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Pinnate refers to feather-like or multi-divided features arising from both sides of a common axis. In biological morphology the term applies to plant or animal structures, but it also is used in other disciplines, for example in describing crystalline structures such as some forms of ice or metal crystals, and in earth sciences, where it may refer to patterns of erosion or stream beds.
The term derives from the Latin word pinna meaning "feather", "wing", or "fin". A similar term is pectinate, which refers to a comb-like arrangement of parts (arising from one side of an axis only). The term "pinnate" is used largely in contrast to palmate, in which the parts or structures radiate out from a common point. The term pennate is cognate with "pinnate" and is variously used, but there is no consistent difference in the meaning or usage of the two words.
Botanically, the term describes an arrangement of discrete structures (such as leaflets, veins, lobes, branches, or appendages) arising at multiple points along a common axis. For example, once-divided leaf blades having leaflets arranged on both sides of a rachis are pinnately compound leaves. Many palms (notably the feather palms) and most cycads and grevilleas have pinnately divided leaves. Most species of ferns have pinnate or more highly divided fronds, and in ferns the leaflets or segments are typically referred to as "pinnae" (singular "pinna"). Plants with pinnate leaves are sometimes colloquially called "feather-leaved". Most of the following definitions are from Jackson's Glossary of Botanical Terms:
- pinnatifid and pinnatipartite: leaves with pinnate lobes that are not discrete, remaining sufficiently connected to each other that they are not separate leaflets.
- pinnatisect: cut all the way to the midrib or other axis, but with the bases of the pinnae not contracted to form discrete leaflets.
- pinnate-pinnatifid: pinnate, with the pinnae being pinnatifid.
- paripinnate: pinnately compound leaves in which leaflets are born in pairs along the rachis without a single terminal leaflet; also called "even-pinnate".
- imparipinnate: pinnately compound leaves in which there is a lone terminal leaflet rather than a terminal pair of leaflets; also called "odd-pinnate".
- bipinnate: pinnately compound leaves in which the leaflets are themselves pinnately compound; also called "twice-pinnate".
- tripinnate: pinnately compound leaves in which the leaflets are themselves bipinnate; also called "thrice-pinnate".
- tetrapinnate: pinnately compound leaves in which the leaflets are themselves tripinnate.
- unipinnate: solitary compound leaf with a row of leaflets arranged along each side of a common rachis.
- The term pinnula is the Latin diminutive of pinna; either as such or in the Anglicised form: pinnule, it is differently defined by various authorities. Some apply it to the leaflets of a pinna, especially the leaflets of bipinnate or tripinnate leaves. Others also or alternatively apply it second or third order divisions of a bipinnate or tripinnate leaf.
In animals, pinnate is used to describe feather-like structures:
- Some muscles can be unipinnate or bipinnate muscles
- The fish, Platax pinnatus, is known as the pinnate spadefish or pinnate batfish. Plus the "pinnate bird" is also covered in feathers that look like leaves.
The term is also used to describe certain waterway systems in which all major tributary streams enter the main channels by flowing in one direction at an oblique angle.
- Nicholas Eastaugh, Valentine Walsh, Tracey Chaplin, Ruth Siddall. "Pigment Compendium: A Dictionary of Historical Pigments". Pub: Butterworth-Heinemann 2008. ISBN 978-0750689809
- Charles Seymour Wright, Raymond Edward Priestley. Glaciology. Pub: Harrison and Sons, for the Committee of the Captain Scott Arctic fund, 1922
- Journal of the Electrochemical Society, Volume 100, 1953, page 165: "The zinc is recovered electrolytically as "flake" powder consisting of pinnate crystals."
- Ravi P. Gupta. Remote Sensing Geology. Publisher: Springer 2003. ISBN 978-3540431855
- Collocott, T. C. (ed.) (1974). Chambers Dictionary of science and technology. Edinburgh: W. and R. Chambers. ISBN 0550132023.
- Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928
- Chittenden, Fred J. Ed., Royal Horticultural Society Dictionary of Gardening, Oxford 1951
- Shastri, Varun. Dictionary of Botany. Publisher: Isha Books 2005. ISBN 978-8182052253