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Micropaleontology (also sometimes spelled as micropalaeontology) is the branch of palaeontology that studies microfossils.
Microfossils are fossils generally not larger than four millimeters, and commonly smaller than one millimeter, the study of which requires the use of light or electron microscopy. Fossils which can be studied with the naked eye or low-powered magnification, such as a hand lens, are referred to as macrofossils. Obviously, it can be hard to decide whether or not some organisms should be considered microfossils, as there is no fixed size boundary.
For example, some colonial organisms, such as Bryozoa (especially the Cheilostomata) have relatively large colonies, but are classified on the basis of fine skeletal details of the tiny individuals of the colony.
Microfossils are a common feature of the geological record, from the Precambrian to the Holocene. They are most common in deposits of marine environments, but also occur in brackish water, fresh water and terrestrial sedimentary deposits. While every kingdom of life is represented in the microfossil record, the most abundant forms are protist skeletons or cysts from the Chrysophyta, Pyrrhophyta, Sarcodina, acritarchs and chitinozoans, together with pollen and spores from the vascular plants.
Areas of study
Micropaleontology can be roughly divided into four areas of study on the basis of microfossil composition: (a) calcareous, as in coccoliths and foraminifera, (b) phosphatic, as in the study of some vertebrates, (c) siliceous, as in diatoms and radiolaria, or (d) organic, as in the pollen and spores studied in palynology.
This division reflects differences in the mineralogical and chemical composition of microfossil remains (and therefore in the methods of fossil recovery) rather than any strict taxonomic or ecological distinctions. Most researchers in this field, known as micropaleontologists, are typically specialists in one or more taxonomic groups.
Phosphatic microfossils include Conodonts (tiny oral structures of an extinct chordate group), some scolecodonts ("worm" jaws), Shark spines and teeth, and other Fish remains (collectively called "ichthyoliths").
The study of organic microfossils is called palynology. Organic microfossils include pollen, spores, Chitinozoans (thought to be the egg cases of marine invertebrates), Scolecodonts ("worm" jaws), Acritarchs, Dinoflagellate cysts, and fungal remains.
Sediment or rock samples are collected from either cores or outcrops, and the microfossils they contain are extracted by a variety of physical and chemical laboratory techniques, including sieving, density separation by centrifuge or in heavy liquids, and chemical digestion of the unwanted fraction. The resulting concentrated sample of microfossils is then mounted on a slide for analysis, usually by light microscope. Taxa are then identified and counted. The very large numbers of microfossils that a small sediment sample can often yield allows the collection of statistically robust datasets which can be subjected to multivariate analysis. A typical microfossil study will involve identification of a few hundred specimens from each sample.
Applications of micropaleontology
Microfossils are especially noteworthy for their importance in biostratigraphy. Since microfossils are often extremely abundant, widespread, and quick to appear and disappear from the stratigraphic record, they constitute ideal index fossils from a biostratigraphic perspective. In addition, the planktonic and nektonic habits of some microfossils gives them the added bonus of appearing across a wide range of facies or paleoenvironments, as well as having near-global distribution, making biostratigraphic correlation even more powerful and effective.
Microfossils, particularly from deep-sea sediments, also provide some of the most important records of global environmental change on long, medium or short timescales. Across vast areas of the ocean floor, the shells of planktonic micro-organisms sinking from surface waters provide the dominant source of sediment, and they continuously accumulate (typically at rates of 20-50 million per million years). Study of changes in assemblages of microfossils and of changes in their shell chemistry (e.g., oxygen isotope composition) are fundamental to research on climate change in the geological past.
In addition to providing an excellent tool for sedimentary rock dating and for paleoenvironmental reconstruction – heavily used in both petroleum geology and paleoceanography – micropaleontology has also found a number of less orthodox applications, such as its growing role in forensic police investigation or in determining the provenance of archaeological artefacts.
Micropaleontology is also a tool of Geoarchaeology used in archaeological reconstruction of human habitation sites and environments. Changes in the microfossil population abundance in the stratigraphy of current and former water bodies reflect changes in environmental conditions. Naturally occurring Ostracods in freshwater bodies are impacted by changes in salinity and pH due to human activities. When correlated with other dating techniques, prehistoric environments can be reconstructed. Work on Lake Tanganyika provided a profile of human induced environmental changes of a 4,000 year period.
Similar work in the arid American Southwest has provided information on irrigation canals used by prehistoric peoples from 2100 B.C. to 500 B.C. Other archaeological work in arid climates throughout the Americas has incorporated Micropaleontological analysis to build a more complete picture of prehistoric climate and human activity.
- Brasier, M.D. (1980), Microfossils. Chapman and Hall publishers. ISBN 0-412-44570-0
- Traverse, A. (1988), Paleopalynology. Unwin Hyman ISBN 0-04-561001-0
- Ovechkina, M.N. 2007. Calcareous nannoplankton of the Upper Cretaceous (Campanian and Maastrichtian) of the southern and eastern Russian Platform. Nauka, Moscow. 353 pp. ISBN 978-5-02-035758-7 
- M. Palacios-Fest (2005). "Paleolimnological investigations of anthropogenic environmental change in Lake Tanganyika: IV. Lacustrine paleoecology" (PDF). Journal of Paleolimnology 34: 51–71. doi:10.1007/s10933-005-2397-1.
- M. Palacios-Fest & Owen K. Davis (2008). 8 (PDF). "Las Capas: Early Irrigation and Sedentism in a Southwestern Floodplain". Anthropological Papers No. 28, ed. Jonathan B. Mabry (Center for Desert Archaeology, Tucson, Arizona, USA). pp. 197–220.
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- The Micropalaeontological Society
- Centre for Palynology, University of Sheffield, UK
- The American Association of Stratigraphic Palynologists (AASP)
- Commission Internationale de Microflore du Paléozoique (CIMP), international commission for Palaeozoic palynology.
- The International Nannoplankton Association
- CIMP Subcommission on Acritarchs
- CIMP Chitinozoan Subcommission
- Micropaleontology Press, a division of the Micropaleontology Project, a nonprofit organization with the mission of promoting the science of micropaleontology.
- University of California, Berkeley, microfossil collections.
- The Ocean Drilling Programme (ODP) Micropaleontological reference centres.
- University College, London, Micropalaeontology Unit, including the MIRACLE microfossil image database.
- 2500+ images of Foraminifera with a searchable database
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