The Mydidae (alternative spelling Mydaidae), or Mydas flies, are a cosmopolitan family of flies. It is a small family, with about 471 species described. They are generally large in size, including, in fact, the largest known fly, Gauromydas heros (syn. Mydas heros). Many of the species, in addition to their large size, are mimics of stinging hymenopterans, especially wasps. Most mydids are found in arid and semi-arid regions of the world, but they are also found in other habitats. They are infrequently encountered as the adult lifespan can be quite short. Little is known about their biology, though the larvae of some species appear to be subterranean predators of ants.
For terms see Morphology of Diptera.
Mydids are medium-sized to very large flies (9-60 millimeters in body length). The abdomen is long and cylindrical in section. It is slightly tapered apically in the male, and usually widest at segment 4 in the female. The second segment of the antenna forms a club. Mydids are sparsely pilose, and lack bristles except on the legs. The hindleg is much longer and stronger than either the middle leg or the foreleg and the hind femur is usually swollen and bears ventral spines. The hind tibia has an apical spur or bristles. The wings are long, and narrow to wide. Most of the veins end in the upper margin before the apex.
The classification of the family has changed fairly recently, with the inclusion of a few genera that were previously placed in the family Apioceridae. Apiocerids have long been given the common name "flower-loving flies", but only the group of "flower-loving flies" that actually visit flowers were transferred to the Mydidae. Among the genera transferred is Rhaphiomidas, which includes one of the few Diptera on the United States' Endangered Species List, the Delhi Sands flower-loving fly (Rhaphiomidas terminatus abdominalis).
- Lyons, K. M. and T. Dikow. (2010). Taxonomic revision of Ectyphus Gerstaecker, 1868 and Parectyphus Hesse, 1972 with a key to world Ectyphinae (Insecta, Diptera, Mydidae). ZooKeys 73 25-59.
- Bequaert, M. 1963. Contribution a la connaissance morphologique et a la classification des Mydaidae (Diptera). Bull. Inst. Roy. Sciences Natur. Belg. 37:1-18.
- Oosterbroek, P. 1998. 41. Mydidae. In P. Oosterbroek: The families of Diptera of the Malay Archipelago. Brill: Boston. p. 95.
- Sack P. (1934) 23. Mydaidae. In: Lindner E. (Ed) Die Fliegen der palaearktischen Region. Band 4(5), 1–29. Schweizerbart‘sche Verlagsbuchhandlung, Stuttgart, 1–29.Keys to Palaearctic species but now needs revision (in German).
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