In the compound eye of invertebrates such as insects and crustaceans, the pseudopupil appears as a dark spot which moves across the eye as the animal is rotated. This occurs because the ommatidia which one observes "head-on" (along their optical axes) absorb the incident light, while those to one side reflect it. The pseudopupil therefore reveals which ommatidia are aligned with the axis along which the observer is viewing.
- M. F. Land, G. Gibson, J. Horwood & J. Zeil (1999). "Fundamental differences in the optical structure of the eyes of nocturnal and diurnal mosquitoes" (PDF). Journal of Comparative Physiology A 185 (1): 91–103. doi:10.1007/s003590050369.
- Jochen Zeil & Maha M. Al-Mutairi (1996). "Variations in the optical properties of the compound eyes of Uca lactea annulipes" (PDF). The Journal of Experimental Biology 199 (7): 1569–1577. PMID 9319471.
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