List of birds of Uruguay

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The Southern Lapwing is the national bird of Uruguay.

This is a list of the bird species recorded in Uruguay. The avifauna of Uruguay has 475 species, of which five have been introduced by humans, and 46 are rare or accidental. Eighteen species are globally threatened.

This list's taxonomic treatment (designation and sequence of orders, families, and species) and nomenclature (common and scientific names) follow the conventions of Clements's 5th edition. The family accounts at the beginning of each heading reflects this taxonomy, as do the species counts found in each family account. Introduced and accidental species are included in the total counts for Uruguay.

The following tags have been used to highlight certain categories, but not all species fall into one of these categories. Those that do not are commonly occurring, native species.

  • (A) Accidental A species that rarely or accidentally occurs in Uruguay.
  • (I) Introduced A species introduced to Uruguay as a consequence, direct or indirect, of human actions.


Table of contents

Non-passerines: Rheas . Tinamous . Penguins . Grebes . Albatrosses . Shearwaters and Petrels . Storm-Petrels . Diving petrels . Boobies and Gannets . Cormorants . Darters . Frigatebirds . Bitterns, Herons and Egrets . Storks . Ibises and Spoonbills . Flamingos . Screamers . Ducks, Geese and Swans . New World vultures . Osprey . Hawks, Kites and Eagles . Caracaras and Falcons . Guans, Chachalacas and allies . Limpkins . Rails, Crakes, Gallinules, and Coots . Seriemas . Jacanas . Painted snipe . Oystercatchers . Avocets and Stilts . Plovers and Lapwings . Sandpipers and allies . Seedsnipes . Sheathbills . Skuas and Jaegers . Gulls . Terns . Skimmers . Pigeons and Doves . Parrots, Macaws and allies . Cuckoos and Anis . Barn owls . Typical owls . Potoos . Nightjars . Swifts . Hummingbirds . Trogons and Quetzals . Kingfishers . Woodpeckers and allies .

Passerines: Ovenbirds . Woodcreepers . Typical antbirds . Gnateaters . Cotingas . Tyrant flycatchers . Swallows and Martins . Wagtails and Pipits . Wrens . Mockingbirds and Thrashers . Thrushes and allies . Gnatcatchers . Crows, Jays, Ravens and Magpies . Waxbills and allies . Vireos . New World warblers . Tanagers . Buntings, Sparrows, Seedeaters and allies . Saltators, Cardinals and allies . Troupials and allies . Siskins, Crossbills and allies . Sparrows .

See also       References

Rheas[edit]

Order: Struthioniformes. Family: Rheidae

The rheas are large flightless birds native to South America. Their feet have three toes rather than four which allows them to run faster. One species has been recorded in Uruguay.

Tinamous[edit]

Order: Tinamiformes. Family: Tinamidae

The tinamous are one of the most ancient groups of bird. Although they look similar to other ground-dwelling birds like quail and grouse, they have no close relatives and are classified as a single family Tinamidae within their own order, the Tinamiformes. They are distantly related to the ratites (order Struthioniformes), that includes the rheas, emu, and kiwi. Three species have been recorded in Uruguay.

Penguins[edit]

Order: Sphenisciformes. Family: Spheniscidae

The penguins are a group of aquatic, flightless birds living almost exclusively in the Southern Hemisphere. Most penguins feed on krill, fish, squid, and other forms of sealife caught while swimming underwater. Three species have been recorded in Uruguay.

Grebes[edit]

Order: Podicipediformes. Family: Podicipedidae

Grebes are small to medium-large sized freshwater diving birds. They have lobed toes, and are excellent swimmers and divers. However, they have their feet placed far back on the body, making them quite ungainly on land. Four species have been recorded in Uruguay.

Albatrosses[edit]

Order: Procellariiformes. Family: Diomedeidae

The albatrosses are among the largest of flying birds, and the great albatrosses from the genus Diomedea have the largest wingspans of any extant birds. Six species have been recorded in Uruguay.

Shearwaters and petrels[edit]

Order: Procellariiformes. Family: Procellariidae

The procellariids are the main group of medium-sized 'true petrels', characterised by united nostrils with a medium septum, and a long outer functional primary. Eighteen species have been recorded in Uruguay.

Storm-Petrels[edit]

Order: Procellariiformes. Family: Hydrobatidae

The storm-petrels are relatives of the petrels, and are the smallest of sea-birds. They feed on planktonic crustaceans and small fish picked from the surface, typically while hovering. The flight is fluttering and sometimes bat-like. Five species have been recorded in Uruguay.

Diving petrels[edit]

Order: Procellariiformes. Family: Pelecanoididae

The diving petrels are small auk-like birds found in the southern oceans. They feed on krill, copepods and small fish and squid. One species has been recorded in Uruguay.

Boobies and gannets[edit]

Order: Pelecaniformes. Family: Sulidae

The sulids comprise the gannets and boobies. Both groups comprise medium-to-large coastal sea-birds that plunge-dive for fish. One species has been recorded in Uruguay.

Cormorants[edit]

Order: Pelecaniformes. Family: Phalacrocoracidae

The Phalacrocoracidae is a family of medium-to-large coastal, fish-eating sea-birds that includes cormorants and shags. Plumage colouration varies with the majority having mainly dark plumage, some species being black and white, and a few being colourful. Three species have been recorded in Uruguay.

Darters[edit]

Order: Pelecaniformes. Family: Anhingidae

Darters are frequently referred to as "snake-birds" because of their long thin neck, which gives a snake-like appearance when they swim with their bodies submerged.

The males have black and dark brown plumage, an erectile crest on the nape and a larger bill than the female. The females have a much paler plumage especially on the neck and underparts. The darters have completely webbed feet, and their legs are short and set far back on the body. Their plumage is somewhat permeable, like that of cormorants, and they spread their wings to dry after diving. One species has been recorded in Uruguay.

Frigatebirds[edit]

Order: Pelecaniformes. Family: Fregatidae

Frigatebirds are large sea-birds usually found over tropical oceans. They are large, black-and-white or completely black, with long wings and deeply forked tails. The males have inflatable coloured throat pouches. They do not swim or walk, and cannot take off from a flat surface. Having the largest wingspan-to-body-weight ratio of any bird, they are essentially aerial, able to stay aloft for more than a week. One species has been recorded in Uruguay.

Bitterns, herons and egrets[edit]

Order: Ciconiiformes. Family: Ardeidae

The family Ardeidae contains the bitterns, herons and egrets. Herons and egrets are medium to large sized wading birds with long necks and legs. Bitterns tend to be shorter necked and more wary. Unlike other long-necked birds suck as storks, ibises and spoonbills, members of Ardeidae fly with their necks retracted. Eleven species have been recorded in Uruguay.

Storks[edit]

Order: Ciconiiformes. Family: Ciconiidae

Storks are large, long-legged, long-necked, wading birds with long, stout bills. Storks are mute; bill-clattering is an important mode of stork communication at the nest. Their nests can be large and may be reused for many years. Many species are migratory. Three species have been recorded in Uruguay.

Ibises and spoonbills[edit]

Order: Ciconiiformes. Family: Threskiornithidae

The Threskiornithidae is a family of large terrestrial and wading birds which includes the ibises and spoonbills. They have long, broad wings with 11 primary and about 20 secondary feathers. They are strong fliers and despite their size and weight, very capable soarers. Six species have been recorded in Uruguay.

Flamingos[edit]

Order: Phoenicopteriformes. Family: Phoenicopteridae

Flamingos are gregarious wading birds, usually 3 to 5 feet high, found in both the Western and Eastern Hemispheres. They are more numerous in the latter. Flamingos filter-feed on shellfish and algae. Their oddly shaped beaks are specially adapted to separate mud and silt from the food they consume, and are used upside-down. One species has been recorded in Uruguay.

Screamers[edit]

Order: Anseriformes. Family: Anhimidae

The screamers are a small family of birds related to the ducks. They are large, bulky birds, with a small downy head, long legs and large feet which are only partially webbed. They have large spurs on their wings which are used in fights over mates and territorial disputes. Eight species have been recorded in Uruguay.

Ducks, geese and swans[edit]

Order: Anseriformes. Family: Anatidae

The family Anatidae includes the ducks and most duck-like waterfowl, such as geese and swans. These are birds that are modified for an aquatic existence with webbed feet, flattened bills and feathers that are excellent at shedding water due to an oily coating. Twenty two species have been recorded in Uruguay.

New World vultures[edit]

Order: Falconiformes. Family: Cathartidae

The New World vultures are not closely related to Old World vultures, but superficially resemble them because of convergent evolution. Like the Old World vultures, they are scavengers. However, unlike Old World vultures, which find carcasses by sight, New World vultures have a good sense of smell with which they locate carrion. Four species have been recorded in Uruguay.

Osprey[edit]

Order: Falconiformes. Family: Pandionidae

The Pandionidae family contains only one species, the Osprey. The Osprey is a medium large raptor which is a specialist fish-eater with a worldwide distribution.

Hawks, kites and eagles[edit]

Order: Falconiformes. Family: Accipitridae

Accipitridae is a family of birds of prey and include hawks, eagles, kites, harriers and Old World vultures. These birds have powerful hooked beaks for tearing flesh from their prey, strong legs, powerful talons, and keen eyesight. Nineteen species have been recorded in Uruguay.

Caracaras and falcons[edit]

Order: Falconiformes. Family: Falconidae

Falconidae is a family of diurnal birds of prey. They differ from hawks, eagles, and kites in that they kill with their beaks instead of their feet. Seven species have been recorded in Uruguay.

Guans, Chachalacas and allies[edit]

Order: Galliformes. Family: Cracidae

The Cracidae are large birds, similar in general appearance to turkeys. The guans and curassows live in trees, but the smaller chachalacas are found in more open scrubby habitats. They are generally dull-plumaged, but the curassows and some guans have colourful facial ornaments. Three species have been recorded in Uruguay.

Limpkins[edit]

Order: Gruiformes. Family: Aramidae

The Limpkin resembles a large rail. It has drab brown plumage and a greyer head and neck.

Rails, crakes, gallinules, and coots[edit]

Order: Gruiformes. Family: Rallidae

Rallidae is a large family of small to medium-sized birds which includes the rails, crakes, coots, and gallinules. Typically they inhabit dense vegetation in damp environments near lakes, swamps, or rivers. In general they are shy and secretive birds, difficult to observe. Most species have strong legs, and have long toes which are well adapted to soft, uneven surfaces. They tend to have short, rounded wings and be weak fliers. Fifteen species have been recorded in Uruguay.

Seriemas[edit]

Order: Gruiformes. Family: Cariamidae

The seriemas are terrestrial birds which run rather than fly (though they are able to fly for short distances.) They have long legs, necks, and tails, but only short wings, reflecting their way of life. They are brownish birds with short bills and erectile crests, found on fairly dry open grasslands. One species has been recorded in Uruguay.

Jacanas[edit]

Order: Charadriiformes. Family: Jacanidae

The jacanas are a group of tropical waders in the family Jacanidae. They are found worldwide in the Tropics. They are identifiable by their huge feet and claws which enable them to walk on floating vegetation in the shallow lakes that are their preferred habitat. One species has been recorded in Uruguay.

Painted snipe[edit]

Order: Charadriiformes. Family: Rostratulidae

Painted snipe are short-legged, long-billed birds similar in shape to the true snipes, but more brightly coloured. One species has been recorded in Uruguay.

Oystercatchers[edit]

Order: Charadriiformes. Family: Haematopodidae

The oystercatchers are large and noisy plover-like birds, with strong bills used for smashing or prising open molluscs. Two species have been recorded in Uruguay.

Avocets and stilts[edit]

Order: Charadriiformes. Family: Recurvirostridae

Recurvirostridae is a family of large wading birds, which includes the avocets and the stilts. The avocets have long legs and long up-curved bills. The stilts have extremely long legs and long, thin, straight bills. One species has been recorded in Uruguay.

Plovers and lapwings[edit]

Order: Charadriiformes. Family: Charadriidae

The family Charadriidae includes the plovers, dotterels, and lapwings. They are small to medium-sized birds with compact bodies, short, thick necks and long, usually pointed, wings. They are found in open country worldwide, mostly in habitats near water, although there are some exceptions. Eight species have been recorded in Uruguay.

Sandpipers and allies[edit]

Order: Charadriiformes. Family: Scolopacidae

The Scolopacidae are a large diverse family of small to medium sized shorebirds including the sandpipers, curlews, godwits, shanks, tattlers, woodcocks, snipes, dowitchers and phalaropes. The majority of species eat small invertebrates picked out of the mud or soil. Variation in length of legs and bills enable different species to feed in the same habitat, particularly on the coast, without direct competition for food. Twenty four species which have been recorded in Uruguay.

Seedsnipes[edit]

Order: Charadriiformes. Family: Thinocoridae

The seedsnipes are a small family of birds that resemble sparrows. They have short legs and long wings and are herbivorous waders. One species has been recorded in Uruguay.

Sheathbills[edit]

Order: Charadriiformes. Family: Chionididae

The sheathbills are scavengers of the Antarctic regions. They have white plumage, and look plump and dove-like, but are believed to be similar to the ancestors of the modern gulls and terns. One species has been recorded in Uruguay.

Skuas and jaegers[edit]

Order: Charadriiformes. Family: Stercorariidae

The family Stercorariidae are, in general, medium to large birds, typically with grey or brown plumage, often with white markings on the wings. They nest on the ground in temperate and arctic regions and are long-distance migrants. Six species have been recorded in Uruguay.

Gulls and terns[edit]

Order: Charadriiformes. Family: Laridae

Laridae is a family of medium to large birds seabirds and includes gulls, kittiwakes and terns. They are typically grey or white, often with black markings on the head or wings. They have longish bills and webbed feet. Terns are a group of generally general medium to large sea-birds typically with grey or white plumage, often with black markings on the head. Most terns hunt fish by diving but some pick insects off the surface of fresh water. Terns are generally long-lived birds, with several species living in excess of 25 to 30 years.

Skimmers[edit]

Order: Charadriiformes. Family: Rynchopidae

Skimmers are a small family of tropical tern-like birds. They have an elongated lower mandible which they use to feed by flying low over the water surface and skimming the water for small fish. One species has been recorded in Uruguay.

Pigeons and doves[edit]

Order: Columbiformes. Family: Columbidae

Pigeons and doves are stout-bodied birds with short necks and short slender bills with a fleshy cere. Nine species have been recorded in Uruguay.

Parrots, macaws and allies[edit]

Order: Psittaciformes. Family: Psittacidae

Parrots are small to large birds with a characteristic curved beak shape. Their upper mandibles have slight mobility in the joint with the skull and they have a generally erect stance. All parrots are zygodactyl, having the four toes on each foot placed two at the front and two back. Seven species have been recorded in Uruguay.

Cuckoos and anis[edit]

Order: Cuculiformes. Family: Cuculidae

The family Cuculidae includes cuckoos, roadrunners and anis. These birds are of variable size with slender bodies, long tails and strong legs. Unlike the cuckoo species of the Old World, North American cuckoos are not brood parasites. Eight species have been recorded in Uruguay.

Barn owls[edit]

Order: Strigiformes. Family: Tytonidae

Barn owls are medium to large sized owls with large heads and heart-shaped faces. They have long strong legs with powerful talons. One species has been recorded in Uruguay.

Typical owls[edit]

Order: Strigiformes. Family: Strigidae

Typical owls are small to large solitary nocturnal birds of prey. They have large forward-facing eyes and ears, a hawk-like beak, and a conspicuous circle of feathers around each eye called a facial disk. Nine species have been recorded in Uruguay.

Potoos[edit]

Order: Caprimulgiformes. Family: Nyctibiidae

The potoos (sometimes called Poor-Me-Ones) are large near passerine birds related to the nightjars and frogmouths. They are nocturnal insectivores which lack the bristles around the mouth found in the true nightjars. One species has been recorded in Uruguay.

Nightjars[edit]

Order: Caprimulgiformes. Family: Caprimulgidae

Nightjars are medium-sized nocturnal birds with long wings, short legs and very short bills that usually nest on the ground. Most have small feet, of little use for walking, and long pointed wings. Their soft plumage is camouflaged to resemble bark or leaves. Seven species have been recorded in Uruguay.

Swifts[edit]

Order: Apodiformes. Family: Apodidae

Swifts are small aerial birds, spending the majority of their lives flying. These birds have very short legs and never settle voluntarily on the ground, perching instead only on vertical surfaces. Many swifts have long swept-back wings that resemble a crescent or a boomerang. One species has been recorded in Uruguay.

Hummingbirds[edit]

Order: Trochiliformes. Family: Trochilidae

Hummingbirds are small birds capable of hovering in mid-air due to the rapid flapping of their wings. They are the only birds that can fly backwards. Nine 9 species have been recorded in Uruguay.

Trogons and Quetzals[edit]

Order: Trogoniformes. Family: Trogonidae

The family Trogonidae includes trogons and quetzals. Found in tropical woodlands worldwide, they feed on insects and fruit, and their broad bills and weak legs reflect their diet and arboreal habits. Although their flight is fast, they are reluctant to fly any distance. Trogons have soft, often colourful, feathers with distinctive male and female plumage. One species has been recorded in Uruguay.

Kingfishers[edit]

Order: Coraciiformes. Family: Alcedinidae

Kingfishers are medium-sized birds with large heads, long, pointed bills, short legs, and stubby tails. Three species have been recorded in Uruguay.

Woodpeckers and allies[edit]

Order: Piciformes. Family: Picidae

Woodpeckers are small to medium-sized birds with chisel-like beaks, short legs, stiff tails and long tongues used for capturing insects. Some species have feet with two toes pointing forward, and two backward, while several species have only three toes. Many woodpeckers have the habit of tapping noisily on tree trunks with their beaks. Nine species have been recorded in Uruguay.

Ovenbirds[edit]

Order: Passeriformes. Family: Furnariidae

Ovenbirds comprise a large family of small sub-oscine passerine bird species found in Central and South America. They are a diverse group of insectivores which gets its name from the elaborate "oven-like" clay nests built by some species, although others build stick nests or nest in tunnels or clefts in rock. THirty species have been recorded in Uruguay.

Woodcreepers[edit]

Order: Passeriformes. Family: Dendrocolaptidae

The Dendrocolaptidae are brownish birds and maintain an upright vertical posture, supported by their stiff tail vanes. They feed mainly on insects taken from tree trunks. Four species have been recorded in Uruguay.

Typical antbirds[edit]

Order: Passeriformes. Family: Thamnophilidae

The antbirds are a large family of small passerine birds of subtropical and tropical Central and South America. They are forest birds, and tend to feed on insects at or near the ground. A sizable minority of them specialize in following columns of army ants to eat the small invertebrates that leave hiding to flee the ants. Many species lack bright colour; brown, black and white being the dominant tones. Two species have been recorded in Uruguay.

Gnateaters[edit]

Order: Passeriformes. Family: Conopophagidae

The gnateaters are round, short-tailed, and long-legged birds, which are closely related to the antbirds. Three species have been recorded in Uruguay.

Cotingas[edit]

Order: Passeriformes. Family: Cotingidae

The cotingas are birds of forests or forest edges of tropical South America. Comparatively little is known about this diverse group, although all have broad bills with hooked tips, rounded wings, and strong legs. The males of many of the species are brightly coloured, or decorated with plumes or wattles. One species has been recorded in Uruguay.

Tyrant flycatchers[edit]

Order: Passeriformes. Family: Tyrannidae

Tyrant flycatchers are passerine birds which occur throughout North and South America. They superficially resemble the Old World flycatchers, but are more robust with stronger bills. They do not have the sophisticated vocal capabilities of the songbirds. Most, but not all, have plain colouring. As the name implies, most are insectivorous. Fifty seven species have been recorded in Uruguay.

Becards and tityras[edit]

Order: Passeriformes. Family: Tityridae

Swallows and martins[edit]

Order: Passeriformes. Family: Hirundinidae

The Hirundinidae family is a group of passerines characterized by their adaptation to aerial feeding. Their adaptations include a slender streamlined body, long pointed wings and short bills with wide gape. The feet are designed for perching rather than walking, and the front toes are partially joined at the base. Eleven species have been recorded in Uruguay.

Wagtails and pipits[edit]

Order: Passeriformes. Family: Motacillidae

The Motacillidae are a family of small passerine birds with medium to long tails. They include the wagtails, longclaws and pipits. They are slender, ground feeding insectivores of open country. Four species have been recorded in Uruguay.

Wrens[edit]

Order: Passeriformes. Family: Troglodytidae

The wrens are mainly small and inconspicuous except for their loud songs. These birds have short wings and a thin down-turned bill. Several species often hold their tails upright. All are insectivorous. Two species have been recorded in Uruguay.

Mockingbirds and Thrashers[edit]

Order: Passeriformes. Family: Mimidae

The mimids are a family of passerine birds that includes thrashers, mockingbirds, tremblers, and the New World catbirds. These birds are notable for their vocalizations, especially their ability to mimic a wide variety of birds and other sounds heard outdoors. Their colouring tends towards dull greys and browns. Three species have been recorded in Uruguay.

Thrushes and allies[edit]

Order: Passeriformes. Family: Turdidae

The thrushes are a group of passerine birds that occur mainly in the Old World. They are plump, soft plumaged, small to medium-sized insectivores or sometimes omnivores, often feeding on the ground. Many have attractive songs. Four species have been recorded in Uruguay.

Gnatcatchers[edit]

Order: Passeriformes. Family: Polioptilidae

These dainty birds resemble Old World warblers in their build and habits, moving restlessly through the foliage seeking insects. The gnatcatchers and gnatwrens are mainly soft bluish grey in colour, and have the typical insectivore's long sharp bill. They are birds of fairly open woodland or scrub, and nest in bushes or trees. One species has been recorded in Uruguay.

Crows, jays, ravens and magpies[edit]

Order: Passeriformes. Family: Corvidae

The Corvidae family includes crows, ravens, jays, choughs, magpies, treepies, nutcrackers, and ground jays. Corvids are above average in size for the bird order Passeriformes. Some of the larger species show high levels of learning behavior. Three species have been recorded in Uruguay.

Waxbills and allies[edit]

Order: Passeriformes. Family: Estrildidae

The estrildid finches are small passerine birds of the Old World tropics and Australasia. They are gregarious and often colonial seed-eaters with short thick but pointed bills. They are all similar in structure and habits, but have a wide variation in plumage colours and pattern. One species has been recorded in Uruguay.

Vireos[edit]

Order: Passeriformes. Family: Vireonidae

The vireos are a group of small to medium sized passerine birds restricted to the New World. They are typically greenish in colour and resemble wood warblers apart from their heavier bills. Two species have been recorded in Uruguay.

New World warblers[edit]

Order: Passeriformes. Family: Parulidae

The New World warblers are a group of small, often colourful, passerine birds restricted to the New World. Most are arboreal, but some are terrestrial. Most members of this family are insectivores. Five species have been recorded in Uruguay.

Tanagers[edit]

Order: Passeriformes. Family: Thraupidae

The tanagers are a large group of small to medium-sized passerine birds restricted to the New World, mainly in the tropics. Many species are brightly coloured. They are seed eaters, but have a preference for fruit and nectar. Most have short, rounded wings.

Buntings, sparrows, and allies[edit]

Order: Passeriformes. Family: Emberizidae

The emberizids are a large family of passerine birds. They are seed-eating birds with a distinctively shaped bill. In Europe, most species are named as buntings. In North America, most of the species in this family are known as Sparrows, but these birds are not closely related to the Old World sparrows which are in the family Passeridae. Many emberizid species have distinctive head patterns. Three species have been recorded in Uruguay.

Saltators, Cardinals and allies[edit]

Order: Passeriformes. Family: Cardinalidae

The cardinals are a family of passerine birds that are robust, seed-eating birds, with strong bills. They are typically associated with open woodland. The sexes usually have distinct plumages. Five species have been recorded in Uruguay.

Troupials and allies[edit]

Order: Passeriformes. Family: Icteridae

The icterids are a group of small to medium, often colourful, passerine birds restricted to the New World and include the grackles, New World blackbirds, and New World orioles. Most species have black as the predominant plumage colour, often enlivened by yellow, orange or red. Seventeen species have been recorded in Uruguay.

Siskins, crossbills and allies[edit]

Order: Passeriformes. Family: Fringillidae

Finches are seed-eating passerine birds, that are small to moderately large and have a strong beak, usually conical and in some species very large. All have 12 tail feathers and 9 primaries. These birds have a bouncing flight with alternating bouts of flapping and gliding on closed wings, and most sing well. Six species have been recorded in Uruguay.

Sparrows[edit]

Order: Passeriformes. Family: Passeridae

Sparrows are small passerine birds. Sparrows tend to be small, plump, brown or grey birds with short tails and short powerful beaks. Sparrows are seed-eaters, and they also consume small insects. One species has been recorded in Uruguay.

See also[edit]

References[edit]

External links[edit]

  • Birds of Uruguay Birdlist, multi-lingual website by country with standardized codes for abundance and seasonal presence.