The station began broadcasting in 1965 as the first FM radio station in Lenawee County, and has been owned by the Koehn family (d/b/a Lenawee Broadcasting Company) since its inception. Over the years the station's music has evolved from a mix of MOR and country music to exclusively adult contemporary with hits from the mainstream AC and Hot AC formats, with oldies played on Fridays during "Friday Flashbacks." Like many other AC stations, WLEN plays continuous Christmas music between Thanksgiving and Christmas, taking a break on Fridays for the station's regular "Friday Flashbacks" programming of oldies from the '50s through the '80s.
WLEN features NBC radio news on the hour. All music programming is live and local with the exception of two weekend specialty shows: Magic of the 80's on Saturday nights and the weekend version of the John Tesh Radio Show on Sunday nights. The station also features a popular Tejano music show, "Radio Picoso", on Sunday afternoons, hosted by David Atkinson.
Other WLEN features include national and local newscasts, "Classified" (a Tradio-style feature), "Partyline - Now!" (a Saturday-morning phone-in talk show), and weekly specialty features focusing on neighboring communities such as Tecumseh, Onsted, Blissfield, Hudson, the Irish Hills area, and Deerfield.