Temporal range: Early Pleistocene–Late Pleistocene
|Skull from Atapuerca|
The range of this bear has been found to encompass both Europe and Asia, demonstrating the ability of the species to adapt to many Pleistocene environments.
U. deningeri is a descendant of U. savini and an ancestor of U. spelaeus.
Ursus deningeri has a combination of primitive and derived characters that distinguishes it from all other Pleistocene bears. Its mandible is slender like that of living brown bears and Ursus etruscus. It also has derived characters of cave bears (Ursus spelaeus) and is considered to be the descendant of Ursus savini and very close to the common ancestor of brown bears.
Sites and specimen ages:
- Nalaikha, Mongolia: ~1.8 Mya to 800,000 years ago
- West Runton Freshwater Bed, Cromer Forest Bed Formation, Norfolk, England: ~800,000–100,000 years ago
- Cueva del Agua, Granada, Spain: ~800,000–100,000 years ago
- Venosa bed excavations 1974–1976, Basilicata, Italy: ~800,000–100,000 years ago
- Emirkaya-2, Central Anatolia, Turkey: ~800,000–100,000 years ago
- Sima de los Huesos, Atapuerca, Spain: >300,000 years ago
- Darband Cave, Alborz, Caspian, Iran: ~300,000–200,000 years ago
Biglari, F., V. Jahani (2011). "The Pleistocene Human Settlement in Gilan, Southwest Caspian Sea: Recent Research.". Eurasian Prehistory 8 (1–8 (1–2): 3–28.
- Nuria García and Juan Luis Arsuaga, Department of Paleontology, Ciudad University, 2001[full citation needed]
- Publication in PNAS.
- Dabney & al. 2013. PNAS. "Complete mitochondrial genome sequence of a Middle Pleistocene cave bear reconstructed from ultrashort DNA fragments" doi: 10.1073/pnas.1314445110 PNAS
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