
3 African forest elephant 2.7[32] 6.0[32] 6.2 (20.3)[33]
4 White rhinoceros 2[34][35] 4.5[36] 4.4 (14.4)[37]
5 Indian rhinoceros 1.9[38][39] 4.0[40] 4.2 (13.8)[41]
6 Hippopotamus 1.8[42][43] 4.5[44] 5.05 (16.5)[45]
7 Black rhinoceros 1.1[46] 2.9[47] 4 (13.1)[48]
8 Javan rhinoceros 1.75[49][50] 2.3[51] 3.8 (12.5)[52]
9 Giraffe 1.0[1] 2[53] 5.15 (16.9)[54]
10 Gaur 0.95[55] 1.5[55] 3.8 (12.5)[56]

Vertebrates[edit]

Mammals (Mammalia)[edit]

Main article: List of largest mammals
Size of Paraceratherium (dark grey) compared to a human and other rhinos (though one study suggests Palaeoloxodon namadicus may have been a larger land mammal)

The blue whale is the largest mammal of all time, with the largest known specimen being 33.6 m (110.2 ft) long and the largest weighted specimen being 190 tonnes.[12][57][58] The extinct whale species Perucetus colossus was shorter than the blue whale, at 17.0–20.1 meters (55.8–65.9 ft) but it is estimated to have rivaled or surpassed it in weight, at 85–340 tonnes. At the highest estimates, this would make Perucetus the heaviest known animal in history.[8]

The largest land mammal extant today is the African bush elephant. The largest extinct land mammal known was long considered to be Paraceratherium orgosensis, a rhinoceros relative thought to have stood up to 4.8 m (15.7 ft) tall, measured over 7.4 m (24.3 ft) long and may have weighed about 17 tonnes.[59][60] In 2015, a study suggested that one example of the proboscidean Palaeoloxodon namadicus may have been the largest land mammal ever, based on extensive research of fragmentary leg bone fossils from one individual, with a maximum estimated size of 22 tonnes.[61][59]

Stem-mammals (Synapsida)[edit]

The Late Triassic Lisowicia bojani, from what is now southern Poland, probably was the largest of all non-mammalian synapsids (most of which became extinct 250 million years ago), at 4.5 m (15 ft) in length, 2.6 m (8.5 ft) in height and 9 t (20,000 lb) in weight.[62] However, one study suggested a more conservative weight of 4.87 tonnes to 7.02 tonnes for the adult taxon, with an average body mass of 5.88 tonnes.[63] The largest carnivorous synapsid was Anteosaurus at 5–6 m (16–20 ft) and 500–600 kg (1,100–1,300 lb).[64]

Caseasaurs (Caseasauria)
The herbivorous Alierasaurus was the largest caseid and the largest amniote to have lived at the time, with an estimated length around 6–7 m (20–23 ft).[65] Another huge caseasaur is Cotylorhynchus hancocki, with an estimated length and weight of at least 6 m (20 ft)[66] and more than 500 kg (1,100 lb).[67]
Sphenacodontids (Sphenacodontidae)
The biggest carnivorous synapsid of Early Permian was Dimetrodon, which could reach 4.6 m (15 ft) and 250 kg (550 lb).[68] The largest members of the genus Dimetrodon were also the world's first fully terrestrial apex predators.[69]
Tappenosaurids (Tappenosauridae)
The Middle Permian Tappenosaurus was estimated at 5.5 m (18 ft) in length which is comparable in size with the largest dinocephalians.[70]
Therapsids (Therapsida)
The plant-eating dicynodont Lisowicia bojani is the largest-known of all non-mammalian synapsids, at 4.5 m (15 ft) and 9,000 kg (20,000 lb).[62][71][72] The largest carnivorous therapsid was the aforementioned Anteosaurus from what is now South Africa during Middle Permian epoch. It reached 5–6 m (16–20 ft) long, and about 500–600 kg (1,100–1,300 lb) in weight.[64]

Reptiles (Reptilia)[edit]

Main article: List of largest reptiles
The saltwater crocodile is the largest living reptile.

The largest living reptile, a representative of the order Crocodilia, is the saltwater crocodile (Crocodylus porosus) of Southern Asia and Australia, with adult males being typically 3.9–5.5 m (13–18 ft) long. The largest confirmed saltwater crocodile on record was 6.32 m (20.7 ft) long, and weighed about 1,360 kg (3,000 lb).[1] Unconfirmed reports of much larger crocodiles exist, but examinations of incomplete remains have never suggested a length greater than 7 m (23 ft).[73] Also, a living specimen estimated at 7 m (23 ft) and 2,000 kg (4,400 lb) has been accepted by the Guinness Book of World Records.[74] However, due to the difficulty of trapping and measuring a very large living crocodile, the accuracy of these dimensions has yet to be verified. A specimen named Lolong caught alive in the Philippines in 2011 (died February 2013) was found to have measured 6.17 m (20.2 ft) in length.[75][76][77][78][79]

The Komodo dragon (Varanus komodoensis), also known as the "Komodo monitor", is a large species of lizard found in the Indonesian islands of Komodo, Rinca, Flores, Gili Motang, Nusa kode and Padar. A member of the monitor lizard family (Varanidae), it is the largest living species of lizard, growing to a maximum length of more than 3 metres (9.8 feet) in rare cases and weighing up to approximately 166 kilograms (366 pounds).[80]

Largest living reptiles[edit]

The following is a list of the largest living reptile species ranked by average weight, which is dominated by the crocodilians. Unlike mammals, birds, or fish, the mass of large reptiles is frequently poorly documented and many are subject to conjecture and estimation.[1]

Rank Animal Average mass
[kg (lb)] 		Maximum mass
[kg (lb)] 		Average total length
[m (ft)]
1 Saltwater crocodile 450 (1,000)[81][82] 2,000 (4,400)[83][84] 4.5 (14.8)[81][85]
2 Nile crocodile 410 (900)[86] 1,090 (2,400)[1] 4.2 (13.8)[86]
3 Orinoco crocodile 380 (840)[citation needed] 1,100 (2,400)[citation needed] 4.1 (13.5)[87][88]
4 Leatherback sea turtle 364 (800)[89][90] 932 (2,050)[1] 2.0 (6.6)[1]
5 American crocodile 336 (740)[91] 1,000 (2,200)[92] 4.0 (13.1)[93][94]
6 Black caiman 300 (661)[citation needed] 1,000 (2,200)[citation needed] 3.9 (12.8)[95][96][97][98]
7 Gharial 250 (550)[99] 1,000 (2,200)[100] 4.5 (14.8)[99]
8 American alligator 240 (530)[101][102] 1,000 (2,200)[1] 3.4 (11.2)[102]
9 Mugger crocodile 225 (495)[101] 700 (1,500)[103] 3.3 (10.8)[102]
10 False gharial 210 (460)[104] 590 (1,300)[105] 4.0 (13.1)[106]
11 Aldabra giant tortoise 205 (450)[107] 360 (790)[1] 1.4 (4.6)[108]
12 Loggerhead sea turtle 200 (441)[citation needed] 545 (1,202)[citation needed] 0.95 (3.2)[108]
13 Green sea turtle 190 (418.9)[109] 395 (870.8)[86] 1.12 (3.67)[86]
14 Slender-snouted crocodile 180 (400)[110] 325 (720)[110] 3.3 (10.8)[110]
15 Galapagos tortoise 175 (390)[111] 417 (919)[112] 1.5 (4.9)[113]

Dinosaurs (Dinosauria)[edit]

Main article: Dinosaur size
See also: Largest prehistoric animals
Bruhathkayosaurus matleyi, potentially the largest animal to have walked the earth.
A human and the longest known dinosaurs of the five major clades. Each grid section represents 1 square meter.
  Sauropoda (Supersaurus vivianae)
  Ornithopoda (Shantungosaurus giganteus)
  Theropoda (Spinosaurus aegyptiacus)
  Thyreophora (Stegosaurus ungulatus)
  Marginocephalia (Triceratops prorsus)

Dinosaurs are now extinct, except for birds, which are theropods.

Sauropods (Sauropoda)
The largest dinosaurs, and the largest animals to ever live on land, were the plant-eating, long-necked Sauropoda. The tallest and heaviest sauropod known from a complete skeleton is a specimen of an immature Giraffatitan discovered in Tanzania between 1907 and 1912, now mounted in the Museum für Naturkunde of Berlin. It is 12–13.27 m (39.4–43.5 ft) tall and weighed 23.3–39.5 tonnes.[114][115][116][117][118][119] The longest is a 25 m (82 ft) long specimen of Diplodocus discovered in Wyoming, and mounted in Pittsburgh's Carnegie Natural History Museum in 1907. A Patagotitan specimen found in Argentina in 2014 is estimated to have been 37–40 m (121–131 ft) long and 20 m (66 ft) tall, with a weight of 69–77 tonnes.[120][121]
There were larger sauropods, but they are known only from a few bones. The current record-holders include Argentinosaurus, which may have weighed 100 tonnes; Supersaurus which might have reached 34 m (112 ft) in length and Sauroposeidon which might have been 18 m (59 ft) tall. Some abnormal specimens such as specimen BYU 9024 of the Barosaurus genus could reach an astounding 45-48 meters long, with mass varying from the 'modest' 60-66 tons to the more immense 92-120 tons.[122][123][124] Two other such sauropods include Bruhathkayosaurus and Maraapunisaurus. Both are known only from fragments that no longer exist. Bruhathkayosaurus might have been between 40–44 m (131–144 ft) in length and 175–220 tonnes in weight according to some estimates, with recent estimates being place between 110-170 tons.[125][10] Maraapunisaurus might have been approximately 35–40 m long and 80–120 tonnes or more.[126] Each of these two 'super-sauropods' would have easily rivalled the largest blue whale in size.[10][12]
Rank Animal Average mass
[tonnes] 		Maximum mass
[tonnes] 		Average total length
[m (ft)]
1 Bruhathkayosaurus matleyi 110-170[10] 240[10] 44.1 (144.6)[127][128]
2 Maraapunisaurus fragillimus 80-120[129] 150[130] 35-40 (115-131)[129]
3 Barosaurus lentus BYU 9024 60-66[122][123][131][132] 92-120[124] 45-48 (148-157)[132]
4 Argentinosaurus huinculensis 75-80[117] 100[117][133] 35–39.7 (115–130)[117][134]
5 Mamenchisaurus 50-80[135] 80[135] 26–35 (85–115)[135]
6 Notocolossus gonzalezparejasi 44.9–75.9[133] 75.9[133] 28 (92)[133]
7 Patagotitan mayorum 55-69[136] 77[136] 33–37 (108–121)[136]
8 Puertasaurus reuili 50-60[137] 60 27-30 (89-98)[138][139][140]
9 Sauroposeidon proteles 40-60[141] 60[141] 27–34 (89–112)[141][137][142]
10 Dreadnoughtus schrani 22.1–59.3[143] 59.3[143] 26 (85)[133][143]
Theropods (Theropoda)
The largest theropod known from a nearly complete skeleton is the most complete Tyrannosaurus rex specimen, nicknamed "Sue", which was discovered in South Dakota in 1990 and now mounted in the Field Museum of Chicago at a total length of 12.3 m (40 ft). Body mass estimates have reached over 9,500 kg,[144] though other figures, such as Hartman's 2013 estimate of 8,400 kg,[145] have been lower.
Another giant theropod is Spinosaurus aegyptiacus from the mid-Cretaceous of North Africa. Size estimates have been fluctuating far more over the years, with length estimates ranging from 12.6 to 18 m and mass estimates from 7 to 20.9 t.[146][147] Recent findings favor a length exceeding 15 m [148] and a body mass of 7.5 tons.[149]
Other contenders known from partial skeletons include Giganotosaurus carolinii (est. 12.2–13.2 m and 6-13.8 tonnes) and Carcharodontosaurus saharicus (est. 12-13.3 m and 6.2-15.1 tonnes).[147][150][151][152][153][122]
The largest extant theropod is the common ostrich (see birds, below).
Armored dinosaurs (Thyreophora)
The largest thyreophorans were Ankylosaurus and Stegosaurus, from the Late Cretaceous and Late Jurassic periods (respectively) of what is now North America, both measuring up to 9 m (30 ft) in length and estimated to weigh up to 6 tonnes.[154][155]
Ornithopods (Ornithopoda)
The largest ornithopods were the hadrosaurids Shantungosaurus, a late Cretaceous dinosaur found in the Shandong Peninsula of China, and Magnapaulia from the late Cretaceous of North America. Both species are known from fragmentary remains but are estimated to have reached over 15 m (49 ft) in length[156][157] and were likely the heaviest non-sauropod dinosaurs, estimated at over 23 tonnes.[157]
Ceratopsians (Ceratopsia)
The largest ceratopsians were Triceratops and its ancestor Eotriceratops from the late Cretaceous of North America. Both estimated to have reached about 9 m (30 ft) in length[158] and weighed 12 tonnes.[159][160]

Birds (Aves)[edit]

Main article: List of largest birds
The common ostrich is the largest living bird.

The largest living bird, a member of the Struthioniformes, is the common ostrich (Struthio camelus), from the plains of Africa. A large male ostrich can reach a height of 2.8 m (9.2 ft) and weigh over 156 kg (344 lb).[161] A mass of 200 kg (440 lb) has been cited for the common ostrich but no wild ostriches of this weight have been verified.[162] Eggs laid by the ostrich can weigh 1.4 kg (3.1 lb) and are the largest eggs in the world today.[citation needed]

The largest bird in the fossil record may be the extinct elephant birds (Aepyornithidae) of Madagascar, which were related to the kiwis. Aepyornis exceeded 3 m (9.8 ft) in height and 500 kg (1,100 lb), while Vorombe could reach a similar height and a mass of 732 kg (1,614 lb).[163] The last of the elephant birds became extinct about 300 years ago. Of almost exactly the same upper proportions as the largest elephant birds was Dromornis stirtoni of Australia, part of a 26,000-year-old group called mihirungs of the family Dromornithidae.[164] The largest carnivorous bird was Brontornis, an extinct flightless bird from South America which reached a weight of 350 to 400 kg (770 to 880 lb) and a height of about 2.8 m (9 ft 2 in).[165] The tallest carnivorous bird was Kelenken, which could reach 3 to 3.2 meters in height and 220 to 250 kilograms. The tallest bird ever was the giant moa (Dinornis maximus), part of the moa family of New Zealand that went extinct around 1500 AD. This particular species of moa stood up to 3.7 m (12 ft) tall,[161] but weighed about half as much as a large elephant bird or mihirung due to its comparatively slender frame.[1]

The heaviest bird ever capable of flight was Argentavis magnificens, the largest member of the now extinct family Teratornithidae, found in Miocene-aged fossil beds of Argentina, with a wingspan up to 5.5 m (18 ft), a length of up to 1.25 m (4.1 ft), a height on the ground of up to 1.75 m (5.7 ft) and a body weight of at least 71 kg (157 lb).[1][166][167] Pelagornis sandersi is thought to have had an even larger wingspan of about 6.1–7.4 m (20–24 ft), but is only about 22–40 kg (49–88 lb), half the mass of the former.[168][167]

Heaviest living bird species[edit]

The following is a list of the heaviest living bird species based on maximum reported or reliable mass, but average weight is also given for comparison. These species are almost all flightless, which allows for these particular birds to have denser bones and heavier bodies. Flightless birds comprise less than 2% of all living bird species.[citation needed]

Rank Animal Binomial Name Average mass
[kg (lb)] 		Maximum mass
[kg (lb)] 		Average total length
[cm (ft)] 		Flighted
1 Common ostrich Struthio camelus 104 (230)[169] 156.8 (346)[169] 210 (6.9)[170] No
2 Somali ostrich Struthio molybdophanes 90 (200)[169] 130 (287)[citation needed] 200 (6.6)[169] No
3 Southern cassowary Casuarius casuarius 45 (99)[169] 85 (190)[171] 155 (5.1)[169] No
4 Northern cassowary Casuarius unappendiculatus 44 (97)[169] 75 (170)[169] 149 (4.9)[170] No
5 Emu Dromaius novaehollandiae 33 (73)[169][172] 70 (150)[citation needed] 153 (5)[169] No
6 Emperor penguin Aptenodytes forsteri 31.5 (69)[170][173] 46 (100)[170] 114 (3.7)[170] No
7 Greater rhea Rhea americana 23 (51)[172] 40 (88)[170] 134 (4.4)[169] No
8 Domestic turkey/wild turkey Meleagris gallopavo 13.5 (29.8) [174] 39 (86)[175] 100 - 124.9 (3.3 – 4.1)[citation needed] Yes
9 Dwarf cassowary Casuarius bennetti 19.7 (43)[169] 34 (75)[169] 105 (3.4)[citation needed] No
10 Lesser rhea Rhea pennata 19.6 (43)[169] 28.6 (63)[169] 96 (3.2)[170] No
11 Mute swan Cygnus olor 11.87 (26.2) 23 (51) 100-130 (3.3 - 4.3)[176] Yes
12 Great bustard Otis tarda 10.6 (23.4)[citation needed] 21 (46)[1] 115 (3.8)[citation needed] Yes
13 King penguin Aptenodytes patagonicus 13.6 (30)[170][173] 20 (44)[177] 92 (3)[citation needed] No
14 Kori bustard Ardeotis kori 11.4 (25.1)[170] 20 (44.1)[citation needed] 150 (5)[170] Yes
15 Trumpeter swan Cygnus buccinator 11.6 (25.1) 17.2 (38) 138 - 165 (4.5 - 5.4) Yes
16 Wandering albatross Diomedea exulans 11.9 (24) 16.1 (38)[178] 107 - 135 (3.5 - 4.4) Yes
17 Whooper swan Cygnus cygnus 11.4 (25) 15.5 (32) 140 - 165 (4.5 - 5.4) Yes
18 Dalmatian Pelican Pelecanus crispus 11.5 (25) 15 (33.1)[citation needed] 183 (6)[citation needed] Yes
19 Andean condor Vultur gryphus 11.3 (25)[176] 14.9 (33)[176] 100 - 130 (3.3 - 4.3)[176] Yes

Amphibians (Amphibia)[edit]

A giant salamander, the largest living amphibian.

The largest living amphibian is the South China giant salamander (Andrias sligoi). Formerly considered conspecific with the Chinese giant salamander (A. davidianus), the maximum size of this nearly human-sized river-dweller is 64 kg (141 lb) and almost 1.83 m (6.0 ft).[1] Before amniotes became the dominant tetrapods, several giant amphibian proto-tetrapods existed and were certainly the dominant animals in their ecosystems. The largest known was the crocodile-like Prionosuchus, which reached a length of 9 m (30 ft).[179]

Frogs (Anura)
The massive and destructive cane toad ranks as the largest toad in the world.
The largest member of the largest order of amphibians is the African goliath frog (Conraua goliath). The maximum size this species is verified to attain is a weight of 3.8 kg (8.4 lb) and a snout-to-vent length of 39 cm (15 in).[1] The largest of the toads, the cane toad (Rhinella marina), is also the second largest member of the frog order. This infamous, often invasive species can grow to maximum mass of 2.65 kg (5.8 lb) and measure a maximum of 33 cm (13 in) from snout-to-vent.[1] Rivaling the previous two species, the African bullfrog (Pyxicephalus adspersus) can range up to a weight of 2 kg (4.4 lb) and 25.5 cm (10.0 in) from snout to vent.[180] Another large frog is the largest frog in North America, the American bullfrog, which can reach weights of up to 0.8 kg (1.8 lb) and snout-to-vent-length (SVL) of 20 cm (7.9 in). However, the toad Beelzebufo ampinga, found in fossil from the Cretaceous era in what is now Madagascar, was estimated to grow to 41 cm (16 in) long and weigh up to 4.5 kg (9.9 lb), making it the largest frog ever known.[181] But in more recent studies, animals of this species have been estimated to have grown to at least 23.2 cm (9.1 in) (snout-vent length), which is around the size a modern African bullfrog can reach.[182] The largest tree frog is the Australasian white-lipped tree frog (Litoria infrafrenata), the females of which can reach a length of 14 cm (5.5 in) from snout to vent and can weigh up to 115 g (4.1 oz).[183] The family Leptodactylidae, one of the most diverse anuran families, also has some very large members. The largest is the Surinam horned frog (Ceratophrys cornuta), which can reach 20 cm (7.9 in) in length from snout to vent and weigh up to 0.48 kg (1.1 lb).[184] While not quite as large as Ceratophrys cornuta, Leptodactylus pentadactylus is often heavier; it can reach 18.5 cm (7.3 in) long and weigh 0.60 kilograms (1.3 pounds). The largest dendrobatid is the Colombian golden poison frog (Phyllobates terribilis), which can attain a length of 6 cm (2.4 in) and nearly 28.3 g (1.00 oz).[185] Most frogs are classified under the suborder Neobatrachia, although nearly 200 species are part of the suborder Mesobatrachia, or ancient frogs. The largest of these are the little-known Brachytarsophrys or Karin Hills frogs, of South Asia, which can grow to a maximum snout-to-vent length of 17 cm (6.7 in) and a maximum weight of 0.54 kg (1.2 lb).[186]
Caecilians (Gymnophiona)
The largest of the worm-like caecilians is the Colombian Thompson's caecilian (Caecilia thompsoni), which reaches a length of 1.5 m (4.9 ft), a width of about 4.6 cm (1.8 in) and can weigh up to about 1 kg (2.2 lb).[1]
Salamanders (Urodela)
Besides the previously mentioned Chinese and South China giant salamanders, the closely related Japanese giant salamander (Andrias japonicus) is also sometimes cited as the largest living amphibian, but salamanders of a greater size than 1.53 m (5.0 ft) and 36 kg (79 lb) have never been verified for this species. Another giant of the amphibian world is the North American hellbender (Cryptobranchus alleganiensis), which can measure up to 0.76 m (2.5 ft). The recently described reticulated siren of the southeastern United States rivals the hellbender in size, although it is more lean in build.[187] The largest of the newts is the Iberian ribbed newt (Pleurodeles waltl), which can grow up to 30 cm (12 in) in length.[188]

Fish[edit]

Main article: List of largest fish

Invertebrate chordates[edit]

Tunicates (Tunicata)[edit]

Further information: Tunicata

The largest tunicates are Synoicum pulmonaria, found at depths of 20 and 40 metres (66 and 131 ft), and are up to 14 centimetres (6 in) in diameter. It is also present in the northwestern Atlantic Ocean, around the coasts of Greenland and Newfoundland, but is less common here than in the east, and occurs only at depths between 10 and 13 metres (33 and 43 ft).[189]

Entergonas (Enterogona)
The largest entergonas Synoicum pulmonaria it is usually found at depths between about 20 and 40 metres (66 and 131 ft) and can grow to over a metre (yard) in length. It is also present in the northwestern Atlantic Ocean, around the coasts of Greenland and Newfoundland, but is less common here than in the east, and occurs only at depths between 10 and 13 metres (33 and 43 ft).[189]
Pleurogonas (Pleurogona)
The largest pleurogonas: Pyura pachydermatina .[190] In colour it is off-white or a garish shade of reddish-purple. The stalk is two thirds to three quarters the length of the whole animal which helps distinguish it from certain invasive tunicates not native to New Zealand such as Styela clava and Pyura stolonifera.[191] It is one of the largest species of tunicates and can grow to over a metre (yard) in length.[192]
Aspiraculates (Aspiraculata)
The largest aspiraculates: Oligotrema large and surrounded by six large lobes; the cloacal syphon is small. They live exclusively in deep water and range in size from less than one inch (2 cm) to 2.4 inches (6 cm).

Thaliacea[edit]

The Pyrosoma atlanticum is the largest thaliacean.

The largest thaliacean, Pyrosoma atlanticum, is cylindrical and can grow up to 60 cm (2 ft) long and 4–6 cm wide. The constituent zooids form a rigid tube, which may be pale pink, yellowish, or bluish. One end of the tube is narrower and is closed, while the other is open and has a strong diaphragm. The outer surface or test is gelatinised and dimpled with backward-pointing, blunt processes. The individual zooids are up to 8.5 mm (0.33 in) long and have a broad, rounded branchial sac with gill slits. Along the side of the branchial sac runs the endostyle, which produces mucus filters. Water is moved through the gill slits into the centre of the cylinder by cilia pulsating rhythmically. Plankton and other food particles are caught in mucus filters in the processes as the colony is propelled through the water. P. atlanticum is bioluminescent and can generate a brilliant blue-green light when stimulated.[193][194]

Doliolida (Doliolida)
The largest doliolida: Doliolida [195] The doliolid body is small, typically 1–2 cm long, and barrel-shaped; it features two wide siphons, one at the front and the other at the back end, and eight or nine circular muscle strands reminiscent of barrel bands. Like all tunicates, they are filter feeders. They are free-floating; the same forced flow of water through their bodies with which they gather plankton is used for propulsion - not unlike a tiny ramjet engine. Doliolids are capable of quick movement. They have a complicated lifecycle consisting of sexual and asexual generations. They are nearly exclusively tropical animals, although a few species are found as far north as northern California.[citation needed]
Salps (Salpida)
The largest salps: Cyclosalpa bakeri 15cm (6ins) long. There are openings at the anterior and posterior ends of the cylinder which can be opened or closed as needed. The bodies have seven transverse bands of muscle interspersed by white, translucent patches. A stolon grows from near the endostyle (an elongated glandular structure producing mucus for trapping food particles). The stolon is a ribbon-like organ on which a batch of aggregate forms of the animal are produced by budding. The aggregate is the second, colonial form of the salp and is also gelatinous, transparent and flabby. It takes the shape of a radial whorl of individuals up to about 20cm (4in) in diameter. It is formed of approximately 12 zooids linked side by side in a shape that resembles a crown.[193][196] are largest thetyses: Thetys vagina Individuals can reach up to 30 cm (12 in) long.[citation needed]
Larvaceans (Larvacea)
The largest larvaceans: Appendicularia 1 cm (0.39 in) in body length (excluding the tail).[citation needed]

Cephalochordates (Leptocardii)[edit]

Further information: Leptocardii

The largest lancelet is the European lancelet (Branchiostoma lanceolatum) "primitive fish". It can grow up to 6 cm (2.5 in) long.[197]

Invertebrate non-chordates[edit]

Echinoderms (Echinodermata)[edit]

At up to 3 m (9.8 ft) long, the sea cucumber Synapta maculata is the longest known echinoderm.

The largest species of echinoderm in terms of bulk is probably the starfish species Thromidia gigas, of the class Asteroidea, which reaches a weight of over 6 kg (13 lb),[198] but it might be beaten by some giant sea cucumbers such as Thelenota anax. However, at a maximum span of 63 cm (25 in), Thromidia gigas is quite a bit shorter than some other echinoderms.[1] The longest echinoderm known is the conspicuous sea cucumber Synapta maculata, with a slender body that can extend up to 3 m (9.8 ft). In comparison, the biggest sea star is the brisingid sea star Midgardia xandaros, reaching a span of 1.4 m (4.6 ft), despite being quite slender.[1] Evasterias echinosoma is another giant echinoderm and can measure up to 1 m (3.3 ft) across and weigh 5.1 kg (11 lb).[1]

Crinoids (Crinoidea)
The largest species of crinoid is the unstalked feather-star Heliometra glacialis, reaching a total width of 78 cm (31 in) and an individual arm length of 35 cm (14 in). A width of 91.4 cm (36.0 in) was claimed for one unstalked feather-star but is not confirmed.[1] The genus Metacrinus has a stalk span of 61 cm (24 in) but, due to its bulk and multiple arms, it is heavier than Heliometra.[1] In the past, crinoids grew much larger, and stalk lengths up to 40 m (130 ft) have been found in the fossil record.[199]
Sea urchins and allies (Echinoidea)
The largest sea urchin is the species Sperosoma giganteum from the deep northwest Pacific Ocean, which can reach a shell width of about 30 cm (12 in).[200] Another deep sea species Hygrosoma hoplacantha is only slightly smaller.[200] The largest species found along the North America coast is the Pacific red sea urchin (Mesocentrotus franciscanus) where the shell can reach 19 cm (7.5 in).[201] If the spines enter into count, the biggest species may be a Diadematidae like Diadema setosum, with a test up to 10 cm (3.9 in) only, but its spines can reach up to 30 cm (12 in) in length.[202]
Sea cucumbers (Holothuroidea)
The bulkiest species of sea cucumber are Stichopus variegatus and Thelenota anax, weighing several pounds, being about 21 cm (8.3 in) in diameter, and reaching a length of 1 m (3.3 ft) when fully extended. Synapta maculata can reach an extended length of 3 m (9.8 ft), but is extremely slender (3-5cm) and weigh much less than Stichopodids.[1]
Brittle stars (Ophiuroidea)
The largest known specimen of brittle star is the basket star Astrotoma agassizii. This species can grow to have a span of 1 m (3.3 ft).[1] Sometimes, Gorgonocephalus stimpsoni is considered the largest but the maximum this species is can measure 70 cm (28 in) and a disk diameter of about 14.3 cm (5.6 in). Outside from euryalids, the biggest ophiurid brittle star may be Ophiopsammus maculata (6–7 inches).[203]
Sea stars (Asteroidea)
The heaviest sea star is Thromidia gigas from the Indo-Pacific, which can surpass 6 kg (13 lb) in weight, but only has a diameter of about 65 cm (2.13 ft).[198][200] Despite its relatively small disk and weight, the long slender arms of Midgardia xandaros from the Gulf of California makes it the sea star with the largest diameter at about 1.4 m (4.5 ft).[200] Mithrodia clavigera may also become wider than 1 m (39 in) in some cases, with stout arms.[citation needed]

Flatworms (Platyhelminthes)[edit]

Further information: Platyhelminthes
Monogenean flatworms (Monogenea)
The largest known members of this group of very small parasites are among the genus of capsalids, Listrocephalos, reaching a length of 2 cm (0.79 in).[204]
Flukes (Trematoda)
The largest known species of fluke is Fasciolopsis buski, which most often attacks humans and livestock. One of these flukes can be up to 7.5 cm (3.0 in) long and 2 cm (0.79 in) thick.[205]
Tapeworms (Cestoda)
The largest known species of tapeworm is the whale tapeworm, Polygonoporus giganticus, which can grow to over 30 m (98 ft).[206][207]

Segmented worms (Annelida)[edit]

The largest of the segmented worms (including earthworms, leeches, and polychaetes) is the African giant earthworm (Microchaetus rappi). Although it averages about 1.36 m (4.5 ft) in length, this huge worm can reach a length of as much as 6.7 m (22 ft) and can weigh over 1.5 kg (3.3 lb).[208] Only the giant Gippsland earthworm, Megascolides australis, and a few giant polychaetes, including the notorious Eunice aphroditois, reach nearly comparable sizes, reaching 4 and 3.6 m (13 and 12 ft), respectively.[1]

Ribbon worms (Nemertea)[edit]

The largest nemertean is the bootlace worm, Lineus longissimus. A specimen found washed ashore on a beach in St. Andrews, Scotland in 1864 was recorded at a length of 55 m (180 ft).[209]

Mollusks (Mollusca)[edit]

A 7 m (23 ft) giant squid, the second largest of all invertebrates, encased in ice in the Melbourne Aquarium.

Both the largest mollusks and the largest of all invertebrates (in terms of mass) are the largest squids. The colossal squid (Mesonychoteuthis hamiltoni) is projected to be the largest invertebrate.[210] Current estimates put its maximum size at 12 to 14 m (39 to 46 ft) long and 750 kg (1,650 lb),[211] based on analysis of smaller specimens. In 2007, authorities in New Zealand announced the capture of the largest known colossal squid specimen. It was initially thought to be 10 m (33 ft) and 450 kg (990 lb). It was later measured at 4.2 m (14 ft) long and 495 kg (1,091 lb) in weight. The mantle was 2.5 m (8.2 ft) long when measured.[212][213]

The giant squid (Architeuthis dux) was previously thought to be the largest squid, and while it is less massive and has a smaller mantle than the colossal squid, it may exceed the colossal squid in overall length including tentacles. One giant squid specimen that washed ashore in 1878 in Newfoundland reportedly measured 16.8 m (55 ft) in total length (from the tip of the mantle to the end of the long tentacles), head and body length 6.1 m (20 ft), 4.6 m (15 ft) in circumference at the thickest part of mantle, and weighed about 900 kg (2,000 lb). This specimen is still often cited as the largest invertebrate that has ever been examined.[1][214][215] However, no animals approaching this size have been scientifically documented and, according to giant squid expert Steve O'Shea, such lengths were likely achieved by greatly stretching the two tentacles like elastic bands.[216]

Aplacophorans (Aplacophora)
The largest known of these worm-like, shell-less mollusks are represented in the genus Epimenia, which can reach 30 cm (12 in) long. Most aplacophorans are less than 5 cm (2.0 in) long.[217]
Chitons (Polyplacophora)
The largest of the chitons is the gumboot chiton, Cryptochiton stelleri, which can reach a length of 33 cm (13 in) and weigh over 2 kg (4.4 lb).[218]
The mouth of a mostly hidden giant clam, the largest bivalve
Bivalves (Bivalvia)
The largest of the bivalve mollusks is the giant clam, Tridacna gigas. Although even larger sizes have been reported for this passive animal, the top verified size was for a specimen from the Great Barrier Reef. This creature weighed 270 kg (600 lb), had an axial length of 1.14 m (3.7 ft) and depth of 0.75 m (2.5 ft).[1] The largest bivalve ever was Platyceramus platinus, a Cretaceous giant that reached an axial length of up to 3 m (nearly 10 ft).[219]
Gastropods (Gastropoda)
The "largest" of this most diverse and successful mollusk class of slugs and snails can be defined in various ways.
The living gastropod species that has the largest (longest) shell is Syrinx aruanus with a maximum shell length of 0.91 m (3.0 ft), a weight of 18 kg (40 lb) and a width of 96 cm (38 in).[220][221] Another giant species is Melo amphora, which in a 1974 specimen from Western Australia, measured 0.71 m (2.3 ft) long, had a maximum girth of 0.97 m (3.2 ft) and weighed 16 kg (35 lb).[1]
The largest shell-less gastropod is the giant black sea hare (Aplysia vaccaria) at 0.99 m (3.2 ft) in length and almost 14 kg (31 lb) in weight.
The largest of the land snails is the giant African snail (Achatina achatina) at up to 1 kg (2.2 lb) and 35 cm (14 in) long.
Cephalopods (Cephalopoda)
(See Cephalopod size.) While generally much smaller than the giant Architeuthis and Mesonychoteuthis, the largest of the octopuses, the giant Pacific octopus (Enteroctopus dofleini), can grow to be very large. The largest confirmed weight of a giant octopus is 74 kg (163 lb),[222] with a 7 m (23 ft) arm span (with the tentacles fully extended) and a head-to-tentacle-tip length of 3.9 m (13 ft).[223] Specimens have been reported up to 125 kg (276 lb) but are unverified. A weight of 10 - 50kg is a much more common size.[1]

Roundworms (Nematoda)[edit]

The largest roundworm, Placentonema gigantissima,[224] is a parasite found in the placentas of sperm whales which can reach up to 9 m (30 ft) in length.[225]

Velvet worms (Onychophora)[edit]

The largest velvet worm known is Solórzano's velvet worm (Peripatus solorzanoi). An adult female was recorded to have a body length of 22 cm (approximately 8.7 in).[226]

Arthropods (Arthropoda)[edit]

Japanese spider crab.

The largest arthropod known to have existed is the eurypterid (sea scorpion) Jaekelopterus, reaching up to 2.5 m (8.2 ft) in body length, followed by the millipede relative Arthropleura at around 2.1 m (6.9 ft) in length.[227] Among living arthropods, the Japanese spider crab (Macrocheira kaempferi) is the largest in overall size, the record specimen, caught in 1921, had an extended arm span of 3.8 m (12 ft) and weighed about 19 kg (42 lb).[1] The heaviest is the American lobster (Homarus americanus), the largest verified specimen, caught in 1977 off of Nova Scotia weighed 20 kg (44 lb) and its body length was 1.1 m (3.6 ft).[1] The largest land arthropod and the largest land invertebrate is the coconut crab (Birgus latro), up to 40 cm (1.3 ft) long and weighing up to 4 kg (8.8 lb) on average. Its legs may span 1 m (3.3 ft).[1]

Arachnids (Arachnida)[edit]

Both spiders and scorpions include contenders for the largest arachnids.

Giant huntsman spider
Spiders (Araneae)
The largest species of arachnid by length is probably the giant huntsman spider (Heteropoda maxima) of Laos, which in 2008 replaced the Goliath birdeater (Theraphosa blondi) of northern South America as the largest known spider by leg-span.[228] However the most massive arachnids, of comparable dimensions and possibly even greater mass, are the Chaco golden knee (Grammostola pulchripes), and the Brazilian salmon pink (Lasiodora parahybana). The huntsman spider may span up to 29 cm (11 in) across the legs, while in the New World tarantulas like Theraphosa can range up to 26 cm (10 in).[1] In Grammostola, Theraphosa and Lasiodora, the weight is projected to be up to at least 150 g (5.3 oz) and body length is up to 10 cm (3.9 in).[229]
Scorpions (Scorpiones)
The largest of the scorpions is the species Heterometrus swammerdami of the Indian subcontinent, which have a maximum length of 29.2 cm (11.5 in) and weigh around 60 g (2.1 oz). Another extremely large scorpion is the African emperor scorpion (Pandinus imperator), which can weigh 57 g (2.0 oz) but is not known to exceed a length of 23 cm (9.1 in).[1] However, they were dwarfed by Pulmonoscorpius kirktonensis, a giant extinct species of scorpion from Scotland, at an estimated length of 0.7 m (2.3 ft),[230] and the aquatic Brontoscorpio, at up to 94 cm (3.08 ft) which is only known from a free finger.[231]
Pseudoscorpions (Pseudoscorpiones)
The largest pseudoscorpion is Garypus titanius, from Ascension island, which can be 12 mm (0.47 in) long.[232]

Crustaceans (Crustacea)[edit]

The largest crustacean is the Tasmanian giant crab (Pseudocarcinus gigas), with a weight of 13 kilograms (29 lb) and a carapace width of up to 46 centimetres (18 in). It is the only species in the genus Pseudocarcinus.[233] Males reach more than twice the size of females.[234] At a length of up to 40 centimetres (16 in), Lysiosquillina maculata is the largest mantis shrimp in the world.[235] Tasmanian giant freshwater crayfish (Astacopsis gouldi) 5 kilograms (11 lb) in weight and over 80 centimetres (31 in) long have been known in the past, but now, even individuals over 2 kilograms (4.4 lb) are rare.[236] The species is only found in Tasmanian rivers flowing north into the Bass Strait below 400 metres (1,300 ft) above sea level, and is listed as an endangered species on the IUCN Red List.[237]

Branchiopods (Branchiopoda)
The largest of these primarily freshwater crustaceans is probably Branchinecta gigas, which can reach a length 10 cm (3.9 in).[238]
Barnacles and allies (Maxillopoda)
The largest species is Pennella balaenopterae, a copepod and ectoparasite specialising in parasitising marine mammals. The maximum size attained is 32 cm (about 13 in).[239] The largest of the barnacles is the giant acorn barnacle, Balanus nubilis, reaching 7 cm (2.8 in) in diameter and 12.7 cm (5.0 in) high.[240]
Ostracods (Ostracoda)
The largest living representative of these small and little-known but numerous crustaceans is the species Gigantocypris australis females of which reaching a maximum length of 3 cm (1.2 in).
Amphipods, isopods, and allies (Peracarida)
Giant isopod
The largest species is the giant isopod (Bathynomus pergiganteus), which can reach a length of 45 cm (18 inches) and a weight of 1.7 kg (3.7 lb).[241]
Remipedes (Remipedia)
The largest of these cave-dwelling crustaceans is the species Godzillius robustus, at up to 4.5 cm (1.8 in).[242]

Horseshoe crabs (Xiphosura)[edit]

The four modern horseshoe crabs are of roughly the same sizes, with females measuring up to 60 cm (2.0 ft) in length and 5 kg (11 lb) in weight.[243]

Sea spiders (Pycnogonida)[edit]

The largest of the sea spiders is the deep-sea species Colossendeis colossea, attaining a leg span of nearly 60 cm (2.0 ft).[244]

Trilobites (Trilobita)[edit]

Some of these extinct marine arthropods exceeded 60 cm (24 in) in length. A nearly complete specimen of Isotelus rex from Manitoba attained a length over 70 cm (28 in), and an Ogyginus forteyi from Portugal was almost as long. Fragments of trilobites suggest even larger record sizes. An isolated pygidium of Hungioides bohemicus implies that the full animal was 90 cm (35 in) long.[245][246]

Myriapods (Myriapoda)[edit]

Centipedes (Chilopoda)
Scolopendra gigantea
The biggest of the centipedes is Scolopendra gigantea of the neotropics, reaching a length of 33 cm (13 in).[247]
Millipedes (Diplopoda)
Two species of millipede both reach a very large size: Archispirostreptus gigas of East Africa and Scaphistostreptus seychellarum, endemic to the Seychelles islands. Both of these species can slightly exceed a length of 28 cm (11 in) and measure over 2 cm (0.79 in) in diameter.[1] The largest ever known was the Arthropleura, a gigantic prehistoric specimen that reached nearly 189 cm (74 in).
Symphylans (Symphyla)
The largest known symphylan is Hanseniella magna, originating in Tasmanian caves, which can reach lengths from 25 mm (0.98 in) up to 30 mm (1.2 in).[248]

Insects (Insecta)[edit]

Goliath beetle

Insects, a class of Arthropoda, are easily the most numerous class of organisms, with over one million identified species, and probably many undescribed species. The heaviest insect is almost certainly a species of beetle, which incidentally is the most species-rich order of organisms. Although heavyweight giant wetas (Deinacrida heteracantha) are known, the elephant beetles of Central and South America, (Megasoma elephas) and (M. actaeon), the Titan beetle (Titanus giganteus) of the neotropical rainforest or the Goliath beetles, (Goliathus goliatus) and (G. regius), of Africa's rainforest are thought to reach a higher weight.[249] The most frequently crowned are the Goliath beetles, the top known size of which is at least 100 g (3.5 oz) and 11.5 cm (4.5 in).[1] The elephant beetles and titan beetle can reach greater lengths than the Goliath, at up to 13.1 and 15.2 cm (5.2 and 6.0 in), respectively, but this is in part thanks to their rather large horns. The Goliath beetle's wingspan can range up to 25 cm (9.8 in).[1]

Some moths and butterflies have much larger areas than the heaviest beetles, but weigh a fraction as much.

The longest insects are the stick insects, see below.

Representatives of the extinct dragonfly-like order Meganisoptera, such as the Carboniferous Meganeura monyi of what is now France and the Permian Meganeuropsis permiana of what is now North America, are the largest insect species known to have existed. These creatures had a wingspan of some 75 cm (30 in) and a mass of over 1 pound (450 g), making them about the size of a crow.[1]

Cockroaches and termites (Blattodea)
Giant burrowing cockroach
The largest cockroach by body mass is the Australian giant burrowing cockroach (Macropanesthia rhinoceros), also known as the rhinoceros cockroach. This species can attain a length of 8.3 cm (3.3 in) and a weight of 36 g (1.3 oz). It does not have wings.[250] The Brazilian giant cockroach (Blaberus giganteus) of the neotropics reaches greater sizes of up to 10 cm in length and 15 cm in wingspan, although it is not as massive and heavy as the burrowing species. The termites, traditionally classified in their own order (Isoptera), have recently been re-considered to belong in Blattodea. The largest of the termites is the African species Macrotermes bellicosus. The queen of this species can attain a length of 14 cm (5.5 in) and breadth of 5.5 cm (2.2 in) across the abdomen; other adults, on the other hand, are about a third of the size.[1]
Beetles (Coleoptera)
The beetles are the largest order of organisms on earth, with about 400,000 species so far identified. The most massive species are the Goliathus, Megasoma and Titanus beetles already mentioned. Another fairly large species is the Hercules beetle (Dynastes hercules) of the neotropic rainforest with a maximum overall length of at least 19 cm (7.5 in) including the extremely long pronotal horn. The weight in this species does not exceed 16.5 g (0.58 oz).[1] The longest overall beetle is a species of longhorn beetle, Batocera wallacei, from New Guinea, which can attain a length of 26.6 cm (10.5 in), about 19 cm (7.5 in) of which is comprised by the long antennae.[1]
Earwigs (Dermaptera)
Since 1798, the largest of the earwigs has been the Saint Helena giant earwig (Labidura herculeana), endemic to the island of its name, measuring up to 8 cm (3.1 in) in length.[251] As of 2014, with the declaring of the organism extinct by the IUCN,[252] this may no longer be the case, although some believe a small number individuals are still extant.[253]
True flies (Diptera)
Gauromydas heros, the largest fly
The largest species of this order, which includes the common housefly, is the neotropical species Gauromydas heros, which can reach a length of 6 cm (2+3⁄8 in) and a wingspan of 10 cm (3.9 in).[1] Species of crane fly, the largest of which is Holorusia brobdignagius, can attain a length of 23 cm (9.1 in) but are extremely slender and much lighter in weight than Gauromydas.
Mayflies (Ephemeroptera)
The largest mayflies are members of the genus Proboscidoplocia from Madagascar. These insects can reach a length of 7 cm (2.8 in).[254]
True bugs (Hemiptera)
Giant water bug walking over land
The largest species of this diverse order is usually listed as the giant water bug in the genus Lethocerus, with L. maximus from the Neotropics being the absolutely largest.[255] They can surpass 12 cm (4.7 in) in length,[256] with some suggesting that the maximum size is 15 cm (5.9 in).[257] It is more slender and less heavy than most other insects of this size (principally the huge beetles). The largest cicada is Megapomponia imperatoria, which has a head-body length of about 7 cm (2.8 in) and a wingspan of 18–20 cm (7–8 in).[258][259] The cicadas of the genus Tacua can also grow to comparably large sizes. The largest type of aphid is the giant oak aphid (Stomaphis quercus), which can reach an overall length of 2 cm (0.79 in).[260] The biggest species of leafhopper is Ledromorpha planirostris, which can reach a length of 2.8 cm (1.1 in).[261]
Megachile pluto, the largest bee.
Ants and allies (Hymenoptera)
The largest of the ants, and the heaviest species of the order, are the females of the African Dorylus helvolus, reaching a length of 5.1 cm (2.0 in) and a weight of 8.5 g (0.30 oz).[1] The ant that averages the largest for the mean size within the whole colony is a ponerine ant, Dinoponera gigantea, from South America, averaging up to 3.3 cm (1.3 in) from the mandibles to the end of abdomen.[1] Workers of the bulldog ant (Myrmecia brevinoda) of Australia are up to 3.7 cm (1.5 in) in total length, although much of this is from their extremely large mandibles.[1] The largest of the bee species, also in the order Hymenoptera, is Megachile pluto of Indonesia, the females of which can be 3.8 cm (1.5 in) long, with a 6.3 cm (2.5 in) wingspan. Nearly as large, the carpenter bees can range up to 2.53 cm (1.00 in).[1] The largest wasp is probably the so-called tarantula hawk species Pepsis pulszkyi of South America, at up to 6.8 cm (2.7 in) long and 11.6 cm (4.6 in) wingspan, although many other Pepsis approach a similar size. The giant scarab-hunting wasp Megascolia procer may rival the largest tarantula hawks in weight and wingspan, though its body is not as long.[1]
Moths and allies (Lepidoptera)
Queen Alexandra's birdwing.
The Hercules moth (Coscinocera hercules), in the family Saturniidae, is endemic to New Guinea and northern Australia, and its wings have the largest documented surface area (300 square centimeters) of any living insect,[262][263] and a maximum wingspan which is confirmed to 28 cm (11 in) while unconfirmed specimens have spanned up to 35.5 cm (14.0 in). The largest species overall is often claimed to be either the Queen Alexandra's birdwing (Ornithoptera alexandrae), a butterfly from Papua New Guinea, or the Atlas moth (Attacus atlas), a moth from Southeast Asia. Both of these species can reach a length of 8 cm (3.1 in), a wingspan of 28 cm (11 in) and a weight of 12 g (0.42 oz). One Atlas moth allegedly had a wingspan of 30 cm (12 in) but this measurement was not verified.[1] The larvae in the previous species can weigh up to 58 and 54 g (2.0 and 1.9 oz), respectively. However, there are no reported measurements of surface area that would exceed the Hercules moth, and the white witch (Thysania agrippina) of Central and South America, has the largest recorded wingspan of the order, and indeed of any living insect, though the white witch is exceeded in surface area by the Hercules moth. The verified record-sized Thysania spanned 30.8 cm (12.1 in) across the wings, although specimens have been reported to 36 cm (14 in).[1] The heaviest mature moths have been cited in the giant carpenter moth (Xyleutes boisduvali) of Australia, which has weighed up to 20 g (0.71 oz) although the species does not surpass 25.5 cm (10.0 in) in wingspan.[1]
Mantises (Mantodea)
The largest species of this order is Toxodera denticulata from Java, which has been measured up to 20 cm (7.9 in) in overall length.[264] However, an undescribed species from the Cameroon jungle is allegedly much larger than any other mantis and may rival the larger stick insects for the longest living insect.[265] Among widespread mantis species, the largest is the Chinese mantis (Tenodera aridifolia). The females of this species can attain a length of up to 10.6 cm (4.2 in).
Scorpionflies (Mecoptera)
The largest scorpionfly, the common scorpionfly (Panorpa communis), can reach a body length of about 30 millimetres (1.2 in).[266]
Alderflies and allies (Megaloptera)
This relatively small insect order includes some rather large species, many of which are noticeable for their elongated, imposing mandibles. The dobsonflies reach the greatest sizes of the order and can range up to 12.5 cm (4.9 in) in length.[267]
Net-winged insects (Neuroptera)
Blue eyes lacewing
These flying insects reach their largest size in Palparellus voeltzkowi, which can have a wingspan over 16 cm (6.3 in).[268] The largest lacewing is the "blue eyes lacewing" (Nymphes myrmeleonides) of Australia, which can measure up to 4 cm (1.6 in) in length and span 11 cm (4.3 in) across the wings.[269] Some forms of this ancient order could grow extremely large during the Jurassic period and may have ranked among the largest insects ever.[270] Found in the Early Cretaceous sedimentary rocks, Makarkinia adamsi had wings nearly 140–160 mm (5.5–6.3 in) in length.[271]
Dragonflies and damselflies (Odonata)
The largest species of Odonata is the damselfly Megaloprepus caerulatus of the neotropics, attaining a size of as much as 19 cm (7.5 in) across the wings and a body length of over 12 cm (4.7 in).[1] The largest species of dragonfly is Tetracanthagyna plagiata of Southeast Asia, spanning up to 17.6 cm (6.9 in) and measuring up to 11.8 cm (4.6 in) long; while bulkier and heavier than Megaloprepus (at up to 7 g (0.25 oz)), it is smaller in its linear dimensions.[1]
Grasshoppers and allies (Orthoptera)
Giant weta
The largest of this widespread, varied complex of insects are the giant wetas of New Zealand, which is now split among 12 species. The largest of these is the Little Barrier Island giant weta (Deinacrida heteracantha), the largest specimen was weighed at 71.3 g (2.52 oz), one of the largest insects weights ever known. These heavyweight insects can be over 9 cm (3.5 in) long.[1] The largest grasshopper species is often considered to be the Australian giant grasshopper (Valanga irregularis), which ranges up to 9 cm (3.5 in) in length.[272] The American eastern lubber grasshopper (Romalea guttata) can allegedly range up to 10 cm (3.9 in) in length.[273] However, the greatest grasshopper sizes known, to 12 cm (4.7 in), have been cited in the South American giant grasshopper (Tropidacris violaceus). The longest members of this order (although much lighter than the giant wetas) is the katydid Macrolyristes corporalis of Southeast Asia which can range up to 21.5 cm (8.5 in) with its long legs extended and can have a wingspan of 20 cm (7.9 in).[274]
Stick insects (Phasmatodea)
The longest known stick insects are also the longest known insects, notably species in the tribe Pharnaciini, but they are generally relatively lightweight because of their slender shape. The longest is an unnamed species of Phryganistria discovered in China in 2016, where a specimen held at the Insect Museum of West China in Chengdu has a total length of 62.4 cm (24.6 in).[275] The second-longest species is the Australian Ctenomorpha gargantua, females of which have been measured at over 60 cm (24 in) in total length.[276] Other very large species, formerly believed to be longest but now considered third longest is Sadyattes chani; a specimen held in the Natural History Museum in London has a total length of 56.7 cm (22.3 in).[277][278] These measurements are, however, with the front legs fully extended; it has a body length measuring 35.7 cm (14.1 in).[279] Another very large species is Phobaeticus kirbyi where the total length (including extended legs) is up to 54.6 cm (21.5 in) and the body alone up to 32.8 cm (12.9 in).[280] Another of the longest insect in terms of total length is Phobaeticus serratipes of Malaysia and Singapore, measuring up to 55.5 cm (21.9 in).[281] Another extremely long stick insect is Pharnacia maxima, which measured 51 cm (20 in) with its legs extended.[1] The spiny stick insect (Heteropteryx dilatata) of Malaysia does not reach the extreme lengths of its cousins, the body reaching up to 16 cm (6.3 in) long, but it is much bulkier. The largest Heteropteryx weighed about 65 g (2.3 oz) and was 3.5 cm (1.4 in) wide across the thickest part of the body.[1]
Lice (Phthiraptera)
These insects, which live parasitically on other animals, are as a rule quite small. The largest known species is the hog louse, Haematopinus suis, a sucking louse that lives on large livestock like pigs and cattle. It can range up to 6 mm (0.24 in) in length.[282]
Stoneflies (Plecoptera)
Pteronarcys californica
The largest species of stonefly is Pteronarcys californica of western North America, a species favored by fishermen as lures. This species can attain a length of 5 cm (2.0 in) and a wingspan of over 9.5 cm (3.7 in).[283]
Booklice (Psocoptera)
The largest of this order of very small insects are the barklice of the genus Psocus, the top size of which is about 1 cm.[284]
Fleas (Siphonaptera)
The largest species of flea is Hystrichopsylla schefferi. This parasite is known exclusively from the fur of the mountain beaver (Aplodontia rufa) and can reach a length of 1.2 cm (0.47 in).[1]
Silverfishes and allies (Thysanura)
These ancient flightless insects, some of which feed on human household objects, can range up to 4.3 cm (1.7 in) in length. A 350 million year old form was known to grow quite large, at up to 6 cm (2.4 in).[citation needed]
Thrips (Thysanoptera)
Members of the genus Phasmothrips are the largest kinds of thrips. The maximum size these species attain is approximately 1.3 cm (0.51 in) in length.[285]
Caddisflies (Trichoptera)
The largest of the small, moth-like caddisflies is Eubasilissa maclachlani. This species can range up to 7 cm (2.8 in) across the wings.[286]
Angel insects (Zoraptera)
The largest angel insect species, Hubbard's angel insect (Zorotypus hubbardi), grows up to 3 mm (0.12 in) in length.[287][288]

Cnidarians (Cnidaria)[edit]

The lion's mane jellyfish, one of the longest extant animals

The lion's mane jellyfish (Cyanea capillata) is the largest cnidarian species, of the class Scyphozoa. The largest known specimen of this giant, found washed up on the shore of Massachusetts Bay in 1870,[289][290] had a bell diameter of 2.5 m (8.2 ft), a weight of 150 kg (330 lb). The tentacles of this specimens were as long as 37 m (121 ft) and were projected to have a tentacular spread of about 75 m (246 ft) making it one of the longest extant animals.[1]

Corals and sea anemones (Anthozoa)
The largest individual species are the sea-anemones of the genus Discoma, which can attain a mouth disc diameter of 60 cm (2.0 ft).[291] Longer, but much less massive overall, are the anemones of the genus Ceriantharia, at up to 2 m (6.6 ft) tall.[292] Communities of coral can be truly massive, a single colony of the genus Porites can be over 10 m (33 ft), but the actual individual organisms are quite small.
Hydrozoans (Hydrozoa)
The colonial siphonophore Praya dubia can attain lengths of 40–50 m (130–160 ft).[293] The Portuguese man o' war's (Physalia physalis) tentacles can attain a length of up to 50 m (160 ft).[294] On 6 April 2020 the Schmidt Ocean Institute announced the discovery of a giant Apolemia siphonophore in submarine canyons near Ningaloo Coast, measuring 15 m (49 ft) diameter with a ring approximately 47 m (154 ft) long, claiming it was possibly the largest siphonophore ever recorded.[295][296]

Sponges (Porifera)[edit]

Despite its inert appearance, the heavyweight Xestospongia muta is indeed an animal

The largest known species of sea sponge is the giant barrel sponge, Xestospongia muta. These massively built sponges can reach 2.4 m (8 ft) in height and can be about the same thickness at the thickest part of the "body".[297] Some of these creatures have been estimated to be over 2,400 years of age.[298]

Calcareous sponges (Calcarea)
The largest known of these small, inconspicuous sponges is probably the species Pericharax heteroraphis, attaining a height of 30 cm (0.98 ft). Most calcareous sponges do not exceed 10 cm (3.9 in) tall.[citation needed]
Hexactinellid sponges (Hexactinellida)
A relatively common species, Rhabdocalyptus dawsoni, can reach a height of 1 m (3.3 ft) once they are of a very old age.[299] This is the maximum size recorded for a hexactinellid sponge.[citation needed]

See also[edit]

References[edit]

  1. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj Wood, Gerald The Guinness Book of Animal Facts and Feats (1983) ISBN 978-0-85112-235-9
  2. ^ Davies, Ella (20 April 2016). "The longest animal alive may be one you never thought of". BBC Earth. Retrieved 14 February 2018.
  3. ^ "Largest mammal". Guinness World Records.
  4. ^ Motani, Ryosuke; Pyenson, Nicholas D. (29 February 2024). "Downsizing a heavyweight: factors and methods that revise weight estimates of the giant fossil whale Perucetus colossus". PeerJ. 12: e16978. doi:10.7717/peerj.16978. ISSN 2167-8359. PMC 10909350. PMID 38436015.
  5. ^ "How Large Are Blue Whales Really? Size Comparison". Archived from the original on 11 November 2021. Retrieved 18 October 2019 – via www.youtube.com.
  6. ^ "How big are blue whales? And what does 'big' mean? By palaeozoologist on DeviantArt". February 2014.
  7. ^ McClain, Craig R.; Balk, Meghan A.; Benfield, Mark C.; Branch, Trevor A.; Chen, Catherine; Cosgrove, James; Dove, Alistair DM; Gaskins, Leo C.; Helm, Rebecca R. (13 January 2015). "Sizing ocean giants: patterns of intraspecific size variation in marine megafauna". PeerJ. 3: E715. doi:10.7717/peerj.715. ISSN 2167-8359. PMC 4304853. PMID 25649000.
  8. ^ a b Bianucci, Giovanni; Lambert, Olivier; Urbina, Mario; Merella, Marco; Collareta, Alberto; Bennion, Rebecca; Salas-Gismondi, Roberto; Benites-Palomino, Aldo; Post, Klaas; de Muizon, Christian; Bosio, Giulia; Di Celma, Claudio; Malinverno, Elisa; Paolo Pierantoni, Pietro; Maria Villa, Igor; Amson, Eli (2 August 2023). "A heavyweight early whale pushes the boundaries of vertebrate morphology". Nature. 620 (7975): 824–829. Bibcode:2023Natur.620..824B. doi:10.1038/s41586-023-06381-1. PMID 37532931. S2CID 260433513. Retrieved 2 August 2023.
  9. ^ Pester, Patrick (8 March 2024). "Colossus the enormous 'oddball' whale is not the biggest animal to ever live, scientists say". Lve Science. Retrieved 11 March 2024.
  10. ^ a b c d e Paul, Gregory S.; Larramendi, Asier (11 April 2023). "Body mass estimate of Bruhathkayosaurus and other fragmentary sauropod remains suggest the largest land animals were about as big as the greatest whales". Lethaia. 56 (2): 1–11. Bibcode:2023Letha..56..2.5P. doi:10.18261/let.56.2.5. ISSN 0024-1164. S2CID 259782734.
  11. ^ Emanuelson, Karen (2006). "Neonatal Care and Hand Rearing". Biology, Medicine, and Surgery of Elephants. pp. 223–241. doi:10.1002/9780470344484.ch16. ISBN 978-0-470-34448-4.
  12. ^ a b c Zimmer, Carl (29 February 2024). "Researchers Dispute Claim That Ancient Whale Was Heaviest Animal Ever - A new study argues that Perucetus, an ancient whale species, was certainly big, but not as big as today's blue whales". The New York Times. Archived from the original on 29 February 2024. Retrieved 2 March 2024.
  13. ^ a b c d e f g h i j k l m n o p q r Mark Tandy. Lives of Whales Archived 16 June 2017 at the Wayback Machine. Iwcoffice.org
  14. ^ Blue Whale. The Marine Mammal Center
  15. ^ North Pacific Right Whale | Marine education | Alaska Sea Grant. Seagrant.uaf.edu (15 February 2008)
  16. ^ a b c Stewart, et al, National Audubon Society Guide to Marine Mammals of the World. Knofp (2002), ISBN 978-0-375-41141-0
  17. ^ Bowhead Whales, Balaena mysticetus. Marinebio.org (30 September 2011)
  18. ^ Maine Department of Inland Fisheries and Wildlife – Maine Endangered Species Program/Northern Right Whale Archived 8 September 2011 at the Wayback Machine. Maine.gov
  19. ^ a b North Atlantic Right Whale. Animal Info (2 November 2005)
  20. ^ a b Whitehead, H. (2002). Sperm whale Physeter macrocephalus. In Perrin, W., Würsig B. and Thewissen, J.. Encyclopedia of Marine Mammals. Academic Press. pp. 1165–1172. ISBN 0-12-551340-2
  21. ^ WDC - Sperm Whale
  22. ^ Humpback Whale. Animal Info (1 February 2005)
  23. ^ David J. Schmidly; William B. Davis (2004). The mammals of Texas. University of Texas Press. ISBN 9780292702417.
  24. ^ "Sei Whale Species Guide". Whale and Dolphin Conservation (WDC). Retrieved 2 March 2016.
  25. ^ Bernd G. Würsig; J. G. M. Thewissen (2002). Encyclopedia of Marine Mammals. Gulf Professional Publishing. p. 70. ISBN 978-0-12-551340-1.
  26. ^ African Elephant Really Two Wildly Different Species. News.nationalgeographic.com (22 December 2010)
  27. ^ ADW: Loxodonta africana: Information. Animaldiversity.ummz.umich.edu
  28. ^ Georges Frei. Weight and Size of elephants in zoo and circus. Upali.ch
  29. ^ Larramendi, A. (2016). "Shoulder height, body mass and shape of proboscideans" (PDF). Acta Palaeontologica Polonica. 61. doi:10.4202/app.00136.2014. S2CID 2092950. Archived (PDF) from the original on 24 August 2016.
  30. ^ African Elephant. The Animal Files
  31. ^ a b Shoshani, J. and Eisenberg, J. F. Elephas maximus. Archived 30 April 2013 at the Wayback Machine Mammalian Species (1982) 182:1–8
  32. ^ a b Forest elephant videos, photos and facts – Loxodonta cyclotis Archived 24 October 2014 at the Wayback Machine. ARKive
  33. ^ Forest Elephant Loxodonta cyclotis – Appearance/Morphology: Measurement and Weight (Literature Reports). Wildpro.twycrosszoo.org Archived 11 May 2013 at the Wayback Machine
  34. ^ "White Rhino - Species - WWF". World Wildlife Fund. Retrieved 14 June 2017.
  35. ^ White rhinoceros videos, photos and facts – Ceratotherium simum Archived 9 May 2015 at the Wayback Machine. ARKive (6 August 2004)
  36. ^ "African Rhinoceros". viuzza.net. Archived from the original on 14 July 2015. Retrieved 14 June 2017.
  37. ^ White Rhinoceroses, White Rhinoceros Pictures, White Rhinoceros Facts. Animals.nationalgeographic.com
  38. ^ Indian rhinoceros videos, photos and facts – Rhinoceros unicornis Archived 4 October 2011 at the Wayback Machine. ARKive
  39. ^ Laurie, W. A.; Lang, E. M. and Groves, C. P. Rhinocerus unicorns Archived 29 June 2017 at the Wayback Machine. Mammalian Species (1983) 211:1–6
  40. ^ Boitani, Luigi, Simon & Schuster's Guide to Mammals. Simon & Schuster/Touchstone Books (1984), ISBN 978-0-671-42805-1
  41. ^ Indian rhinoceros Archived 26 September 2011 at the Wayback Machine. Ultimateungulate.com
  42. ^ Eltringham, S.K. (1999). The Hippos. Poyser Natural History Series. London: Academic Press. ISBN 0-85661-131-X.
  43. ^ Hippopotamus. Learnanimals.com
  44. ^ "Hippopotamus amphibius (hippopotamus)". Animal Diversity Web.
  45. ^ Hippopotamuses, Hippopotamus Pictures, Hippopotamus Facts. Animals.nationalgeographic.com
  46. ^ Black Rhinoceroses, Black Rhinoceros Pictures, Black Rhinoceros Facts. Animals.nationalgeographic.com
  47. ^ Hillman-Smith, A. K. Kes; Groves, Colin P. (1994). "Diceros bicornis". Mammalian Species (455): 1–8. doi:10.2307/3504292. JSTOR 3504292. S2CID 253955264.
  48. ^ ADW: Diceros bicornis: Information. (9 April 2009)
  49. ^ Javan Rhinoceros. Animal Info (26 November 2005)
  50. ^ Javan Rhino. Onehornedrhino.org Archived 13 October 2011 at the Wayback Machine
  51. ^ Javan rhinoceros videos, photos and facts – Rhinoceros sondaicus Archived 27 September 2011 at the Wayback Machine. ARKive
  52. ^ EDGE :: Mammal Species Information Archived 8 November 2017 at the Wayback Machine Edgeofexistence.org (12 November 2010)
  53. ^ Owen-Smith, R. Norman, Megaherbivores: The Influence of Very Large Body Size on Ecology. Cambridge University Press (1992), ISBN 978-0-521-42637-4
  54. ^ Giraffe. The Animal Files
  55. ^ a b Smith, A. T., Xie, Y. (eds.) (2008) A Guide to the Mammals of China. Princeton University Press, Princeton Oxfordshire. p. 472 ISBN 0691099847
  56. ^ Walrus: Physical Characteristics Archived 20 November 2018 at the Wayback Machine. Seaworld.org
  57. ^ "Largest mammal".
  58. ^ "Balaenoptera musculus (Blue whale)". Animal Diversity Web.
  59. ^ a b Larramendi, A. (2016). "Shoulder height, body mass and shape of proboscideans" (PDF). Acta Palaeontologica Polonica. 61. doi:10.4202/app.00136.2014. S2CID 2092950.
  60. ^ Fortelius, M.; Kappelman, J. (1993). "The largest land mammal ever imagined". Zoological Journal of the Linnean Society. 108: 85–101. doi:10.1111/j.1096-3642.1993.tb02560.x.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  61. ^ "An Ancient Elephant May Have Been Biggest Land Mammal Ever". 17 July 2015. Retrieved 21 April 2017.
  62. ^ a b St. Fleur, Nicholas (4 January 2019). "An Elephant-Size Relative of Mammals That Grazed Alongside Dinosaurs". The New York Times. Retrieved 7 January 2019.
  63. ^ Romano, Marco; Manucci, Fabio (14 June 2019). "Resizing Lisowicia bojani: volumetric body mass estimate and 3D reconstruction of the giant Late Triassic dicynodont". Historical Biology. 33 (4): 474–479. doi:10.1080/08912963.2019.1631819. ISSN 0891-2963. S2CID 196679837.
  64. ^ a b Anteosaurus. Palaeos.com
  65. ^ Romano, Marco; Citton, Paolo; Maganuco, Simone; Sacchi, Eva; Caratelli, Martina; Ronchi, Ausonio; Nicosia, Umberto (2019). Somerville, I. D. (ed.). "New basal synapsid discovery at the P outcrop of Torre del Porticciolo ( A lghero, I taly)". Geological Journal. 54 (3): 1554–1566. doi:10.1002/gj.3250. ISSN 0072-1050. S2CID 133755506.
  66. ^ "Permian Stratigraphy – International Commission on Stratigraphy International Union of Geological Sciences" (PDF). Archived (PDF) from the original on 3 December 2018. Retrieved 15 September 2022.
  67. ^ Reisz, Robert R.; Fröbisch, Jörg (16 April 2014). "The Oldest Caseid Synapsid from the Late Pennsylvanian of Kansas, and the Evolution of Herbivory in Terrestrial Vertebrates". PLOS ONE. 9 (4): e94518. Bibcode:2014PLoSO...994518R. doi:10.1371/journal.pone.0094518. PMC 3989228. PMID 24739998.
  68. ^ Berman, D.S.; Reisz, R.R.; Martens, T.; Henrici, A.C. (2001). "A new species of Dimetrodon (Synapsida: Sphenacodontidae) from the Lower Permian of Germany records first occurrence of genus outside of North America" (PDF). Canadian Journal of Earth Sciences. 38 (5): 803–812. Bibcode:2001CaJES..38..803B. doi:10.1139/cjes-38-5-803.
  69. ^ Brink, Kirstin S.; Reisz, Robert R. (16 October 2014). "Hidden dental diversity in the oldest terrestrial apex predator Dimetrodon". Nature Communications. 5: 3269. Bibcode:2014NatCo...5.3269B. doi:10.1038/ncomms4269. PMID 24509889.
  70. ^ Olson, E.C. (1955). "Parallelism in the evolution of the Permian reptilian faunas of the Old and New Worlds". Fieldiana. 37 (13): 395. Retrieved 15 September 2022.
  71. ^ Sulej, Tomasz; Niedzwiedzki, Grzegorz (4 January 2019). "An elephant-sized Late Triassic synapsid with erect limbs". Science. 363 (6422): 78–80. Bibcode:2019Sci...363...78S. doi:10.1126/science.aal4853. PMID 30467179.
  72. ^ "Gigantic mammal 'cousin' discovered". Science Daily. 23 November 2018. Retrieved 7 January 2019.
  73. ^ Crocodilian Biology Database – FAQ – Which is the largest species of crocodile? Flmnh.ufl.edu
  74. ^ Boloji.com – A Study in Diversity Archived 30 December 2010 at the Wayback Machine. News.boloji.com
  75. ^ ""Lolong" holds world record as largest croc in the world". Protected Areas and Wildlife Bureau. 17 November 2011. Archived from the original on 19 April 2012. Retrieved 23 June 2012.
  76. ^ Britton, Adam (12 November 2011). "Accurate length measurement for Lolong". Croc Blog. Retrieved 23 June 2012.
  77. ^ "NatGeo team confirms Lolong the croc is world's longest". GMA News Online. 9 November 2011. Retrieved 23 June 2012.
  78. ^ "Philippine town claims world's largest crocodile title". The Telegraph. 9 November 2011. Archived from the original on 10 November 2011. Retrieved 23 June 2012.
  79. ^ "'Lolong' claims world's largest croc title". ABS-CBNnews.com. Agence France-Presse. 9 November 2011. Archived from the original on 4 January 2015. Retrieved 23 June 2012.
  80. ^ Ciofi, Claudio (March 1999). "The Komodo Dragon". Scientific American. 280 (3): 84–91. Bibcode:1999SciAm.280c..84C. doi:10.1038/scientificamerican0399-84.
  81. ^ a b "Saltwater Crocodile". National Geographic. 11 November 2010. Archived from the original on 21 May 2011.
  82. ^ "Crocodylus porosus- Salt-water Crocodile, Estuarine Crocodile". Australian Government- Department of the Environment. Retrieved 3 June 2015.
  83. ^ Grigg, G. & Gans, C. "Morphology & Physiology of Crocodylia" (PDF). Australian Government- Department of the Environment. Retrieved 17 May 2016.
  84. ^ "World's Top 5 Largest Crocodiles Ever Recorded". Our Planet. 16 December 2017.
  85. ^ Webb, G. J.; Hollis, G. J. & Manolis, S. C. (1991). "Feeding, growth, and food conversion rates of wild juvenile saltwater crocodiles (Crocodylus porosus)". Journal of Herpetology. 25 (4): 462–473. doi:10.2307/1564770. JSTOR 1564770.
  86. ^ a b c d "BBC Nature - Nile crocodile videos, news and facts".
  87. ^ Orinoco crocodile videos, photos and facts – Crocodylus intermedius Archived 5 February 2019 at the Wayback Machine. ARKive
  88. ^ WAZA. "Orinoco Crocodile". Archived from the original on 10 November 2013. Retrieved 2 May 2013.
  89. ^ Leatherback Sea Turtle. euroturtle.org Archived 3 April 2012 at the Wayback Machine
  90. ^ "Client Validation". www.vanaqua.org. Archived from the original on 7 September 2011. Retrieved 14 June 2017.
  91. ^ Jake Fishman. "ADW: Crocodylus acutus: INFORMATION". Animal Diversity Web.
  92. ^ ANIMAL BYTES – American Crocodile Archived 2 November 2013 at the Wayback Machine. Seaworld.org
  93. ^ "American Crocodile". National Geographic. 10 September 2010. Archived from the original on 21 May 2011.
  94. ^ http://www.myfwc.com/media/664081/AmericanCrocodilesinFL.pdf Archived 7 May 2016 at the Wayback Machine [bare URL PDF]
  95. ^ Crocodilian Species – Black Caiman (Melanosucus niger). Crocodilian.com
  96. ^ "Black caiman videos, photos and facts - Melanosuchus niger - ARKive". Archived from the original on 4 February 2013. Retrieved 2 May 2013.
  97. ^ SeaWorld Parks & Entertainment. "Caimans". Archived from the original on 31 July 2013. Retrieved 3 May 2013.
  98. ^ http://www.iucncsg.org/365_docs/attachments/protarea/06_M-24b37cab.pdf [bare URL PDF]
  99. ^ a b "Gharial". Archived from the original on 18 October 2011. Retrieved 26 September 2011.
  100. ^ Gavials (Gharials), Gavial (Gharial) Pictures, Gavial (Gharial) Facts. Animals.nationalgeographic.com
  101. ^ a b "American Alligator". ScienceDaily. Archived from the original on 18 March 2015. Retrieved 28 February 2018.
  102. ^ a b c "American Alligator". 25 April 2016.
  103. ^ Lang, J. W.; Andrews, H.; Whitaker, R. (1989). "Sex determination and sex ratios in Crocodylus palustris". American Zoologist. 29 (3): 935–952. doi:10.1093/icb/29.3.935.
  104. ^ http://www.zoonegaramalaysia.my/RPFalseGharial.pdf Archived 27 January 2018 at the Wayback Machine [bare URL PDF]
  105. ^ Ahmad, A. A.; Dorrestein, G. M.; Oh, S. J. W. Y.; Hsu, C. D. (1 August 2017). "Multi-organ Metastasis of Fibrolamellar Hepatocellular Carcinoma in a Malayan Gharial (Tomistoma schlegelii)". Journal of Comparative Pathology. 157 (2): 80–84. doi:10.1016/j.jcpa.2017.06.007. ISSN 0021-9975. PMID 28942308.
  106. ^ "Tomistoma Task Force". tomistoma.org.
  107. ^ Chris Ng. "ADW: Dipsochelys dussumieri: INFORMATION". Animal Diversity Web.
  108. ^ a b Hughes, G. M.; Gaymer, R.; Moore, M.; Woakes, A. J. (1971). "Respiratory exchange and body size in the Aldabra giant tortoise". The Journal of Experimental Biology. 55 (3): 651–665. doi:10.1242/jeb.55.3.651. PMID 5160860.
  109. ^ "Information About Sea Turtles: Green Sea Turtle – Sea Turtle Conservancy". Retrieved 18 October 2019.
  110. ^ a b c "African Slender-Snouted Crocodile - The Maryland Zoo in Baltimore". The Maryland Zoo in Baltimore.
  111. ^ ADW: Geochelone nigra: Information. Animaldiversity.ummz.umich.edu
  112. ^ White Matt (18 August 2015). "2002: Largest Tortoise". Official Guinness World Records. Retrieved 27 November 2015.
  113. ^ San Diego Zoo's Animal Bytes: Galápagos Tortoise. Sandiegozoo.org
  114. ^ Mazzetta, G.V.; et al. (2004). "Giants and bizarres: Body size of some southern South American Cretaceous dinosaurs". Historical Biology. 16 (2–4): 1–13. Bibcode:2004HBio...16...71M. CiteSeerX 10.1.1.694.1650. doi:10.1080/08912960410001715132. S2CID 56028251.
  115. ^ Janensch, W. (1950). "The skeleton reconstruction of Brachiosaurus brancai". pp. 97–103.
  116. ^ Paul, G.S. (1988). "The brachiosaur giants of the Morrison and Tendaguru with a description of a new subgenus, Giraffatitan, and a comparison of the world's largest dinosaurs". Hunteria. 2 (3): 1–14.
  117. ^ a b c d Benson, R. B. J.; Campione, N. S. E.; Carrano, M. T.; Mannion, P. D.; Sullivan, C.; Upchurch, P.; Evans, D. C. (2014). "Rates of dinosaur body mass evolution indicate 170 million years of sustained ecological innovation on the avian stem lineage". PLOS Biology. 12 (5): e1001853. doi:10.1371/journal.pbio.1001853. PMC 4011683. PMID 24802911.
  118. ^ Taylor, M.P. (2009). "A re-evaluation of Brachiosaurus altithorax Riggs 1903 (Dinosauria, Sauropod) and its generic separation from Giraffatitan brancai (Janensch 1914)" (PDF). Journal of Vertebrate Paleontology. 29 (3): 787–806. Bibcode:2009JVPal..29..787T. doi:10.1671/039.029.0309. S2CID 15220647.
  119. ^ "The World of Dinosaurs". Museum für Naturkunde. Retrieved 18 November 2018.
  120. ^ Morgan, James (17 May 2014). "BBC News - 'Biggest dinosaur ever' discovered". BBC News. Retrieved 30 September 2014.
  121. ^ "Giant dinosaur slims down... a bit". BBC News. 10 August 2017. Retrieved 18 November 2018.
  122. ^ a b c Seebacher, F. (2001). "A new method to calculate allometric length-mass relationships of dinosaurs". Journal of Vertebrate Paleontology. 21 (1): 51–60. CiteSeerX 10.1.1.462.255. doi:10.1671/0272-4634(2001)021[0051:ANMTCA]2.0.CO;2. ISSN 0272-4634. S2CID 53446536.
  123. ^ a b Henderson, Donald (2013). "Sauropod Necks: Are They Really for Heat Loss?". PLOS ONE. 8 (10): e77108. Bibcode:2013PLoSO...877108H. doi:10.1371/journal.pone.0077108. PMC 3812985. PMID 24204747.
  124. ^ a b "Volumetric analysis of Barosaurus' size". thesauropodomorphlair. 28 January 2020.
  125. ^ Mortimer, M. (2001), "Re: Bruhathkayosaurus" Archived 22 May 2017 at the Wayback Machine, discussion group, The Dinosaur Mailing List, 19 June 2001. Accessed 23 May 2008.
  126. ^ Paul, Gregory S. (2019). "Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" (PDF). Annals of the Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. S2CID 210840060.
  127. ^ Mortimer, M. (2001), "Re: Bruhathkayosaurus", discussion group, The Dinosaur Mailing List, 19 June 2001. Accessed 23 May 2008.
  128. ^ Mortimer, M. (2004), "Re: Largest Dinosaurs" Archived 13 September 2019 at the Wayback Machine, discussion group, The Dinosaur Mailing List, 7 September 2004. Accessed 23 May 2008.
  129. ^ a b Paul, Gregory S. (2019). "Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" (PDF). Annals of the Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. S2CID 210840060.
  130. ^ Carpenter, K. (2006). "Biggest of the big: a critical re-evaluation of the mega-sauropod Amphicoelias fragillimus." In Foster, J.R. and Lucas, S.G., eds., 2006, Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36: 131–138.
  131. ^ Paul, G.S., 2016, The Princeton Field Guide to Dinosaurs 2nd edition, Princeton University Press p. 213
  132. ^ a b "The size of the BYU 9024 animal". svpow.com. 16 June 2019. Archived from the original on 16 April 2022.
  133. ^ a b c d e González Riga, Bernardo J.; Lamanna, Matthew C.; Ortiz David, Leonardo D.; Calvo, Jorge O.; Coria, Juan P. (2016). "A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot". Scientific Reports. 6: 19165. Bibcode:2016NatSR...619165G. doi:10.1038/srep19165. PMC 4725985. PMID 26777391.
  134. ^ Paul, G.S. (1997). "Dinosaur models: the good, the bad, and using them to estimate the mass of dinosaurs" (PDF). In Wolberg, D.L.; Stump, E.; Rosenberg, G.D. (eds.). DinoFest International Proceedings. Dinofest International. The Academy of Natural Sciences. pp. 129–154. Archived (PDF) from the original on 4 March 2016.
  135. ^ a b c Paul, Gregory S. (2019). "Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" (PDF). Annals of the Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. S2CID 210840060.
  136. ^ a b c Carballido, José L.; Pol, Diego; Otero, Alejandro; Cerda, Ignacio A.; Salgado, Leonardo; Garrido, Alberto C.; Ramezani, Jahandar; Cúneo, Néstor R.; Krause, Javier M. (1860). "A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs". Proceedings of the Royal Society B: Biological Sciences. 284 (1860): 20171219. doi:10.1098/rspb.2017.1219. PMC 5563814. PMID 28794222.
  137. ^ a b Paul, G.S. (2016) The Princeton Field Guide to Dinosaurs. 2nd ed. Princeton University Press p. 206
  138. ^ Holtz, Tom (2012) Genus List for Holtz (2007) Dinosaurs
  139. ^ Holtz, Thomas R. (2014). "Supplementary Information to Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages".
  140. ^ Hartman, Scott (2013). "The biggest of the big". Skeletal Drawing. Retrieved 4 November 2018.
  141. ^ a b c Wedel, Mathew J.; Cifelli, R.L.; Sanders, R..K. (2000). "Osteology, paleobiology, and relationships of the sauropod dinosaur Sauroposeidon" (PDF). Acta Palaeontologica Polonica. 45: 343–388. Archived (PDF) from the original on 5 July 2008. Retrieved 27 July 2009.
  142. ^ Wedel, Mathew J.; Cifelli, Richard L. (Summer 2005). "Sauroposeidon: Oklahoma's Native Giant" (PDF). Oklahoma Geology Notes. 65 (2): 40–57. Archived from the original (PDF) on 20 September 2008.
  143. ^ a b c Bates, Karl T.; Falkingham, Peter L.; Macaulay, Sophie; Brassey, Charlotte; Maidment, Susannah C.R. (2015). "Downsizing a giant: re-evaluating Dreadnoughtus body mass". Biol Lett. 11 (6): 20150215. doi:10.1098/rsbl.2015.0215. PMC 4528471. PMID 26063751.
  144. ^ Hutchinson, John R.; Bates, Karl T.; Molnar, Julia; Allen, Vivian; Makovicky, Peter J. (2011). "A computational analysis of limb and body dimensions in Tyrannosaurus rex with implications for locomotion, ontogeny, and growth". PLOS ONE. 6 (10): 1–20. Bibcode:2011PLoSO...626037H. doi:10.1371/journal.pone.0026037. PMC 3192160. PMID 22022500.
  145. ^ Hartman, Scott (7 July 2013). "Mass estimates: North vs South redux". Retrieved 17 August 2015.
  146. ^ Dal Sasso, Christiano; Maganuco, Simone; Buffetaut, Eric; Mendez, Marco A. (2005). "New information on the skull of the enigmatic theropod Spinosaurus, with remarks on its size and affinities" (PDF). Journal of Vertebrate Paleontology. 25 (4): 888–896. doi:10.1671/0272-4634(2005)025[0888:niotso]2.0.co;2. S2CID 85702490. Archived from the original (PDF) on 29 April 2011. Retrieved 17 August 2015.
  147. ^ a b Therrien, F.; Henderson, D.M. (2007). "My theropod is bigger than yours...or not: estimating body size from skull length in theropods". Journal of Vertebrate Paleontology. 27 (1): 108–115. doi:10.1671/0272-4634(2007)27[108:MTIBTY]2.0.CO;2. ISSN 0272-4634. S2CID 86025320.
  148. ^ Ibrahim, Nizar; Sereno, Paul C.; Dal Sasso, Christiano; Maganuco, Simone; et al. (2014). "Semiaquatic adaptations in a giant predatory dinosaur". Science. 345 (6204): 1613–1616. Bibcode:2014Sci...345.1613I. doi:10.1126/science.1258750. PMID 25213375. S2CID 34421257.
  149. ^ "Discoveries - Paul Sereno - Paleontologist - The University of Chicago". paulsereno.uchicago.edu. Retrieved 14 June 2017.
  150. ^ Coria, R. A.; Salgado, L. (1995). "A new giant carnivorous dinosaur from the Cretaceous of Patagonia". Nature. 377 (6546): 225–226. Bibcode:1995Natur.377..224C. doi:10.1038/377224a0. S2CID 30701725.
  151. ^ Coria, R. A. and Currie, P. J. (2006). "A new carcharodontosaurid (Dinosauria, Theropoda) from the Upper Cretaceous of Argentina". Geodiversitas, 28 (1): 71–118. pdf link Archived 2 October 2013 at the Wayback Machine
  152. ^ Holtz, Thomas R. Jr. (2012) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2011 Appendix.
  153. ^ Sereno, P. C.; Dutheil, D. B.; Iarochene, M.; Larsson, H. C. E.; Lyon, G. H.; Magwene, P. M.; Sidor, C. A.; Varricchio, D. J.; Wilson, J. A. (1996). "Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation" (PDF). Science. 272 (5264): 986–991. Bibcode:1996Sci...272..986S. doi:10.1126/science.272.5264.986. PMID 8662584. S2CID 39658297.
  154. ^ Chemistry - Stegosaurus. Chemistrydaily.com Archived 26 January 2016 at the Wayback Machine
  155. ^ Ankylosaurus Facts. sciencekids.co.nz
  156. ^ Zhao, M. Z.; Zhang, H. Z. (2006). "The weighted transience and recurrence of Markov processes". Acta Mathematica Sinica, English Series. 23: 111–126. doi:10.1007/s10114-005-0808-x. S2CID 119700784.
  157. ^ a b Morris, William J. (1981). "A new species of hadrosaurian dinosaur from the Upper Cretaceous of Baja California: Lambeosaurus laticaudus". Journal of Paleontology. 55 (2): 453–462. JSTOR 1304231.
  158. ^ Dodson, P. (1996). The Horned Dinosaurs. Princeton University Press, Princeton, New Jersey. ISBN 978-0-691-02882-8.
  159. ^ Alexander, R. M. (1985). "Mechanics of posture and gait of some large dinosaurs". Zoological Journal of the Linnean Society. 83: 1–25. doi:10.1111/j.1096-3642.1985.tb00871.x.
  160. ^ Wu, X. C.; Brinkman, D. B.; Eberth, D. A.; Braman, D. R. (2007). "A new ceratopsid dinosaur (Ornithischia) from the uppermost Horseshoe Canyon Formation (upper Maastrichtian), Alberta, Canada". Canadian Journal of Earth Sciences. 44 (9): 1243–1265. Bibcode:2007CaJES..44.1243W. doi:10.1139/E07-011.
  161. ^ a b birding.com records Archived 2 March 2007 at the Wayback Machine
  162. ^ Avian Medicine: Principles and Application. avianmedicine.net Archived 19 April 2009 at the Wayback Machine
  163. ^ Hansford, J. P.; Turvey, S. T. (26 September 2018). "Unexpected diversity within the extinct elephant birds (Aves: Aepyornithidae) and a new identity for the world's largest bird". Royal Society Open Science. 5 (9): 181295. Bibcode:2018RSOS....581295H. doi:10.1098/rsos.181295. PMC 6170582. PMID 30839722.
  164. ^ Murray, Peter F.; Vickers-Rich, Patricia (2004). Magnificent Mihirungs: The Colossal Flightless Birds of the Australian Dreamtime. Indiana University Press. ISBN 978-0-253-34282-9
  165. ^ Alvarenga, H.; Chiappe, L.; Bertelle, S. (3 May 2011), Phorusrhacids: the Terror Birds, in Dyke, G.; Kaiser, G., Living Dinosaurs: The Evolutionary History of Modern Birds, Wiley, pp. 187–208, ISBN 978-0-470-65666-2
  166. ^ Chatterjee, S.; Templin, R. J.; Campbell, K. E. (2007). "The aerodynamics of Argentavis, the world's largest flying bird from the Miocene of Argentina". Proceedings of the National Academy of Sciences. 104 (30): 12398–12403. Bibcode:2007PNAS..10412398C. doi:10.1073/pnas.0702040104. PMC 1906724. PMID 17609382.
  167. ^ a b Ksepka, Daniel T. (22 July 2014). "Flight performance of the largest volant bird". Proceedings of the National Academy of Sciences. 111 (29): 10624–10629. Bibcode:2014PNAS..11110624K. doi:10.1073/pnas.1320297111. PMC 4115518. PMID 25002475.
  168. ^ Osborne, Hannah (7 July 2014). "Pelagornis Sandersi: World's Biggest Bird Was Twice as Big as Albatross with 24ft Wingspan". ibtimes.co.uk. Retrieved 14 June 2017.
  169. ^ a b c d e f g h i j k l m n o Davies, Stephen, Ratites and Tinamous. Oxford University Press (2002), ISBN 978-0-19-854996-3
  170. ^ a b c d e f g h i j del Hoyo, et al.,Handbook of the Birds of the World. Volume 1: Ostrich to Ducks. Lynx Edicons (1992), ISBN 978-84-87334-10-8
  171. ^ Kofron, Christopher P. (1999). "Attacks to humans and domestic animals by the southern cassowary (Casuarius casuarius johnsonii) in Queensland, Australia". Journal of Zoology. 249 (4): 375–381. doi:10.1111/j.1469-7998.1999.tb01206.x.
  172. ^ a b CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), ISBN 978-0-8493-4258-5.
  173. ^ a b Marion, Remi, Penguins: A Worldwide Guide. Sterling Publishing Co. (1999), ISBN 0-8069-4232-0
  174. ^ Ramsey, Tanya Lewis, Lydia. "The turkeys we eat today weigh twice as much as they did a few decades ago". Business Insider.{{cite web}}: CS1 maint: multiple names: authors list (link)
  175. ^ "Turkey Facts - Turkey for Holidays - University of Illinois Extension". extension.illinois.edu. Archived from the original on 8 July 2018. Retrieved 2 September 2022.
  176. ^ a b c d del Hoyo, J; Elliot, A; Sargatal, J (1996). Handbook of the Birds of the World 3. Barcelona: Lynx Edicions. ISBN 84-87334-20-2.
  177. ^ Leopard Seals Group Penguin Slideshow Ppt Presentation Archived 18 February 2018 at the Wayback Machine. Authorstream.com (31 March 2009)
  178. ^ Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. ISBN 978-0-85112-235-9.
  179. ^ Cox, C. B.; Hutchinson, P. (1991). "Fishes and amphibians from the Late Permian Pedrado Fogo Formation of northern Brazil" (PDF). Palaeontology. 34: 561–573. INIST 19854877. Archived from the original (PDF) on 24 March 2012.
  180. ^ African Bullfrog. Honoluluzoo.org Archived 3 September 2011 at the Wayback Machine
  181. ^ Giant "Frog From Hell" Fossil Found in Madagascar. News.nationalgeographic.com (28 October 2010)
  182. ^ Evans, Susan E.; Groenke, Joseph R.; Jones, Marc E. H.; Turner, Alan H.; Krause, David W. (28 January 2014). "New Material of Beelzebufo, a Hyperossified Frog (Amphibia: Anura) from the Late Cretaceous of Madagascar". PLOS ONE. 9 (1): e87236. Bibcode:2014PLoSO...987236E. doi:10.1371/journal.pone.0087236. PMC 3905036. PMID 24489877.
  183. ^ White Lipped Tree Frog. The Animal Files
  184. ^ Surinam horned frog (Ceratophrys cornuta) – Videos Peru – Peru Videos. Bullafina.com (11 June 2008) [dead link]
  185. ^ Golden Poison Dart Frogs, Golden Poison Dart Frog Pictures, Golden Poison Dart Frog Facts. Animals.nationalgeographic.com
  186. ^ Zweifel, Richard G. (1998). Cogger, H. G. and Zweifel, R. G.. ed. Encyclopedia of Reptiles and Amphibians. San Diego: Academic Press. ISBN 0-12-178560-2.
  187. ^ Platt, John R. "Swampy Thing: The Giant New Salamander Species Discovered in Florida and Alabama". Scientific American Blog Network. Retrieved 19 December 2018.
  188. ^ The Largest Newt As a Pet. Buzzle.com
  189. ^ a b André, Frédéric; Tourenne, Murielle; Foveau, Aurélie (8 August 2011). "Synoicum pulmonaria (Ellis & Solander, 1786)" (in French). DORIS. Retrieved 26 July 2015.
  190. ^ Lambert, Gretchen; Lambert, Charles C. (1996). "Spicule Formation in the New Zealand Ascidian Pyura pachydermatina (Chordata, Ascidiacea)". Connective Tissue Research. 34 (4): 263–269. doi:10.3109/03008209609005270. PMID 9084635.
  191. ^ "Pyura". Biosecurity in New Zealand. Ministry for Primary Industries, New Zealand Government. Archived from the original on 9 February 2013. Retrieved 19 April 2013.
  192. ^ Ruppert, Edward E.; Fox, Richard, S.; Barnes, Robert D. (2004). Invertebrate Zoology (7th ed.). Cengage Learning. p. 941. ISBN 978-81-315-0104-7.{{cite book}}: CS1 maint: multiple names: authors list (link)
  193. ^ a b Cyclosalpa bakeri - Ritter, 1905 Archived 21 March 2016 at the Wayback Machine JelliesZone, by David Wrobel. Retrieved 10 April 2016.
  194. ^ Pyrosoma atlanticum Marine Species Identification Portal. Retrieved 11 November 2011.
  195. ^ Doliolida World Register of Marine Species. Retrieved 17 November 2011.
  196. ^ Pelagic Tunicates Archived 23 June 2016 at the Wayback Machine JelliesZone, by David Wrobel. Retrieved 10 April 2016.
  197. ^ Branchiostoma lanceolatum Marine Species Information Portal. Retrieved 14 November 2011.
  198. ^ a b Mah, Christopher (27 July 2008). "What Are the World's LARGEST Starfish?". The Echinoblog.
  199. ^ Ponsonby, David and Dussart, George The Anatomy of the Sea, Raincoast Books (2005) ISBN 0-8118-4633-4 p. 129
  200. ^ a b c d Gilpin, D. (2006). Echinoderms - Starfish, Urchins, and Other Echinoderms. Capstone. p. 41. ISBN 978-0-7565-1611-6.
  201. ^ "Strongylocentrotus franciscanus". Animal Diversity Web. Retrieved 2 December 2016.
  202. ^ "Long-spined black sea urchin". Wild Singapore. Retrieved 2 December 2016.
  203. ^ Mah, Christopher (27 April 2009). "The World's BIGGEST Brittle Stars!". The Echinoblog.
  204. ^ "Neoentobdella gen. nov for species of Entobdella Blainville in Lamarck, 1818 (Monogenea, Capsalidae, Entobdellinae) from stingray hosts, with descriptions of two new species" (PDF). Acta Parasitologica. 50 (1): 32–48. 2005.
  205. ^ DPDx – Fasciolopsiasis. Dpd.cdc.gov Archived 27 September 2011 at the Wayback Machine
  206. ^ The Persistent Parasites. Time Magazine (8 April 1957)
  207. ^ Hargis, William J. Parasitology and pathology of marine organisms of the world ocean Archived 15 October 2011 at the Wayback Machine. National Oceanic and Atmospheric Administration (1985)
  208. ^ The Mighty Worm. Worm Digest (2 October 2005) Archived 19 February 2009 at the Wayback Machine
  209. ^ Carwardine, M. 1995. The Guinness Book of Animal Records. Guinness Publishing. p. 232.
  210. ^ Photo in the News: Colossal Squid Caught off Antarctica. News.nationalgeographic.com (28 October 2010)
  211. ^ "The UnMuseum - The Colossal Squid". www.unmuseum.org. Retrieved 14 June 2017.
  212. ^ Kathy Marks. NZ's colossal squid to be microwaved Archived 29 September 2007 at the Wayback Machine. The New Zealand Herald (23 March 2007)
  213. ^ "How big is the colossal squid?". Te papa. Archived from the original on 17 December 2008.
  214. ^ Giant Squids, Architeuthis dux. Marinebio.org
  215. ^ Giant Squid, Giant Squid Pictures, Giant Squid Facts. Animals.nationalgeographic.com (4 December 2006)
  216. ^ O'Shea, S. 2003. "Giant Squid and Colossal Squid Fact Sheet". The Octopus News Magazine Online.
  217. ^ Salvini-Plawen, L. V. (1997). "Systematic Revision of the Epimeniidae (Mollusca: Solenogastres)". Journal of Molluscan Studies. 63 (2): 131–155. doi:10.1093/mollus/63.2.131.
  218. ^ Gumboot Chiton. alaska.gov
  219. ^ Kauffman, E. G.; Harries, P. J.; Meyer, C.; Villamil, T.; Arango, C.; Jaecks, G. (2007). "Paleoecology of Giant Inoceramidae (Platyceramus) on a Santonian (Cretaceous) Seafloor in Colorado". Journal of Paleontology. 81: 64–81. doi:10.1666/0022-3360(2007)81[64:POGIPO]2.0.CO;2. S2CID 130048023.
  220. ^ John D. Taylor and Emily A. Glover. Food of giants – field observations on the diet of Syrinx aruanus (Linnaeus, 1758) (Turbinellidae) the largest living gastropod, in F. E. Wells, D. I. Walker and D. S. Jones (eds.) 2003. The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth.
  221. ^ Largest snails in the world – Giant African snail. largestfastestsmartest.co.uk
  222. ^ Cosgrove, James Albert (1987). Aspects of the natural history of Octopus dofleini, the giant Pacific octopus (Thesis). hdl:1828/12155.
  223. ^ [Octopus – Species].[full citation needed]
  224. ^ Gubanov, NM (21 April 1951). "Гигантская нематода из плаценты китообразных Placentonema gigantissima nov. gen., nov. sp" [Giant nematoda from the placenta of Cetacea; Placentonema gigantissima nov. gen., nov. sp] (PDF). Doklady Akademii Nauk SSSR (in Russian). 77 (6): 1123–1125. PMID 14822893. Archived from the original (PDF) on 15 December 2013.
  225. ^ Natural History Collections: Nematoda. Nhc.ed.ac.uk
  226. ^ Morera-Brenes, B.; Monge-Nájera, J. (2010). "A new giant species of placented worm and the mechanism by which onychophorans weave their nets (Onychophora: Peripatidae)". Revista de Biología Tropical. 58 (4): 1127–1142. arXiv:1511.00983. doi:10.15517/rbt.v58i4.5398. PMID 21246983. S2CID 6456946.
  227. ^ Tetlie, O. E.; Poschmann, M.; Braddy, S. J. (2008). "Giant claw reveals the largest ever arthropod". Biology Letters. 4 (1): 106–109. doi:10.1098/rsbl.2007.0491. PMC 2412931. PMID 18029297.
  228. ^ "First Contact". panda.org. Archived from the original on 24 February 2017. Retrieved 14 June 2017.
  229. ^ Poisonous Animals: Tarantula (Grammostola, Phrixothrichus). Library.thinkquest.org
  230. ^ Jeram, Andrew J. (1993). "Scorpions from the Viséan of East Kirkton, West Lothian, Scotland, with a revision of the infraorder Mesoscorpionina". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 84 (3–4): 283–299. Bibcode:1993EESTR..84..283J. doi:10.1017/S0263593300006106. ISSN 1755-6929. S2CID 131416804.
  231. ^ Erik N. Kjellesvig-Waering (1972). "Brontoscorpio anglicus: A Gigantic Lower Paleozoic Scorpion from Central England". Journal of Paleontology. 46 (1): 39–42. JSTOR 1302906.
  232. ^ Jan Beccaloni (2009). "Pseudoscorpiones". Arachnids. Natural History Museum. p. 271. ISBN 978-0-520-26140-2.
  233. ^ P. K. L. Ng; D. Guinot; P. J. F. Davie (2008). "Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world" (PDF). Raffles Bulletin of Zoology. 17: 1–286. Archived from the original (PDF) on 6 June 2011.
  234. ^ D. R. Currie; T. M. Ward (2009). South Australian Giant Crab (Pseudocarcinus gigas) Fishery (PDF). South Australian Research and Development Institute. Fishery Assessment Report for PIRSA. Retrieved 9 December 2013.
  235. ^ Roy Caldwell. "Species: Lysiosquillina maculata". Roy's List of Stomatopods for the Aquarium. University of California Museum of Paleontology. Retrieved 14 February 2009.
  236. ^ "Tasmanian Giant Freshwater Lobster (Astacopsis gouldi)". Department of the Environment, Water, Heritage and the Arts. 9 February 2007. Retrieved 16 March 2010.
  237. ^ T. Walsh & N. Doran (2010). "Astacopsis gouldi". IUCN Red List of Threatened Species. 2010: e.T2190A9337732. doi:10.2305/IUCN.UK.2010-3.RLTS.T2190A9337732.en.
  238. ^ Branchinecta gigas (crustacean). Britannica Online Encyclopaedia
  239. ^ Pennella balaenopterae. Animaldiversity.ummz.umich.edu
  240. ^ Giant Acorn Barnacle. Oregon Coast Aquarium Archived 9 October 2011 at the Wayback Machine
  241. ^ Knight, J.D. Giant Isopod – Deep Sea Creatures on Sea and Sky. Seasky.org
  242. ^ Remipedia: Species – robustus, Godzillius. Crustacea.net (2 October 2002)
  243. ^ Horseshoe Crabs, Limulus polyphemus at. Marinebio.org
  244. ^ Sea spiders Facts, information, pictures. work=Encyclopedia.com (22 October 2004)
  245. ^ Gutierrez-Marco, J. C.; Sa, A. A.; Garcia-Bellido, D. C.; Rabano, I.; Valerio, M. (2009). "Giant trilobites and trilobite clusters from the Ordovician of Portugal". Geology. 37 (5): 443–446. Bibcode:2009Geo....37..443G. doi:10.1130/G25513A.1.
  246. ^ Giant Trilobites in Portugal Could Be Biggest Portugal – Discovery News. Dsc.discovery.com (7 May 2009). Archived 10 May 2009 at the Wayback Machine
  247. ^ Scolopendra gigantea. Arachnoboards.com (13 August 2003)
  248. ^ Minelli, Alessandro; Sergei I. Golovatch (2001). "Myriapods" (PDF). In Simon A. Levin (ed.). Encyclopedia of Biodiversity. Academic Press. pp. 291–303. ISBN 978-0122268656. Archived from the original (PDF) on 21 February 2014.
  249. ^ "Largest". Archived from the original on 18 July 2011. Retrieved 10 June 2009.{{cite web}}: CS1 maint: bot: original URL status unknown (link)(2011). source: The University of Florida Book of Insect Records "Largest". Archived from the original on 20 August 2014. Retrieved 10 June 2009.
  250. ^ Creature Features – Giant Burrowing Cockroach Archived 18 February 2014 at the Wayback Machine. Abc.net.au
  251. ^ The Giant Earwig of St. Helena Labidura herculeana. Earwig Research Centre. Earwigs-online.de
  252. ^ Trust), David Pryce (St Helena National; Liza White (Environment and Natural Resources Directorate, St Helena Government) (22 August 2014). "IUCN Red List of Threatened Species: Labidura herculeana". IUCN Red List of Threatened Species.
  253. ^ "Gone for good: world's largest earwig declared extinct". Mongabay Environmental News. 19 November 2014. Retrieved 12 October 2021.
  254. ^ Madagascan mayfly hyper-diversity. The BioFresh blog (24 May 2011)
  255. ^ P. J. Perez-Goodwyn (2006). Taxonomic revision of the subfamily Lethocerinae Lauck & Menke (Heteroptera: Belostomatidae)". Stuttgarter Beiträge zur Naturkunde. A (Biologie) 695: 1–71.
  256. ^ Haddad Jr; Schwartz; Schwartz; and Carvalho (2010). Bites Caused by Giant Water Bugs Belonging to Belostomatidae Family (Hemiptera, Heteroptera) in Humans: A Report of Seven Cases. Wilderness & Environmental Medicine 21: 130–133.
  257. ^ BBC News (26 May 2011). Giant water bug photographed devouring baby turtle. Retrieved 27 August 2014.
  258. ^ Burton, Maurice; Burton, Robert (2002). International Wildlife Encyclopedia. 4, Chickaree - crabs (3rd ed.). Tarrytown, NY: Marshall Cavendish Corporation. p. 455. ISBN 0-7614-7270-3
  259. ^ Flindt, R. (2006). Amazing Numbers in Biology, p. 10. ISBN 978-3540301462
  260. ^ Giant Oak Aphid hunt is on. The Telegraph (8 August 2007)
  261. ^ Ledromorpha planirostris Archived 14 May 2012 at the Wayback Machine. Bugs.bio.usyd.edu.au
  262. ^ Robert G. Foottit & Peter H. Adler. 2009. Insect Biodiversity: Science and Society. Blackwell Publishing Ltd. ISBN 978-1-405-15142-9
  263. ^ Rainier Flindt. 2006. Amazing Numbers in Biology. Springer-Verlag, Berlin. ISBN 3-540-30146-1
  264. ^ Live Pet Mantis Hobby Archived 4 December 2020 at the Wayback Machine. Bugsincyberspace.com
  265. ^ Craig Glenday (2009). Guinness World Records 2009. Random House Digital, Inc. p. 53. ISBN 978-0-553-59256-6.
  266. ^ "Scorpion Flies - Panorpa communis - UK Safari". www.uksafari.com.
  267. ^ Dobsonfly. Real Monstrosities (26 January 2011)
  268. ^ Palparellus voeltzkowi (Kolbe, 1906). Researcharchive.calacademy.org
  269. ^ Bio-Ditrl, Department of Biological Sciences, University of Alberta Archived 6 July 2011 at the Wayback Machine
  270. ^ Michael S. Engel (2005). "A remarkable kalligrammatid lacewing from the Upper Jurassic of Kazakhstan (Neuroptera: Kalligrammatidae)". Transactions of the Kansas Academy of Science. 108 (1): 59–62. doi:10.1660/0022-8443(2005)108[0059:arklft]2.0.co;2. JSTOR 3628206. S2CID 86240200.
  271. ^ Bechly, G.; Makarkin, V. N. (2016). "A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the Americas". Cretaceous Research. 58: 135–140. Bibcode:2016CrRes..58..135B. doi:10.1016/j.cretres.2015.10.014.
  272. ^ Giant Grasshoppers – The largest grasshopper – Valanga irregularis. Brisbaneinsects.com
  273. ^ Eastern Lubber Grasshopper – Florida eco travel guide. Wildflorida.com
  274. ^ Giant Long-Legged Katydid Archived 7 March 2016 at the Wayback Machine. Hmns.org
  275. ^ "Longest Insect discovered in China". Archived from the original on 8 May 2016.
  276. ^ Brock, Paul D.; Hasenpusch, Jack W. (2009). The Complete Field Guide to Stick and Leaf Insects of Australia. CSIRO Publishing. p. 106. ISBN 9780643094185.
  277. ^ "World's longest insect revealed". Natural History Museum. 16 October 2008. Archived from the original on 19 October 2008. Retrieved 16 October 2008.
  278. ^ Seow-Choen, F. (1995). The longest insect in the world. Malayan Nat. 48: 12.
  279. ^ Hennemann, F.H. & Conle, O.V. (October 2008). "Revision of Oriental Phasmatodea: The tribe Pharnaciini Günther, 1953, including the description of the world's longest insect, and a survey of the family Phasmatidae Gray, 1835 with keys to the subfamilies and tribes (Phasmatodea: "Anareolatae": Phasmatidae)" (PDF). Zootaxa. 1906. Auckland, New Zealand: Magnolia Press: 1–316. doi:10.11646/zootaxa.1906.1.1. ISSN 1175-5326. Retrieved 16 October 2008.
  280. ^ Brock, P.D. 1999. The amazing world of stick and leaf-insects. Cravitz Printing Co., Essex, England.
  281. ^ Seow-Choen, F. (1995). "The longest insect in the world". Malayan Nat. 48: 12.
  282. ^ ADW: Haematopinus suis: Information. Animaldiversity.ummz.umich.edu
  283. ^ Pteronarcys californica – aka Giant Stonefly or Giant Salmonfly. Riverwood Blog – Fly Fishing Gear & Guided Fishing Trips in Oregon (20 April 2009) Archived 2 April 2012 at the Wayback Machine
  284. ^ National Barkfly (Outdoor Psocoptera) Recording Scheme. Brc.ac.uk
  285. ^ List of largest insects. Paulsquiz.com Archived 14 July 2015 at the Wayback Machine
  286. ^ Diptera.info – Discussion Forum: The LARGEST caddisfly of the world.
  287. ^ Foottit, Robert G.; Adler, Peter H. (2018). Insect Biodiversity: Science and Society. Wiley. pp. 199–207. ISBN 978-1-118-94560-5.
  288. ^ Engel, Michael S. (2005). "Zoraptera". Tree of Life Web Project. Retrieved 9 March 2019.
  289. ^ Rare sighting of a lion’s mane jellyfish in Tramore Bay. waterford-today.ie
  290. ^ "Lion's Mane Jellyfish". jellyfishfacts.net. Archived from the original on 26 October 2020. Retrieved 14 June 2017.
  291. ^ Sea Anemones, Sea Anemone Pictures Archived 16 January 2017 at the Wayback Machine. Northrup.org
  292. ^ Tube Anemones – Ceriantharia. Seawater.no
  293. ^ Praya picture. Lifesci.ucsb.edu
  294. ^ Portuguese Man-of-Wars, Portuguese Man-of-War Pictures, Portuguese Man-of-War Facts. Animals.nationalgeographic.com
  295. ^ "Longest Giant Stringy Sea Creature Ever Recorded Looks like It Belongs in Outer Space". interestingengineering.com. 9 April 2020. Retrieved 10 April 2020.
  296. ^ Schmidt Ocean Institute (9 April 2020). "New species discovered during exploration of abyssal deep sea canyons off Ningaloo". EurekAlert!. Retrieved 12 April 2020.
  297. ^ Biggest, Smallest, Fastest, Deepest: Marine Animal Records – Marine Biology: Life in the Ocean. Care2.com (4 March 2009)
  298. ^ Xestospongia muta. Encyclopedia of Life
  299. ^ Yahel, G.; Eerkes-Medrano, D. I.; Leys, S. P. (24 November 2006). "Size independent selective filtration of ultraplankton by hexactinellid glass sponges". Aquatic Microbial Ecology. 45: 181–194. doi:10.3354/AME045181. ISSN 0948-3055. Wikidata Q56915355.
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