Taxonomy of Lucanidae
The Lucanidae are a family of beetles that include the stag beetles. The family can be further subdivided in a taxonomy. The classification presented here follows Smith (2006),[1] with the exception of the tribal classification within the Lucaninae.
Subfamily Aesalinae[edit]
Subfamily Aesalinae
- Tribe Aesalini
- Genus Aesalus (Fabricius, 1801)
- Aesalus asiaticus Lewis, 1883
- Aesalus himalayicus Kurosawa, 1985
- Aesalus imanishii Inahara & Ratti, 1991
- Aesalus meridionalis Bartolozzi, 1989
- Aesalus neotropicalis Bates, 1886
- Aesalus saburoi
- Aesalus sawaii Fujita & Ichikawa, 1985
- Aesalus scarabaeoides Panzer, 1793
- Aesalus sichuanensis
- Aesalus smithi Bates, 1889
- Aesalus trogoides Albers, 1883
- Aesalus ulanowskii Ganglbauer, 1886
- Genus Cretaesalus (fossil)
- Genus Echinoaesalus Zelenka, 1993
- Echinoaesalus barriesi (Zelenka, 1993)
- Echinoaesalus hidakai (Araya, 1993)
- Echinoaesalus jaechi (Zelenka,1993)
- Echinoaesalus matsuii (Araya, 1993)
- Echinoaesalus sabahensis (Zelenka, 1994)
- Echinoaesalus schuhi (Zelenka, 1994)
- Echinoaesalus timidus (Krikken, 1975)
- Echinoaesalus yongi (Araya, 1993)
- Genus Lucanobium Howden & Lawrence, 1974
- Lucanobium squamosum Howden & Lawrence, 1974
- Genus Aesalus (Fabricius, 1801)
- Tribe Ceratognathini
- Genus Ceratognathus Westwood, 1838
- Ceratognathus abdominalis Parry, 1870
- Ceratognathus bitumulatus Carter, 1925
- Ceratognathus flabellatus Boileau, 1905
- Ceratognathus frenchi Blackburn, 1897
- Ceratognathus froggatti Blackburn, 1894
- Ceratognathus gilesi Blackburn, 1895
- Ceratognathus macrognathus Boileau, 1905
- Ceratognathus mentiferus Westwood, 1863
- Ceratognathus minutus Lea, 1929
- Ceratognathus niger Westwood, 1838
- Ceratognathus ocularis Carter, 1925
- Ceratognathus rufipennis Westwood, 1872
- Ceratognathus tasmanus Benesh, 1943
- Ceratognathus westwoodi Thomson, 1862
- Genus Hilophyllus Paulsen & Mondaca, 2006
- Hilophyllus argentinensis (Martinez, 1981)
- Hilophyllus martinezi Paulsen & Mondaca, 2006
- Hilophyllus penai (Martinez, 1976)
- Genus Mitophyllus Parry, 1843
- Mitophyllus alboguttatus (Bates, 1867)
- Mitophyllus dispar (Sharp, 1882)
- Mitophyllus foveolatus (Broun, 1880)
- Mitophyllus gibbosus (Broun, 1884)
- Mitophyllus insignis Broun, 1921
- Mitophyllus irroratus Parry, 1842
- Mitophyllus macrocerus (Broun, 1886)
- Mitophyllus parrianus Westwood, 1863
- Mitophyllus reflexus Broun, 1908
- Genus Ceratognathus Westwood, 1838
- Tribe Nicagini
- Genus Nicagus LeConte, 1861
- Nicagus obscurus (LeConte, 1848)
- Nicagus occultus Paulsen & Smith, 2005
- Nicagus japonicus Nagel, 1928
- Genus Holloceratognathus Nikolajev, 1998
- Holloceratognathus cylindricus (Broun, 1895)
- Holloceratognathus helotoides (Thomson, 1862)
- Holloceratognathus passaliformis (Holloway, 1962)
- Genus Nicagus LeConte, 1861
Subfamily Lampriminae[edit]
Subfamily Lampriminae
- Genus Dendroblax White, 1846
- Dendroblax earlianus White, 1846
- Genus Homolamprima
- Homolamprima crenulata (MacLeay, 1885)
- Genus Lamprima
- Lamprima adolphinae (Gestro, 1875)
- Lamprima aenea Fabricius, 1792
- Lamprima aurata Latreille, 1817
- Lamprima imberbis Barber, 1926
- Lamprima insularis Macleay, 1885
- Genus Phalacrognathus
- Phalacrognathus muelleri MacLeay, 1885
- Genus Streptocerus Dejean, 1833
- Streptocerus speciosus Fairmaire, 1850
- Genus Dendroblax White, 1846
Subfamily Lucaninae[edit]
Subfamily Lucaninae Latreille, 1804
- Tribe Aegini
- Tribe Allotopini
- Genus Allotopus
- Allotopus moellenkampi (Fruhstorfer, 1894)
- Allotopus rosenbergi (Vollenhoven, 1872)
- Genus Mesotopus
- Genus Allotopus
- Tribe Chiasognathini
- Genus Casignetus
- Genus Chiasognathus Stephens, 1831
- Chiasognathus beneshi Lacroix, 1978
- Chiasognathus grantii Stephens, 1831
- Chiasognathus jousselini Reiche, 1850
- Chiasognathus latreillei Solier, 1851
- Chiasognathus mniszechii Kiesche 1919
- Genus Protognathinus
- Genus Sphaenognathus
- Tribe Cladognathini
- Genus Aphanognathus
- Aphanognathus laterotarsoides (Houlbert, 1915)
- Genus Capreolucanus
- Capreolucanus sicardi Didier, 1928
- Genus Cladophyllus
- Genus Eligmodontus
- Genus Gonometopus
- Genus Macrodorcas Motschulsky, 1862
- Macrodorcas adachii
- Macrodorcas amamiana
- Macrodorcas barbata
- Macrodorcas bisignata
- Macrodorcas bomansi
- Macrodorcas davidi
- Macrodorcas dierli
- Macrodorcas elsiledis
- Macrodorcas fulvonotata
- Macrodorcas humilis
- Macrodorcas intricata
- Macrodorcas kobayashii
- Macrodorcas koyamai
- Macrodorcas melliana
- Macrodorcas mochizukii
- Macrodorcas moellenkampi
- Macrodorcas montivaga
- Macrodorcas pieli
- Macrodorcas pseudaxis
- Macrodorcas recta Motschulsky, 1858
- Macrodorcas ruficrus
- Macrodorcas songiana
- Macrodorcas striatipennis Motschulsky, 1862
- Macrodorcas ursulae
- Macrodorcas vernicata
- Macrodorcas yaeyamaensis
- Genus Palaeognathus
- Genus Prismognathus Motschulsky, 1860
- Prismognathus alessandrae
- Prismognathus angularis Waterhouse, 1874
- Prismognathus branczicki
- Prismognathus cheni
- Prismognathus dauricus Motschulsky, 1860
- Prismognathus davidis
- Prismognathus delislei
- Prismognathus formosanus
- Prismognathus klapperichi
- Prismognathus lucidus
- Prismognathus morimotoi
- Prismognathus nigerrimus
- Prismognathus nigricolor
- Prismognathus piluensis
- Prismognathus platycephalus
- Prismognathus ruficephalus
- Prismognathus sinensis
- Prismognathus subnitens
- Prismognathus tokui
- Genus Prosopocoilus Westwood, 1845
- Genus Pseudorhaetus
- Genus Rhaetulus
- Genus Rhaetus
- Rhaetus westwoodi (Parry, 1862)
- Genus Tetrarthrius
- Genus Weinreichius
- Weinreichius perroti Lacroix, 1978
- Genus Aphanognathus
- Tribe Colophonini
- Genus Colophon
- Tribe Cyclommatini
- Genus Cyclommatus Parry, 1863
- Tribe Dendeziini
- Genus Dendezia
- Genus Oonotus Parry, 1864
- Genus Xiphodontus
- Tribe Dorcini
- Genus Bartolozziolucanus
- Genus Cantharolethrus
- Genus Cyclommatus
- Genus Dorcasoides
- Genus Dorcus MacLeay, 1819
- Dorcus akahorii
- Dorcus alcides
- Dorcus alexisi
- Dorcus amamianus
- Dorcus antaeus
- Dorcus anteus
- Dorcus arfakianus
- Dorcus binodulosus
- Dorcus brevis
- Dorcus bucephalus
- Dorcus carinulatus
- Dorcus consentaneus
- Dorcus curvidens
- Dorcus cylindricus
- Dorcus davidis
- Dorcus emikoae
- Dorcus eurycephalus
- Dorcus formosanus
- Dorcus gracilicornis
- Dorcus garndis
- Dorcus hopei
- Dorcus hyperion
- Dorcus japonicus
- Dorcus kyanrauensis
- Dorcus metacostatus
- Dorcus miwai
- Dorcus musimon
- Dorcus parallelus
- Dorcus parallelipipedus - lesser stag beetle. Linnaeus, 1758
- Dorcus parryi
- Dorcus parvulus
- Dorcus peyronis
- Dorcus prochazkai
- Dorcus ratiocinativus
- Dorcus rectus
- Dorcus reichei
- Dorcus rudis
- Dorcus sawaii
- Dorcus schenklingi
- Dorcus sewertzowi
- Dorcus suturalis
- Dorcus taiwanicus
- Dorcus tanakai
- Dorcus taurus
- Dorcus tenuecostatus
- Dorcus titanus
- Dorcus tormosanus
- Dorcus ursulus
- Dorcus vavrai
- Dorcus velutinus
- Dorcus vicinus
- Dorcus yamadai
- Genus Homoderus
- Genus Leptinopterus
- Genus Prosopocoilus
- Genus Rhaetulus
- Genus Serrognathus
- Genus Pycnosiphorus
- Genus Sclerostomus
- Tribe Figulini
- Genus Cardanus
- Genus Dinonigidius
- Genus Figulus
- Genus Ganelius
- Genus Nigidionus
- Genus Nigidius
- Genus Novonigidius
- Genus Penichrolucanus
- Tribe Lissapterini
- Genus Bomansius
- Genus Dorculus
- Genus Geodorcus
- Genus Hoplogonus
- Genus Lissapterus
- Genus Lissotes
- Genus Paralissotes
- Genus Pseudodorcus
- Tribe Lucanini
- Genus Hexarthrius Hope, 1842
- Hexarthrius aduncusJordan & Rothschild, 1894
- Hexarthrius andreasi Schenk, 2003
- Hexarthrius bowringii Parry, 1862
- Hexarthrius buquettii (Hope, 1843)
- Hexarthrius davisoniWaterhouse, 1888
- Hexarthrius forsteri (Hope, 1840)
- Hexarthrius howdeni De Lisle, 1972
- Hexarthrius kirchneri Schenk, 2003
- Hexarthrius mandibularis Deyrolle, 1881
- Hexarthrius melchioritis Séguy, 1954
- Hexarthrius mniszechi(Thomson, 1857)
- Hexarthrius nigritus Lacroix, 1990
- Hexarthrius parryi Hope, 1842
- Hexarthrius rhinoceros(Olivier, 1789)
- Hexarthrius vitalisi Didier, 1925
- Genus Lucanus
- Genus Pseudolucanus
- Pseudolucanus barbarossa (Fabricius, 1801)
- Genus Hexarthrius Hope, 1842
- Tribe Odontolabini
- Tribe Platycerini
- Genus Platyceroides Benesh, 1946
- Genus Platyceropsis Benesh, 1946
- Platyceropsis keeni (Casey, 1895)
- Genus Platycerus Geoffroy, 1762
- Tribe Ryssonotini
- Genus Cacostomus
- Cacostomus floralis (Lea, 1914)
- Cacostomus squamosus Newman, 1840
- Cacostomus subvittata (Moore, 1994)
- Genus Ryssonotus
- Genus Cacostomus
Subfamily Syndesinae[edit]
Subfamily Syndesinae
- Tribe Ceruchini
- Genus Ceruchus MacLeay, 1819
- Ceruchus piceus (Weber, 1801)
- Ceruchus punctatus LeConte, 1869
- Ceruchus striatus LeConte, 1859
- Ceruchus yangi Huang, Imura & Chen, 2011[2]
- Genus Ceruchus MacLeay, 1819
- Tribe Sinodendronini
- Genus Sinodendron Hellwig in Schneider, 1792
- Sinodendron cylindricum (Linnaeus, 1758)
- Sinodendron rugosum Mannerheim, 1843
- Genus Sinodendron Hellwig in Schneider, 1792
- Tribe Syndesini
- Genus Psilodon Perty, 1830
- Psilodon aequinoctiale (Buquet, 1840)
- Psilodon gilberti Boucher, 1993
- Psilodon schuberti Perty, 1830
- Psilodon seguyi (Didier, 1929)
- Psilodon xerophilicum Martinez & Reyes-Castillo, 1985
- Genus Syndesus MacLeay, 1819
- †Syndesus ambericus Woodruff, 2009 (Miocene?; Dominican amber)[3]
- Syndesus cancellatus Montrouzier, 1860
- Syndesus cornutus (Fabricius, 1801)
- Syndesus macleayi Boileau, 1905
- Genus Psilodon Perty, 1830
References[edit]
- ^ Andrew B. T. Smith (2006). "A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification" (PDF). The Coleopterists Bulletin. 60: 144–204. doi:10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2. S2CID 82432509.
- ^ Hao Huang, Yuki Imura & Chang-Chin Chen (2011). "Notes on the genus Ceruchus MacLeay from China, with the description of a new species (Coleoptera: Scarabaeoidea: Lucanidae)" (PDF Excerpt). Zootaxa. 2906: 1–42. doi:10.11646/zootaxa.2906.1.1.
- ^ Woodruff, R.E. (2009). "A new fossil species of stag beetle from Dominican Republic amber, with Australasian connections (Coleoptera: Lucanidae)". Insecta Mundi. 0098: 1–10.