Amelanchier

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Amelanchier
Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Eudicots
(unranked): Rosids
Order: Rosales
Family: Rosaceae
Subfamily: Amygdaloideae[1]
Tribe: Maleae
Subtribe: Malinae
Genus: Amelanchier
Medik.
Species

About 20; see text

Amelanchier (/æməˈlænʃɪər/ am-ə-LAN-sheer),[2] also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry, wild pear, juneberry, saskatoon, sugarplum or wild-plum, and chuckley pear is a genus of about 20 species of deciduous-leaved shrubs and small trees in the Rose family (Rosaceae).

Amelanchier is native to temperate regions of the Northern Hemisphere, growing primarily in early successional habitats. It is most diverse taxonomically in North America, especially in the northeastern United States and adjacent southeastern Canada, and at least one species is native to every U.S. state except Hawaii and to every Canadian province and territory. Two species also occur in Asia, and one in Europe. The taxonomic classification of shadbushes has long perplexed botanists, horticulturalists, and others, as suggested by the range in number of species recognized in the genus, from 6 to 33, in two recent publications.[3][4] A major source of complexity comes from the occurrence of hybridization, polyploidy, and apomixis (asexual seed production), making species difficult to characterize and identify.[5]

The various species of Amelanchier grow to 0.2–20 m tall; some are small trees, some are multistemmed, clump-forming shrubs, and yet others form extensive low shrubby patches (clones). The bark is gray or less often brown, and in tree species smooth or fissuring when older. The leaves are deciduous, cauline, alternate, simple, lanceolate to elliptic to orbiculate, 0.5–10 x 0.5–5.5 cm, thin to coriaceous, with surfaces above glabrous or densely tomentose at flowering, and glabrous or more or less hairy beneath at maturity. The inflorescences are terminal, with 1–20 flowers, erect or drooping, either in clusters of one to four flowers, or in racemes with 4–20 flowers. The flowers have five white (rarely somewhat pink, yellow, or streaked with red), linear to orbiculate petals, 2.6–25 mm long, with the petals in one species (A. nantucketensis) often andropetalous (bearing apical microsporangia adaxially). The flowers appear in early spring, "when the shad run" according to tradition (leading to names such as "shadbush"). The fruit is a berry-like pome, red to purple to nearly black at maturity, 5–15 mm diameter, insipid to delectably sweet, maturing in summer.[5]

Amelanchier plants are valued horticulturally, and their fruits are important to wildlife.

Selected species[edit]

Amelanchier alnifolia

For North American species, the taxonomy follows the forthcoming Flora of North America;[5][6] for Asian species the Flora of China;[7] and for European species the Flora Europaea.[8]

Since classifications have varied greatly over the past century, species names are often used interchangeably in the nursery trade. Several natural or horticultural hybrids also exist, and many A. arborea and A. canadensis plants that are offered for sale are actually hybrids, or entirely different species.

A taxon called Amelanchier lamarckii (or A. x lamarckii) is very widely cultivated and naturalized in Europe, where it was introduced in the 17th century. It is apomictic, breeding true from seed, and probably of hybrid origin, perhaps descending from a cross between A. laevis and either A. arborea or A. canadensis. While A. lamarckii is known to be of North American origin, probably from eastern Canada, it is not known to occur naturally in the wild in North America.[22][23]

Etymology[edit]

The origin of the generic name Amelanchier is probably derived from amalenquièr, amelanchièr, the Provençal names of the European Amelanchier ovalis.[24]

The name serviceberry comes from the similarity of the fruit to the related European Sorbus; it is also said that their flowers heralded the roads in the Appalachian mountains becoming passable, which meant that the circuit-riding preachers would be coming soon and church services would resume; also, that the ground was thawed enough to dig graves, and funeral services could be had for those who died over the winter.

Juneberry refers to the fruits of certain species becoming ripe in June. The name saskatoon originated from a Cree Indian noun misâskwatômina (misāskwatōmina, misaaskwatoomina) for Amelanchier alnifolia. The city of Saskatoon, Saskatchewan is named after this plant.

Shadberry refers to the shad runs in certain New England streams, which generally took place about when the trees bloomed.

Ecology[edit]

Amelanchier plants are preferred browse for deer and rabbits, and heavy browsing pressure can suppress natural regeneration. Caterpillars of such Lepidoptera as Brimstone Moth, Brown-tail, Grey Dagger, Mottled Umber, Rough Prominent, The Satellite, Winter Moth, and the Red-Spotted Purple and the White Admiral (both Limenitis arthemis), as well as various other herbivorous insects feed on Amelanchier. Many insects and diseases that attack orchard trees also affect this genus, in particular trunk borers and Gymnosporangium rust. In years when late flowers of Amelanchier overlap those of wild roses and brambles, bees may spread bacterial fireblight.

Fruit and leaves of Amelanchier ovalis

Uses and cultivation[edit]

The fruit of several species are excellent to eat raw, tasting somewhat like a blueberry, strongly accented by the almond-like flavour of the seeds. Selections from Amelanchier alnifolia have been chosen for fruit production, with several named cultivars.[25] Other cultivars appear to be derived from hybridization between A. alnifolia and A. stolonifera.[25]

Fruit is harvested locally for pies and jams.[26] The saskatoon berry is harvested commercially. One version of the Native American food pemmican was flavored by serviceberry fruits in combination with minced dried meat and fat.

The wood is brown, hard, close-grained, and heavy. The heartwood is reddish-brown, and the sapwood is lighter in color. It can be used for tool handles and fishing rods. Native Americans used it for arrow shafts.

Propagation is by seed, divisions, and grafting. Serviceberries graft so readily that grafts onto other genera, such as Crataegus and Sorbus, are often successful.

Members of the Pit River Tribe would use the wood to create a sort of body armor, the wood was made into a heavy robe or overcoat and corset armor and used for fighting.[27]

Garden history[edit]

Several species are very popular ornamental shrubs, grown for their flowers, bark, and fall color. All need similar conditions to grow well, requiring good drainage, air circulation (to discourage leaf diseases), watering during drought, and soil appropriate for the species.

George Washington planted specimens of Amelanchier on the grounds of his estate, Mount Vernon, in Virginia.[citation needed]

References[edit]

  1. ^ Potter, D., et al. (2007). Phylogeny and classification of Rosaceae. Plant Systematics and Evolution. 266(1–2): 5–43. [Referring to the subfamily by the name "Spiraeoideae"]
  2. ^ "amelanchier". Oxford English Dictionary (3rd ed.). Oxford University Press. September 2005. 
  3. ^ Landry, P. (1975). Le concept d'espece et la taxonomie du genre Amelanchier (Rosacees). Bull. Soc. Bot. France 122: 43-252.
  4. ^ Phipps, J. B., Robertson, K. R., Smith, P. G., & Rohrer, J. R. (1990). A checklist of the subfamily Maloideae (Rosaceae). Canad. J. Bot. 68: 2209–2269.
  5. ^ a b c University of Maine: Amelanchier Systematics and Evolution
  6. ^ Campbell, C. S., Dibble, A. C., Frye, C. T., & Burgess, M. B. (2008; accepted for publication). Amelanchier. In FNA Editorial Committee, Flora of North America 9. Magnoliophyta: Rosidae (in part): Rosales (in part). Oxford University Press, New York.
  7. ^ Flora of China: Amelanchier
  8. ^ Flora Europaea: Amelanchier
  9. ^ University of Maine: Amelanchier alnifolia var. alnifolia
  10. ^ University of Maine: Amelanchier amabilis
  11. ^ University of Maine: Amelanchier arborea
  12. ^ University of Maine: Amelanchier bartramiana
  13. ^ University of Maine: Amelanchier canadensis var. canadensis
  14. ^ University of Maine: Amelanchier humilis
  15. ^ University of Maine: Amelanchier interior
  16. ^ University of Maine: Amelanchier laevis
  17. ^ Flora Europaea: Amelanchier ovalis
  18. ^ University of Maine: Amelanchier sanguinea
  19. ^ Flora of China: Amelanchier sinica
  20. ^ University of Maine: Amelanchier spicata
  21. ^ University of Maine: Amelanchier utahensis
  22. ^ Bean, W. J. (1976). Trees and Shrubs Hardy in the British Isles 8th ed., vol. 1. John Murray ISBN 0-7195-1790-7.
  23. ^ Rushforth, K. (1999). Trees of Britain and Europe. Collins ISBN 0-00-220013-9.
  24. ^ Jepson Flora: Amelanchier alnifolia
  25. ^ a b American Society for Horticultural Science (1997). The Brooks and Olmo Register of Fruit & Nut Varieties, 3rd ed. ASHS Press. 
  26. ^ http://www.youtube.com/watchv=wokPJzsV730&feature=channel_page
  27. ^ Merriam, C. Hart 1966 Ethnographic Notes on California Indian Tribes. University of California Archaeological Research Facility, Berkeley (p. 222)

External links[edit]

  • Juneberry, in What Am I Eating? A Food Dictionary