2014 in paleontology

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Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2014.

Plants[edit]

Newly described angiosperms[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aerofructus[2]

Gen. et sp. nov

Valid

Herrera & Manchester in Herrera et al.

Paleocene

Cerrejón Formation

 Colombia

A member of Malvaceae described on the basis of fossil fruit. Genus includes new species A. dillhoffii.

Ambiplatanus[3]

Gen. et sp. nov

Valid

Mindell, Karafit & Stockey

Late Cretaceous (Coniacian)

 Canada

A member of Platanaceae. The type species is Ambiplatanus washingtonensis.

Anemocardium[2]

Gen. et sp. nov

Valid

Herrera & Manchester in Herrera et al.

Paleocene

Bogotá Formation

 Colombia

A fossil fruit. Genus includes new species A. margaritae.

Annona eocenica[4]

Sp. nov

Valid

Prasad, Singh & Singh

Early Eocene

Cambay Shale Formation

 India

A member of Annonaceae, a species of Annona.

Annona vastanensis[4]

Sp. nov

Valid

Prasad, Singh & Singh

Early Eocene

Cambay Shale Formation

 India

A member of Annonaceae, a species of Annona.

Antarctoxylon mixai[5]

Sp. nov

Valid

Sakala in Sakala & Vodrážka

Late Cretaceous (Coniacian)

Hidden Lake Formation

Antarctica (James Ross Island)

An angiosperm of uncertain phylogenetic placement, a species of Antarctoxylon.

Aquaephyllum[6]

Gen. et sp. nov

Valid

Gallego et al.

Late Cretaceous (Maastrichtian)

La Colonia Formation

 Argentina

A member of Araceae. The type species is Aquaephyllum auriculatum.

Bombacoxylon pondaungense[7]

Sp. nov

Valid

Licht, Boura & De Franceschi in Licht et al.

Eocene

Pondaung Formation

 Myanmar

A member of Malvaceae, a species of Bombacoxylon.

Bridelia makumensis[8]

Sp. nov

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A member of Phyllanthaceae, a species of Bridelia.

Caesalpinia veraechinataformis[9]

Sp. nov

Valid

Bernardes-de-Oliveira et al.

Eocene-Oligocene

Entre-Córregos Formation

 Brazil

A species of Caesalpinia.

Cascadiacarpa exilis[10]

Sp. nov

Valid

Mindell, Stockey & Beard

Eocene

 Canada

A member of Fagaceae, a species of Cascadiacarpa.

Castanopsis presclerophylla[11]

Sp. nov

Valid

Wu et al.

Late Pliocene

 China

A member of Fagaceae, a species of Castanopsis.

Chachlovia[12]

Gen. et sp. et comb. nov

Valid

Alekseev & Herman in Alekseev, Herman & Shchepetov

Late Cretaceous

 Russia

An angiosperm of uncertain phylogenetic placement. The type species is Chachlovia kiyensis Alekseev sp. nov.; genus also contains Chachlovia dombeyopsoida Herman comb. nov.

Cinnamomum palaeobejolghota[13]

Sp. nov

Valid

Khan & Bera

Middle to late Miocene

Dafla Formation

 India

A member of Lauraceae, a species of Cinnamomum.

Cocos binoriensis[14]

Sp. nov

Valid

Srivastava & Srivastava

Late Cretaceous or Paleocene (Maastrichtian-Danian)

Deccan Intertrappean Beds

 India

A member of Arecaceae, a relative of the coconut tree.

Cratosmilax[15]

Gen. et sp. nov

Valid

De Lima et al.

Early Cretaceous (Aptian-Albian)

Crato Formation

 Brazil

A member of Smilacaceae. The type species is Cratosmilax jacksoni.

Dicotylophyllum hansonium[16]

Sp. nov

Valid

Peppe & Hickey

Paleocene

Fort Union Formation

 United States

An angiosperm of uncertain phylogenetic placement, a species of Dicotylophyllum.

Dicotylophyllum horsecreekium[16]

Sp. nov

Valid

Peppe & Hickey

Paleocene

Fort Union Formation

 United States

An angiosperm of uncertain phylogenetic placement, a species of Dicotylophyllum.

Ficoxylon mogaungense[7]

Sp. nov

Valid

Licht, Boura & De Franceschi in Licht et al.

Eocene

Pondaung Formation

 Myanmar

A member of Moraceae, a species of Ficoxylon.

Hexagyne philippiana[17]

Gen. et sp. nov

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A member of Piperales, probably related to Aristolochiaceae. The type species is Hexagyne philippiana.

Hickeycarpum[2]

Gen. et sp. nov

Valid

Herrera & Manchester in Herrera et al.

Paleocene

Bogotá Formation

 Colombia

A fossil samara. Genus includes new species H. peltatum.

Kajanthus[18]

Gen. et sp. nov

Valid

Mendes et al.

Early Cretaceous (late Aptian–early Albian)

Figueira da Foz Formation

 Portugal

A member of Lardizabalaceae. The type species is Kajanthus lusitanicus.

Kleinhovia bikanerensis[19]

Sp. nov

Valid

Shukla, Mehrotra & Guleria

Early Eocene

Marh Formation

 India

A species of Kleinhovia.

Liliacidites contortus[20]

Sp. nov

Valid

Mildenhall & Bannister in Conran et al.

Latest Oligocene to earliest Miocene

 New Zealand

A morphospecies described on the basis of pollen recovered from a Luzuriaga-like flower.

Lindera neobifaria[13]

Sp. nov

Valid

Khan & Bera

Pliocene

Subansiri Formation

 India

A member of Lauraceae, a species of Lindera.

Luzuriaga peterbannisteri[20]

Sp. nov

Valid

Conran, Bannister, Mildenhall & Lee in Conran et al.

Latest Oligocene to earliest Miocene

 New Zealand

A species of Luzuriaga.

Macginitiea nascens[16]

Sp. nov

Valid

Peppe & Hickey

Paleocene

Fort Union Formation

 United States

A member of Platanaceae, a species of Macginitiea.

Machaerium aiuruoquense[9]

Sp. nov

Valid

Bernardes-de-Oliveira et al.

Eocene-Oligocene

Entre-Córregos Formation

 Brazil

A species of Machaerium.

Machaerium paleogenum[9]

Sp. nov

Valid

Bernardes-de-Oliveira et al.

Eocene-Oligocene

Entre-Córregos Formation

 Brazil

A species of Machaerium.

Meliosma thriviensis[16]

Sp. nov

Valid

Peppe & Hickey

Paleocene

Fort Union Formation

 United States

A member of Sabiaceae, a species of Meliosma.

Meliosma vandaelium[16]

Sp. nov

Valid

Peppe & Hickey

Paleocene

Fort Union Formation

 United States

A member of Sabiaceae, a species of Meliosma.

Microlaurus[21]

Gen. et sp. nov

Valid

Takahashi et al.

Late Cretaceous (early Coniacian)

Ashizawa Formation

 Japan

A member of Lauraceae. The type species is Microlaurus perigynus.

Paliurus microcarpa[22]

Sp. nov

Valid

Li in Li et al.

Middle or late Miocene

Shengxian Formation

 China

A member of Rhamnaceae, a species of Paliurus.

Palmoxylon romeroi[23]

Sp. nov

Valid

Franco, Brea & Herbst

Late Miocene

Chiquimil Formation

 Argentina

A palm tree, a species of Palmoxylon.

Persea mioparviflora[13]

Sp. nov

Valid

Khan & Bera

Pliocene

Subansiri Formation

 India

A member of Lauraceae. Described by Khan & Bera (2014) as a species of Persea, considered by the authors to be most similar to the species Persea parviflora Spreng.[13]

Persea preglaucescens[13]

Sp. nov

Valid

Khan & Bera

Pliocene

Subansiri Formation

 India

A member of Lauraceae. Described by Khan & Bera (2014) as a species of Persea,[13] though the authors consider it most similar to the species Persea glaucescens (Nees) D.G. Long, assigned by other authors to the genus Machilus.

Platimeliphyllum reznikoviorum[24]

Sp. nov

Valid

Maslova in Maslova et al.

Paleogene

Ashut Formation

 Kazakhstan

An angiosperm of uncertain phylogenetic placement, a species of Platimeliphyllum.

Quercus tenuipilosa[25]

Sp. nov

Valid

Hu & Zhou in Hu et al.

Late Pliocene

Ciying Formation

 China

An oak.

Quercus yangyiensis[26]

Sp. nov

Valid

He, Li & Sun in He et al.

Late Pliocene

Yangyi Formation

 China

An oak.

Ripogonum americanum[27]

Sp. nov

Valid

Carpenter et al.

Early Eocene

 Argentina

A member of Liliales, a species of Ripogonum.

Ruprechtioxylon[28]

Gen. et sp. nov

Valid

Cevallos-Ferriz, Martínez-Cabrera & Calvillo-Canadell

Early Miocene

El Cien Formation

 Mexico

A member of Polygonaceae described on the basis of fossil wood. Genus includes new species R. multiseptatus.

Sabalites dindoriensis[29]

Sp. nov

Valid

Srivastava, Srivastava & Dilcher

Late Cretaceous (Maastrichtian) to early Paleocene (Danian)

Deccan Intertrappean Beds

 India

A palm tree belonging to the subfamily Coryphoideae, a species of Sabalites.

Sapindoxylon burmense[7]

Sp. nov

Valid

Licht, Boura & De Franceschi in Licht et al.

Eocene

Pondaung Formation

 Myanmar

A member of Sapindaceae, a species of Sapindoxylon.

Schimoxylon benderi[7]

Sp. nov

Valid

Licht, Boura & De Franceschi in Licht et al.

Eocene

Pondaung Formation

 Myanmar

A member of Theaceae, a species of Schimoxylon.

Shoreoxylon panganense[7]

Sp. nov

Valid

Licht, Boura & De Franceschi in Licht et al.

Eocene

Pondaung Formation

 Myanmar

A member of Dipterocarpaceae, a species of Shoreoxylon.

Soninia[12]

Gen. et 2 sp. nov

Valid

Herman & Shczepetov in Alekseev, Herman & Shchepetov

Late Cretaceous

 Russia

An angiosperm of uncertain phylogenetic placement. The type species is S. integerrima Herman & Shczepetov; genus also contains S. asiatica Alekseev.

Ternstromites paucimissouriensis[16]

Sp. nov

Valid

Peppe & Hickey

Paleocene

Fort Union Formation

 United States

A member of Theaceae, a species of Ternstromites.

Ticodendron palaios[30]

Sp. nov

Valid

Chambers & Poinar

Eocene or Miocene

 Dominican Republic

A species of Ticodendron known from Dominican amber.

Ulmoidicarpum[2]

Gen. et sp. nov

Valid

Herrera & Manchester in Herrera et al.

Paleocene

Cerrejón Formation

 Colombia

A member of Ulmaceae described on the basis of fossil fruits. Genus includes new species U. tupperi.

Other seed plants[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Agathis zamunerae[31]

Sp. nov

Valid

Wilf et al.

Eocene

 Argentina

A member of Araucariaceae, a species of Agathis.

Araucarites ochotensis[32]

Sp. nov

Valid

Golovneva & Shczepetov in Shczepetov & Golovneva

Late Cretaceous

 Russia

A member of Araucariaceae, a species of Araucarites.

Athrotaxites yumenensis[33]

Sp. nov

Valid

Dong et al.

Early Cretaceous

Zhonggou Formation

 China

A member of Cupressaceae, a species of Athrotaxites.

Bicatia[34]

Gen. et 3 sp. nov

Valid

Friis, Pedersen & Crane

Early Cretaceous (late Aptian to middle Albian)

 Portugal
 United States

A Welwitschia-like seed plant. The type species is Bicatia costata; genus also contains Bicatia juncalensis and Bicatia rugosa.

Cordaites barthelii[35]

Sp. nov

Valid

Šimůnek

Carboniferous (Stephanian)

 Czech Republic

A species of Cordaites.

Cosmosperma[36]

Gen. et sp. nov

Valid

Wang et al.

Late Devonian (Famennian)

Wutong Formation

 China

A member of Lagenospermopsida of uncertain phylogenetic placement. The type species is Cosmosperma polyloba.

Cycas fushunensis[37]

Sp. nov

Valid

Su, Quan & Liu

Eocene (Lutetian)

Jijuntun Formation

 China

A cycad, a species of Cycas.

Elatides zhoui[38]

Sp. nov

Valid

Shi et al.

Early Cretaceous

 Mongolia

A member of Cupressaceae, a species of Elatides.

Elatocladus andhrensis[39]

Sp. nov

Valid

Chinnappa, Rajanikanth & Rao

Early Cretaceous

Gangapur Formation

 India

A member of Coniferales, a species of Elatocladus.

Flabellosemen[40]

Gen. et sp. nov

Valid

Tsymbal

Permian

 Russia

A member of Ginkgoales belonging to the family Psygmophyllaceae. The type species is Flabellosemen riparium.

Giblingodendron[41]

Gen. et 2 sp.nov

Valid

Falcon-Lang, Kurzawe & Lucas

Carboniferous (Kasimovian)[42] to Permian (Sakmarian)

Atrasado Formation[42]
Community Pit Formation

 United States

A member of Coniferopsida of uncertain phylogenetic placement, might be a member of Voltziales, Cordaitales or Dicranophyllales.[42] The type species is Giblingodendron nudifolia; genus also includes Giblingodendron aridus.

Ginkgoxylpropinquus saxonicum[43]

Sp. nov

Valid

Süss & Müller

Early Oligocene

 Germany

A member of Ginkgoales, a species of Ginkgoxylpropinquus.

Hubbardiastrobus[44]

Gen. et sp. nov

Valid

Atkinson, Rothwell & Stockey

Early Cretaceous (Valanginian)

 Canada

A member of Cupressaceae. The type species is Hubbardiastrobus cunninghamioides.

Hughmillerites vancouverensis[45]

Sp. nov

Valid

Atkinson, Rothwell & Stockey

Early Cretaceous

 Canada

A member of Cupressaceae, a species of Hughmillerites.

Macdonaldodendron[41]

Gen. et sp.nov

Valid

Falcon-Lang, Kurzawe & Lucas

Carboniferous (Kasimovian)[42] to Permian (Sakmarian)

Atrasado Formation[42]
Community Pit Formation

 United States

A member of Voltziales. The type species is Macdonaldodendron giganticus.

Manifera[46]

Gen. et sp. nov

Valid

Looy & Stevenson

Permian (Kungurian to Roadian)

Flowerpot Shale Member of the San Angelo Formation

 United States

A member of Voltziales belonging to the family Majonicaceae. The type species is Manifera talaris.

Nilssoniopteris binggouensis[47]

Sp. nov

Valid

Na et al.

Early Cretaceous

Binggou Formation

 China

A member of Bennettitales, a species of Nilssoniopteris.

Nilssoniopteris corrugata[48]

Sp. nov

Valid

Ray, Rothwell & Stockey

Early Cretaceous (Valanginian)

 Canada

A member of Bennettitales, a species of Nilssoniopteris.

Pararaucaria collinsonae[49]

Sp. nov

Valid

Steart, Spencer, Kenrick, Needham & Hilton in Steart et al.

Late Jurassic (Tithonian)

Lulworth Formation

 United Kingdom

A member of Cheirolepidiaceae, a species of Pararaucaria.

Peltaspermum morovii[50]

Sp. nov

Valid

Naugolnykh

Permian (Kazanian)

 Russia

A member of Peltaspermaceae.

Phoenicopsis (Culgoweria) ordosensis[51]

Sp. nov

Valid

Li et al.

Middle Jurassic

Yan'an Formation

 China

A member of Czekanowskiales, a species of Phoenicopsis.

Phoenicopsis (Phoenicopsis) neimengensis[52]

Sp. nov

Valid

Wang, Yang & Sun

Early Cretaceous

Huolinhe Formation

 China

A member of Czekanowskiales, a species of Phoenicopsis.

Phyllocladoxylon pooleae[53]

Sp.nov

Valid

Pujana, Santillana & Marenssi

Eocene

La Meseta Formation

Antarctica

A member of Podocarpaceae, a species of Phyllocladoxylon.

Platycladus yunnanensis[54]

Sp. nov

Valid

Wu in Wu et al.

Late Miocene

Nanlin Formation

 China

A member of Cupressaceae, a species of Platycladus.

Podocarpoxylon radiciforme[55]

Sp. nov

Valid

Süss & Müller

Early Oligocene

 Germany

A conifer, a species of Podocarpoxylon.

Protophyllocladoxylon francisiae[53]

Sp.nov

Valid

Pujana, Santillana & Marenssi

Eocene

La Meseta Formation

Antarctica

A member of Podocarpaceae, a species of Protophyllocladoxylon.

Rudixylon[56]

Gen. et sp. nov

Valid

Bomfleur et al.

Middle or early Late Triassic

Fremouw Formation

Antarctica

A member of Petriellales (an order of seed plants of uncertain phylogenetic placement). The type species is Rudixylon serbetianum.

Rugaspermum stipitatus[57]

Sp. nov

Valid

Bhowmik, Parveen & Das

Middle Triassic

 India

A possible member of Gnetales, a species of Rugaspermum.

Septomedullopitys szei[58]

Sp. nov

Valid

Wan, Yang & Wang

Late Permian (early Wuchiapingian)

Wutonggou Formation

 China

A member of Coniferopsida, a species of Septomedullopitys.

Shimakuroxylon[59]

Gen. et comb. nov

Valid

Philippe et al.

Jurassic and Cretaceous

 China
 Indonesia
 Japan

A gymnosperm described on the basis of fossil wood, possibly a member of Pinophyta. A new genus for "Dadoxylon" japonicum Shimakura (1936).

Xinjiangoxylon[60]

Gen. et sp. nov

Valid

Shi et al.

Late Permian (Changhsingian)

Guodikeng Formation

 China

A gymnosperm. The type species is Xinjiangoxylon turpanense.

Pteridophyta[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Arcellites punctatus[61]

Sp. nov

Valid

Friis, Pedersen & Marone

Early Cretaceous (late Barremian–early Aptian)

Almargem Formation

 Portugal

A member of Salviniales of uncertain phylogenetic placement, a species of Arcellites.

Ashicaulis plumites[62]

Sp. nov

Valid

Tian & Wang in Tian et al.

Middle Jurassic

Tiaojishan Formation

 China

A member of Osmundaceae, a species of Ashicaulis.

Ashicaulis wangii[63]

Sp. nov

Valid

Tian et al.

Middle Jurassic

Tiaojishan Formation

 China

A member of Osmundaceae, a species of Ashicaulis.

Chiropteris monteagnellii[64]

Sp. nov

Valid

Kustatscher, Dellantonio & Van Konijnenburg-Van Cittert

Middle Triassic (Ladinian)

 Italy

A pteridophyte of uncertain phylogenetic placement, a species of Chiropteris.

Cladophlebis ladinica[64]

Sp. nov

Valid

Kustatscher, Dellantonio & Van Konijnenburg-Van Cittert

Middle Triassic (Ladinian)

 Italy

A pteridophyte of uncertain phylogenetic placement, a species of Cladophlebis.

Elaphoglossum miocenicum[65]

Sp. nov

Valid

Lóriga et al.

Miocene

 Dominican Republic

A member of Dryopteridaceae found in Dominican amber, a species of Elaphoglossum.

Flabellariopteris[66]

Gen. et sp. nov

Valid

Sun in Sun et al.

Late Triassic

Yangcaogou Formation

 China

A member of Marsileaceae. The type species is Flabellariopteris mii.

Marattiopsis vodrazkae[67]

Sp. nov.

Valid

Kvaček

Late Cretaceous (Campanian)

Hidden Lake Formation

Antarctica (James Ross Island)

A member of Marattiaceae, a species of Marattiopsis.

Marsilea patnii[68]

Sp. nov

Valid

Patil, Kapgate & Zilpe

Late Cretaceous

Deccan Intertrappean Beds

 India

A species of Marsilea.

Phlebopteris fiemmensis[64]

Sp. nov

Valid

Kustatscher, Dellantonio & Van Konijnenburg-Van Cittert

Middle Triassic (Ladinian)

 Italy

A member of Matoniaceae, a species of Phlebopteris.

Phlebopteris hickeyi[69]

Sp. nov

Valid

Hu & Taylor

Early Cretaceous (Albian)

Jarash Formation

 Jordan

A member of Matoniaceae, a species of Phlebopteris.

Pseudobornia schweitzeri[70]

Sp. nov

Valid

Jurina & Orlova in Orlova & Jurina

Late Devonian

 Russia

A species of Pseudobornia.

Salvinia hainanensis[71]

Sp. nov

Valid

Jin & Wang in Wang, Xu & Jin

Eocene

 China

A species of Salvinia.

Tiania[72]

Gen. et comb. nov

Valid

Wang et al.

Late Permian

Xuanwei Formation

 China

A member of Guaireaceae (extinct family belonging to the order Osmundales); a new genus for "Palaeosmunda" yunnanense Tian & Chang (1995).

Zhongmingella[73]

Gen. et comb. nov

Valid

Wang et al.

Late Permian

Wangjiazhai Formation

 China

A member of Guaireaceae (extinct family belonging to the order Osmundales); a new genus for "Palaeosmunda" plenasioides Li (1983).

Other plants[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Andreiella[74]

Gen. et sp. nov

Valid

Bucur

Early Cretaceous (Barremian–?Aptian)

 Romania

A dasycladalean alga belonging to the family Triploporellaceae. Genus includes new species A. rajkae.

Baikalophyton[75]

Gen. et sp. nov

Valid

Naugolnykh & Minina

Devonian

 Russia

A terrestrial plant similar to Sciadophyton. The type species is Baikalophyton ruzhentsevii.

Belzungia barattoloi[76]

Sp. nov

Valid

Radoičić & Özgen-Erdem

Early Eocene

 Turkey

A dasycladalean alga belonging to the family Thyrsoporellaceae.

Boueina iberica[77]

Sp. nov

Valid

Schlagintweit & Wilmsen

Late Cretaceous (Cenomanian)

Altamira Formation

 Spain

A dasycladalean alga belonging to the family Udoteaceae.

Calymperites burmensis[78]

Sp. nov

Valid

Heinrichs et al.

Cretaceous (Albian or Cenomanian)

Burmese amber

 Myanmar

A moss belonging to the group Dicranales (sensu lato).

Ceratolejeunea antiqua[79]

Sp. nov

Valid

Heinrichs & Schäfer-Verwimp in Heinrichs et al.

Early to middle Miocene

 Mexico

A liverwort belonging to the family Lejeuneaceae, a species of Ceratolejeunea.

Changxingia[80]

Gen. et sp. nov

Valid

Wang et al.

Late Devonian

Wutong Formation

 China

A lycopsid. The type species is Changxingia longifolia.

Clypeina tibanai[81]

Sp. nov

Valid

Granier, Dias-Brito & Bucur

Cretaceous (AlbianCenomanian)

Ponta do Mel Formation

 Brazil

A dasycladalean alga belonging to the family Polyphysaceae.

Diplopora obliguspora[82]

Sp. nov

Valid

Senowbari-Daryan

Late Triassic (Norian-Rhaetian)

 Greece

A dasycladalean alga belonging to the family Diploporellaceae.

Gackstroemia cretacea[83]

Sp. nov

Valid

Heinrichs et al.

Early Cretaceous (late Albian)

 Myanmar

A liverwort belonging to the group Jungermanniopsida and the family Lepidolaenaceae, a species of Gackstroemia.

Lepidodendron bellii[84]

Sp. nov

Valid

Álvarez-Vázquez & Wagner

Carboniferous (middle Pennsylvanian, i.e. early Westphalian)

 Canada

A species of Lepidodendron.

Lycopodites baikalensis[85]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Middle Jurassic

Prisayan Formation

 Russia

A member of Lycopodiales, a species of Lycopodites.

Lycopodites subulifolius[85]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Prisayan Formation

 Russia

A member of Lycopodiales, a species of Lycopodites.

Marchantites huolinhensis[86]

Sp. nov

Valid

Li et al.

Early Cretaceous

Huolinhe Formation

 China

A liverwort belonging to the group Marchantiales, a species of Marchantites.

Pantiathallites[87]

Gen. et sp. nov

Valid

Banerjee & Dutta

Early Permian

 India

A metzgeriinean bryophyte. The type species is Pantiathallites gondwanensis.

Planatophyton[88]

Gen. et sp. nov

Valid

Gerrienne et al.

Early or Middle Devonian

Hujiersite Formation

 China

An euphyllophyte. The type species is Planatophyton hujiersitense.

Probolocupsis sarmeikensis[82]

Sp. nov

Valid

Senowbari-Daryan

Late Triassic (Norian-Rhaetian)

 Greece

A dasycladalean alga belonging to the family Triploporellaceae.

Probolocuspis? tenuiparia[82]

Sp. nov

Valid

Senowbari-Daryan

Late Triassic (Norian-Rhaetian)

 Greece

A dasycladalean alga belonging to the family Triploporellaceae.

Rhizomnium dentatum[89]

Sp nov

valid

Heinrichs et al, 2014

Middle Eocene

Baltic amber

 Europe

A Mniaceae moss

Russoella parthica[90]

Sp. nov

Valid

Barattolo, Taherpour Khalil Abad & Ashrafi

Late Cretaceous (Maastrichtian)

 Afghanistan

A dasycladalean alga.

Ufadendron[91]

Gen. et sp. nov

Valid

Naugolnykh

Permian (Kungurian)

Koshelevsk Formation

 Russia

A lycopsid. The type species is Ufadendron ufaensis.

Vetiplanaxis espinosus[92]

Sp. nov

Valid

Hedenäs, Heinrichs & Schmidt

Cretaceous (Albian or Cenomanian)

Burmese amber

 Myanmar

A moss belonging to the class Bryopsida and the order Hypnodendrales.

Vetiplanaxis longiacuminatus[92]

Sp. nov

Valid

Hedenäs, Heinrichs & Schmidt

Cretaceous (Albian or Cenomanian)

Burmese amber

 Myanmar

A moss belonging to the class Bryopsida and the order Hypnodendrales.

Vetiplanaxis oblongus[92]

Sp. nov

Valid

Hedenäs, Heinrichs & Schmidt

Cretaceous (Albian or Cenomanian)

Burmese amber

 Myanmar

A moss belonging to the class Bryopsida and the order Hypnodendrales.

Cnidarians[edit]

Newly described cnidarians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Acropora macrocalyx[93]

Sp. nov

Valid[94]

Wallace & Bosellini

Early Miocene (Burdigalian)

 Italy

A stony coral, a species of Acropora.

Acropora piedmontensis[93]

Sp. nov

Valid[94]

Wallace & Bosellini

Early Miocene (Burdigalian)

 Italy

A stony coral, a species of Acropora.

Acropora salentina[93]

Sp. nov

Valid[94]

Wallace & Bosellini

Oligocene (Chattian)

 Italy

A stony coral, a species of Acropora.

Acropora slovenica[93]

Sp. nov

Valid[94]

Wallace & Bosellini

Oligocene (Rupelian)

Gornji Grad beds

 Slovenia

A stony coral, a species of Acropora.

Arctistrotion[95]

Gen. et 2 sp. et comb. nov

Valid

Fedorowski & Stevens

Carboniferous (Bashkirian)

 United States

A colonial coral belonging to the subclass Rugosa and the family Lithostrotionidae. The type species is Arctistrotion variabilis; genus also contains "Lithostrotionella" wahooensis Armstrong (1972) and Arctistrotion simplex.

Axophyllum cozari[96]

Sp. nov

Valid

Rodríguez & Somerville

Carboniferous (Viséan)

 Spain

A rugose coral belonging to the group Stauriida and the family Axophyllidae.

Axophyllum julianaense[96]

Sp. nov

Valid

Rodríguez & Somerville

Carboniferous (Serpukhovian)

 Spain

A rugose coral belonging to the group Stauriida and the family Axophyllidae.

Axophyllum spinosum[96]

Sp. nov

Valid

Rodríguez & Somerville

Carboniferous (Serpukhovian)

 Spain

A rugose coral belonging to the group Stauriida and the family Axophyllidae.

Axophyllum spiralum[96]

Sp. nov

Valid

Rodríguez & Somerville

Carboniferous (Viséan)

 Spain

A rugose coral belonging to the group Stauriida and the family Axophyllidae.

Cambroctoconus kyrgyzstanicus[97]

Sp. nov

Valid

Peel in Geyer et al.

Cambrian (Amgan)

 Kyrgyzstan

A stem-cnidarian, a species of Cambroctoconus.

Haootia[98]

Gen. et sp. nov

Valid

Liu et al.

Late Ediacaran

Fermeuse Formation

 Canada

A probable cnidarian, possibly related to Medusozoa. The type species is Haootia quadriformis.

Morenaphyllum[96]

Gen. et 2 sp. et comb. nov

Valid

Rodríguez & Somerville

Carboniferous

 Kazakhstan
 Spain

A rugose coral belonging to the group Stauriida and the family Axophyllidae. The type species is M. antolinense; genus also includes new species M. boyerense, as well as "Carcinophyllum lonsdaleiforme" pauciseptata Bykova (1966) (raised to the rank of a distinct species).

Paraheritschioides compositus[95]

Sp. nov

Valid

Fedorowski & Stevens

Carboniferous (Bashkirian)

Wahoo Limestone

 United States

A colonial coral belonging to the subclass Rugosa and the family Kleopatrinidae, a species of Paraheritschioides.

Paraheritschioides katvalae[95]

Sp. nov

Valid

Fedorowski & Stevens

Carboniferous (Moscovian)

Saginaw Bay Formation

 United States

A colonial coral belonging to the subclass Rugosa and the family Kleopatrinidae, a species of Paraheritschioides.

Pareynia viacrucense[96]

Sp. nov

Valid

Rodríguez & Somerville

Carboniferous (Serpukhovian)

 Spain

A rugose coral belonging to the group Stauriida and the family Axophyllidae.

Sinobryon[99]

Gen. et sp. nov

Valid

Baliński, Sun & Dzik

Ordovician (early Floian)

Fenxiang Formation

 China

A hydrozoan, a member of Hydroidolina. The type species is Sinobryon elongatum.

Thamnopora itoae[100]

Sp. nov

Valid

Niko, Ibaraki & Tazawa

Devonian

 Japan

A tabulate coral belonging to the group Favositida and the family Pachyporidae.

Thurispina[101]

Gen. et sp. nov

Valid

Weyer

Devonian (late Famennian)

 Germany

A member of Rugosa. The type species is Thurispina jogi.

Arthropods[edit]

Molluscs[edit]

Echinoderms[edit]

Conodonts[edit]

Newly described conodonts[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Acuminatella binodosa[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella constricta[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella curvata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella denticulata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella? extensa[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella longicarinata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella? prima[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella sagittale[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Acuminatella sinuosa[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Belodina longxianensis minor[103]

Subsp. nov

Valid

Cai, Yuan & Wu

Late Ordovician

Lianglitag Formation

 China

A subspecies of Belodina longxianensis.

Carnepigondolella anitae[102]

Sp. nov

Valid

Orchard

Late Triassic

Ludington Formation

 Canada

A member of Gondolellidae.

Carnepigondolella gibsoni[102]

Sp. nov

Valid

Orchard

Late Triassic

Ludington Formation

 Canada

A member of Gondolellidae.

Carnepigondolella milanae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Carnepigondolella postsamueli[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Carnepigondolella spenceri[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Furnishina holmi[104]

Sp. nov

Valid

Bagnoli & Stouge

Cambrian (late Furongian)

Alum Shale Formation

 Sweden

A species of Furnishina.

Gobduntaulepis[105]

Nom. nov

Valid

Doweld

Early Ordovician

 Uzbekistan

A replacement name for Acantholepis Pianovskij in Pianovskij, Pianovskaja & Aleksandrova, 1989 (preoccupied).

Hindeodus microdentatus[106]

Sp. nov

Valid

Wu et al.

Permian–Triassic boundary

 China

A species of Hindeodus.

Icriodella rhodesi[107]

Sp. nov

Valid[108]

Bergström & Ferretti

Late Ordovician

Keisley Limestone

 United Kingdom

A species of Icriodella.

Kraussodontus ludingtonensis[102]

Sp. nov

Valid

Orchard

Late Triassic

Ludington Formation
Pardonet Formation

 Canada

A member of Gondolellidae.

Kraussodontus margaretae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Kraussodontus roberti[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Kraussodontus rosiae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Kraussodontus urbanae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Kraussodontus vancouverense[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Kraussodontus wendae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Metapolygnathus dylani[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Neogondolella prediscreta[106]

Sp. nov

Valid

Wu et al.

Earliest Triassic

 China

A species of Neogondolella.

Norigondolella norica[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parafurnishius[109]

Gen. et sp. nov

Valid

Yang et al.

Early Triassic (Induan)

Feixianguan Formation

 China

A member of Ellisoniidae. The type species is Parafurnishius xuanhanensis.

Parapetella beattyi[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella broatchae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella clareae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella columbiense[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella cordillerense[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella destinae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella elegantula[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella hillarae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella irwini[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella johnpauli[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella lanei[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella posterolata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella prominens angulare[102]

Subsp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella prominens circulare[102]

Subsp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella prominens prominens[102]

Subsp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella pumilio[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella riteri[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella rubae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Parapetella willifordi[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Polygnathus alkhovikovae[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus aragonensis[111]

Sp. nov

Valid

Martínez-Pérez & Valenzuela-Ríos

Early Devonian

Basibé Formation

 Spain

A polygnathid, a species of Polygnathus.

Polygnathus arthuri[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus bardashevi[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus carlsi[111]

Sp. nov

Valid

Martínez-Pérez & Valenzuela-Ríos

Early Devonian

Basibé Formation

 Spain

A polygnathid, a species of Polygnathus.

Polygnathus communis longanensis[112]

Subsp. nov

Valid

Qie et al.

Carboniferous (latest Tournaisian)

Long’an Formation

 China

A polygnathid, a subspecies of Polygnathus communis.

Polygnathus dujieensis[112]

Sp. nov

Valid

Qie et al.

Carboniferous (latest Tournaisian)

Long’an Formation

 China

A polygnathid, a species of Polygnathus.

Polygnathus ivanowskyii[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus karsteni[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus lezhoevi[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus michaelmurphyi[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus ramoni[111]

Sp. nov

Valid

Martínez-Pérez & Valenzuela-Ríos

Early Devonian

Basibé Formation

 Spain

A polygnathid, a species of Polygnathus.

Polygnathus settedabanicus[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Polygnathus yakutensis[110]

Sp. nov

Valid

Baranov, Slavík & Blodgett

Devonian (early Emsian)

 Russia

A polygnathid, a species of Polygnathus.

Primatella bifida[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella circulare[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella elongata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella mclearni[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella oblonga[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella ovale[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella posteroglobosa[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella rectangulare[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella rhomboidale[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella rotunda[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella stanleyi[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella subquadrata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella triangulare[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Primatella vanlierae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella deflecta[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella karenae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella kathleenae[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella mcrobertsi[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella pardoneti[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella posteroexpansa[102]

Sp. nov

Valid

Orchard

Late Triassic

Ludington Formation

 Canada

A member of Gondolellidae.

Quadralella postlobata[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella praecommunisti curvata[102]

Subsp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella praecommunisti ornata[102]

Subsp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella roysi[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella sigmoidale[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Quadralella willistonense[102]

Sp. nov

Valid

Orchard

Late Triassic

Pardonet Formation

 Canada

A member of Gondolellidae.

Siphonodella nandongensis[113]

Sp. nov

Valid

Li et al.

Early Carboniferous

Baping Formation

 China

A species of Siphonodella.

Westergaardodina asinina[104]

Sp. nov

Valid

Bagnoli & Stouge

Cambrian (late Furongian)

Alum Shale Formation

 Sweden

A species of Westergaardodina.

Fishes[edit]

Amphibians[edit]

Basalmost tetrapods[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Weberepeton[114]

Gen. et sp. nov

Valid

Clément & Lebedev

Late Devonian (late Frasnian)

Probably Smota-Lovat’ Formation

 Russia

A member of Elginerpetontidae. The type species is Weberepeton sondalensis.

Temnospondyls[edit]

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Antarctosuchus[116]

Gen. et sp. nov

Valid

Sidor, Steyer & Hammer

Middle Triassic

Fremouw Formation

Antarctica

A capitosauroid. The type species is Antarctosuchus polyodon.

Leptorophus raischi[117]

Sp. nov

Valid

Schoch

Early Permian

Saar–Nahe Basin

 Germany

A branchiosaurid, a species of Leptorophus.

Megalophthalma[118]

Gen. et sp. nov

Valid[119]

Schoch, Milner & Witzmann

Middle Triassic (late Ladinian)

Erfurt Formation

 Germany

A plagiosaurid. The type species is Megalophthalma ockerti.

Nanobamus[120]

Gen. et sp. nov

Valid

Schoch & Milner

Early Permian

Putnam Formation

 United States

An amphibamid. The type species is Nanobamus macrorhinus.

Chroniosuchians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Dromotectum abditum[121]

Sp. nov

Valid

Shishkin, Novikov & Fortuny

Early Triassic

 Russia

A bystrowianid chroniosuchian, a species of Dromotectum.

Dromotectum largum[122]

Sp. nov

Valid

Liu et al.

Late Permian

Shangshihezi Formation

 China

A bystrowianid chroniosuchian, a species of Dromotectum.

Jiyuanitectum[122]

Gen. et sp. nov

Valid

Liu et al.

Late Permian

Shangshihezi Formation

 China

A bystrowianid chroniosuchian. The type species is Jiyuanitectum flatum.

Vyushkoviana[121]

Gen. et sp. nov

Valid

Shishkin, Novikov & Fortuny

Early Triassic

 Russia

A bystrowianid chroniosuchian. The type species is Vyushkoviana operta.

Lissamphibians[edit]

Research[edit]

  • The humerus bone of a large calyptocephalellid anuran, apparently one of the largest fossil anurans known to date, is described by Otero et al. (2014) from the Eocene of Chile.[123]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Eopelobates deani[124]

Sp. nov

Valid

Roček et al.

Early middle Eocene

Green River Formation

 United States

A relative of the European spadefoot toads, a species of Eopelobates.

Ichthyopterygians[edit]

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Janusaurus[126]

Gen. et sp. nov

Valid

Roberts et al.

Late Jurassic

Agardhfjellet Formation

 Norway
 Russia

An ophthalmosaurid ichthyosaur. The type species is Janusaurus lundi.

Leninia[127]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (early Aptian)

 Russia

An ophthalmosaurine ophthalmosaurid ichthyosaur. The type species is Leninia stellans.

Sisteronia[128]

Gen. et sp. nov

Valid

Fischer et al.

Cretaceous (early Albian to early Cenomanian)

Cambridge Greensand
Gault Formation
Marnes Bleues Formation

 France
 United Kingdom

A platypterygiine ophthalmosaurid ichthyosaur. The type species is Sisteronia seeleyi.

Undorosaurus trautscholdi[129]

Sp. nov

Valid

Arkhangelsky & Zverkov

Late Jurassic

 Russia

An ophthalmosaurine ophthalmosaurid ichthyosaur, a species of Undorosaurus.

Sauropterygians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aristonectes quiriquinensis[130]

Sp. nov

Valid

Otero et al.

Late Cretaceous (late Maastrichtian)

Quiriquina Formation

 Chile

A plesiosaur, a species of Aristonectes.

Cymatosaurus erikae[131]

Sp. nov

Valid

Maisch

Middle Triassic (earliest Anisian)

Röt Formation

 Germany

A species of Cymatosaurus.

Majiashanosaurus[132]

Gen. et sp. nov

Valid

Jiang et al.

Early Triassic (Olenekian)

Nanlinghu Formation

 China

A basal member of Eosauropterygia. The type species is Majiashanosaurus discocoracoidis.

Nothosaurus zhangi[133]

Sp. nov

Valid

Liu et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A nothosaur, a species of Nothosaurus.

Odoiporosaurus[134]

Gen. et sp. nov

Valid

Renesto, Binelli & Hagdorn

Middle Triassic (Anisian)

Besano Formation

 Italy

A pachypleurosaur. The type species is Odoiporosaurus teruzzii.

Pararcus[135]

Gen. et sp. nov

Valid

Klein & Scheyer

Middle Triassic (early Anisian)

Vossenveld Formation

 Netherlands

A placodont. The type species is Pararcus diepenbroeki.

Pliosaurus patagonicus[136]

Sp. nov

Valid

Gasparini & O'Gorman

Late Jurassic (middle Tithonian)

Vaca Muerta Formation

 Argentina

A species of Pliosaurus.

Lepidosaurs[edit]

Newly named rhynchocephalians[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Kawasphenodon peligrensis[137]

Sp. nov

Valid

Apesteguia, Gómez & Rougier

Palaeocene

Salamanca Formation

 Argentina

An opisthodontian sphenodontid, a species of Kawasphenodon.

Priosphenodon minimus[138]

Sp. nov

Valid

Apesteguia & Carballido

Early Cretaceous

 Argentina

An eilenodontine sphenodontid, a species of Priosphenodon.

Newly named lizards[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Blanus mendezi[139]

Sp. nov

Valid

Bolet et al.

Miocene (11.6 Mya)

Vallès-Penedès Basin

 Spain

An amphisbaenian, a species of Blanus.

Blanus thomaskelleri[140]

Sp. nov

Valid[141]

Čerňanský, Rage & Klembara

Early Miocene

 Germany

An amphisbaenian, a species of Blanus.

Calanguban[142]

Gen. et sp. nov

Valid[143]

Simões, Caldwell & Kellner

Early Cretaceous

Crato Formation

 Brazil

A scleroglossan lizard with scincomorph affinities. The type species is Calanguban alamoi.

Dakotaseps[144]

Nom. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade;[145] a replacement name for Dakotasaurus Nydam (2013) (preoccupied).

Funiusaurus[146]

Gen. et sp. nov

Valid

Xu et al.

Late Cretaceous

Qiupa Formation

 China

A member of Polyglyphanodontidae. The type species is Funiusaurus luanchuanensis.

Maioricalacerta[147]

Gen. et sp. nov

Valid

Bailon et al.

Early Pliocene (Zanclean)

 Spain

A member of Lacertidae. The type species is Maioricalacerta rafelinensis.

Pyrenasaurus[148]

Gen. et sp. nov

Valid

Bolet & Augé

Late Eocene

 France
 Spain

A member of Scincoidea, possibly a skink. The type species is Pyrenasaurus evansae.

Newly named snakes[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Adinophis[149]

Gen. et sp. nov

Valid

Pritchard et al.

Late Cretaceous (Maastrichtian)

Maevarano Formation

 Madagascar

A member of Madtsoiidae. The type species is Adinophis fisaka.

Indophis fanambinana[149]

Sp. nov

Valid

Pritchard et al.

Late Cretaceous (Maastrichtian)

Maevarano Formation

 Madagascar

A member of Nigerophiidae, a species of Indophis.

Rukwanyoka[150]

Gen. et sp. nov

Valid

McCartney et al.

Oligocene

Nsungwe Formation

 Tanzania

The most ancient booid snake and boa from Africa. The type species is Rukwanyoka holmani.

Seismophis[151]

Gen. et sp. nov

Valid

Hsiou et al.

Late Cretaceous (Cenomanian)

Alcântara Formation

 Brazil

A snake of uncertain phylogenetic placement, possibly a relative of Najash rionegrina. The type species is Seismophis septentrionalis.

Turtles[edit]

Newly named turtles[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Alienochelys[152]

Gen. et sp. nov

Valid

de Lapparent de Broin et al.

Late Cretaceous (Maastrichtian)

Oulad Abdoun Basin

 Morocco

A sea turtle, a member of Dermochelyoidae. The type species is Alienochelys selloumi.

Allaeochelys libyca[153]

Sp. nov

Valid

Havlik, Joyce & Böhme

Miocene (Langhian)

 Libya

A relative of the pig-nosed turtle, a species of the (possibly paraphyletic) genus Allaeochelys.

Ashleychelys[154]

Gen. et sp. nov

Valid

Weems & Sanders

Oligocene

 United States

A pancheloniid sea turtle. The type species is Ashleychelys palmeri.

Atolchelys[155]

Gen. et sp. nov

Valid

Romano et al.

Early Cretaceous (Barremian)

Morro do Chaves Formation

 Brazil

A bothremydid pleurodiran. The type species is Atolchelys lepida.

Brodiechelys royoi[156]

Sp. nov

Valid

Pérez-García, Gasulla & Ortega

Early Aptian

Arcillas de Morella Formation

 Spain

A xinjiangchelyid, a species of Brodiechelys.

Cheirogaster bacharidisi[157]

Sp. nov

Valid

Vlachos, Tsoukala & Corsini

Pliocene

Gonia Formation

 Greece

A tortoise, originally described as a species belonging to the genus Cheirogaster. Subsequently, transferred by Pérez-García & Vlachos (2014) to the genus Titanochelon.[158]

Eodortoka[159]

Gen. et sp. nov

Valid

Pérez-García, Gasulla & Ortega

Early Cretaceous (Aptian)

Arcillas de Morella Formation

 Spain

A dortokid, a member of the clade Pan-Pleurodira (containing living pleurodirans and all turtles that are more closely related to them than to cryptodirans). The type species is Eodortoka morellana.

Gobiapalone[160]

Gen. et sp. et comb. nov

Valid[161]

Danilov et al.

Late Cretaceous (Cenomanian to Maastrichtian)

Barun Goyot Formation
Bayan Shireh Formation
Nemegt Formation

 Mongolia

A trionychine trionychid. Genus contains new species Gobiapalone breviplastra, as well as "Amyda" orlovi Khosatzky (1976).

Hylaeochelys kappa[162]

Sp. nov

Valid

Pérez-García & Ortega

Late Jurassic (Tithonian)

Freixial Formation

 Portugal

A basal member of Eucryptodira, a species of Hylaeochelys.

Judithemys kranzi[163]

Sp. nov

Valid

Weems

Paleocene (early Thanetian)

Aquia Formation

 United States

A member of (likely paraphyletic) group "Macrobaenidae", a species of Judithemys.

Nemegtemys[160]

Gen. et sp. nov

Valid[161]

Danilov et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

 Mongolia

A cyclanorbine trionychid. The type species is Nemegtemys conflata.

Osonachelus[164]

Gen. et sp. nov

Valid

de Lapparent de Broin et al.

Eocene (late Bartonian)

Vic-Manlleu Marls Formation

 Spain

A cheloniid sea turtle. The type species is Osonachelus decorata.

Procolpochelys charlestonensis[154]

Sp. nov

Valid

Weems & Sanders

Oligocene

 United States

A pancheloniid sea turtle; a species of Procolpochelys.

Riodevemys[165]

Gen. et sp. nov

Valid[166]

Pérez-García, Royo-Torres & Cobos

Late Jurassic

 Spain

A pleurosternid paracryptodiran. The type species is Riodevemys inumbragigas.

Tacuarembemys[167]

Gen. et sp. nov

Valid

Perea et al.

Late Jurassic or earliest Cretaceous

Tacuarembó Formation

 Uruguay

A turtle of uncertain phylogenetic placement. The type species is Tacuarembemys kusterae.

Titanochelon[158]

Gen. et comb. nov

Valid

Pérez-García & Vlachos

Early Miocene to early Pleistocene

 Austria
 Bulgaria
 France
 Germany
 Greece
 Portugal
 Spain
  Switzerland
 Turkey
 Malta?

A tortoise; a new genus for "Testudo" bolivari Hernández-Pacheco (1917). Genus also contains "Testudo" eurysternum Gervais (1848–1852), "Testudo" ginsburgi de Broin (1977), "Testudo" vitodurana Biedermann (1862), "Cheirogaster" steinbacheri Karl (1996), "Testudo" leberonensis Depéret (1890), "Testudo" schafferi Szalai (1931), "Testudo" perpiniana Depéret (1885) and "Cheirogaster" bacharidisi Vlachos et al. (2014). Genus might also contain "Testudo" gymnesicus Bate (1914).

‘Trionyx’ baynshirensis[160]

Sp. nov

Valid[161]

Danilov et al.

Late Cretaceous (Cenomanian to Santonian)

Bayan Shireh Formation

 Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

‘Trionyx’ dissolutus[168]

Sp. nov

Valid

Vitek & Danilov

Late Cretaceous (Cenomanian)

 Uzbekistan

A trionychid, a species of Trionyx sensu lato.

‘Trionyx’ gilbentuensis[160]

Sp. nov

Valid[161]

Danilov et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

 Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

‘Trionyx’ gobiensis[160]

Sp. nov

Valid[161]

Danilov et al.

Late Cretaceous (Maastrichtian)

Nemegt Formation

 Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

‘Trionyx’ shiluutulensis[160]

Sp. nov

Valid[161]

Danilov et al.

Late Cretaceous (Campanian)

 Mongolia

A trionychine trionychid, a species of Trionyx sensu lato.

Archosauromorphs[edit]

Basal archosauromorphs[edit]

Research[edit]

  • A redescription of Tasmaniosaurus triassicus is published by Ezcurra (2014).[169]
  • A revision of anatomy and phylogenetic relationships of the archosauriform Dorosuchus neoetus is published by Sookias et al. (2014).[170]
  • A revision of putative euparkeriids from the Triassic of China is published by Sookias et al. (2014).[171]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aenigmastropheus[172]

Gen. et sp. nov

Valid

Ezcurra, Scheyer & Bulter

Late Permian (Wuchiapingian)

Usili Formation

 Tanzania

A non-archosauriform archosauromorph, probably a protorosaurian. The type species is Aenigmastropheus parringtoni.

Garjainia madiba[173]

Sp. nov

Valid

Gower et al.

Early Triassic (late Olenekian)

Burgersdorp Formation

 South Africa

An erythrosuchid archosauriform, a species of Garjainia.

Hyperodapedon tikiensis[174]

Sp. nov

Valid[175]

Mukherjee & Ray

Late Triassic

Tiki Formation

 India

A rhynchosaur, a species of Hyperodapedon.

Proterosuchus goweri[176]

Sp. nov

Valid[177]

Ezcurra & Butler

Early Triassic

 South Africa

A proterosuchid archosauriform, a species of Proterosuchus.

Pseudochampsa[178]

Gen. et comb. nov

Valid

Trotteyn & Ezcurra

Late Triassic (late Carnian or earliest Norian)

Ischigualasto Formation

 Argentina

A proterochampsid archosauriform; a new genus for "Chanaresuchus" ischigualastensis Trotteyn, Martínez & Alcober (2012).

Pseudosuchians[edit]

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Allodaposuchus palustris[182]

Sp. nov

Valid

Blanco et al.

Late Cretaceous (early Maastrichtian)

Tremp Formation

 Spain

An eusuchian crocodylomorph, a species of Allodaposuchus.

Anthracosuchus[183]

Gen. et sp. nov

Valid

Hastings, Bloch & Jaramillo

Paleocene

Cerrejón Formation

 Colombia

A dyrosaurid crocodyliform. The type species is Anthracosuchus balrogus.

Aplestosuchus[184]

Gen. et sp. nov

Valid

Godoy et al.

Late Cretaceous

Adamantina Formation

 Brazil

A baurusuchid crocodylomorph. The type species is Aplestosuchus sordidus.

Caipirasuchus stenognathus[185]

Sp. nov

Valid

Pol et al.

Late Cretaceous

Adamantina Formation

 Brazil

A sphagesaurid crocodylomorph, a species of Caipirasuchus.

Diplocynodon remensis[186]

Sp. nov

Valid

Martin et al.

Late Palaeocene

 France

A member of Alligatoroidea, a species of Diplocynodon.

Machimosaurus buffetauti[187]

Sp. nov

Valid

Young et al.

Late Jurassic (Kimmeridgian)

 France
 Germany
 Poland?
 United Kingdom?

A teleosaurid crocodylomorph, a species of Machimosaurus. Considered to be a junior synonym of Machimosaurus hugii by Martin, Vincent & Falconnet (2015).[188]

Nundasuchus[189]

Gen. et sp. nov

Valid

Nesbitt et al.

Middle Triassic (Anisian)

Manda Beds

 Tanzania

A member of Archosauriformes of uncertain phylogenetic placement, possibly a pseudosuchian. The type species is Nundasuchus songeaensis.

Polesinesuchus[190]

Gen. et sp. nov

Valid

Roberto-da-Silva et al.

Late Triassic

Santa Maria Formation

 Brazil

An aetosaur. The type species is Polesinesuchus aurelioi.

Rukwasuchus[191]

Gen. et sp. nov

Valid

Sertich & O’Connor

Cretaceous (Aptian to Cenomanian)

Galula Formation

 Tanzania

A peirosaurid crocodyliform. The type species is Rukwasuchus yajabalijekundu.

Sahitisuchus[192]

Gen. et sp. nov

Valid

Kellner, Pinheiro & Campos

Paleocene (Itaboraian)

 Brazil

A sebecid crocodylomorph. The type species is Sahitisuchus fluminensis.

Wannchampsus[193]

Gen. et sp. nov

Valid

Adams

Early Cretaceous (late Aptian)

Twin Mountains Formation

 United States

A paralligatorid neosuchian crocodylomorph. The type species is Wannchampsus kirpachi.

Non-avian dinosaurs[edit]

Research[edit]

  • A study of size changes and rates of anatomical innovation in the theropod lineage ancestral to birds is published by Lee et al. (2014).[194]
  • A study of evolution of body size and forelimb length in birds and nov-avian coelurosaurian theropods is published by Puttick, Thomas and Benton (2014).[195]
  • A phylogenetic analysis of bird and non-avian coelurosaurian theropod relationships and a study of rates of morphological evolution and changes in morphological disparity across the dinosaur-bird transition is published by Brusatte et al. (2014).[196]
  • A description of abelisaurid teeth from the Late Jurassic Lourinhã Formation of Portugal and a phylogenetic analysis of theropod relationships based on dental characters is published by Hendrickx and Mateus (2014).[197]
  • A study of theropod diversity in the Cretaceous (Aptian-Albian) of Tunisia is published by Fanti et al. (2014).[198]
  • A juvenile specimen of Megaraptor namunhuaiquii is described by Porfiri et al. (2014).[199]
  • A study of European fossil record of Ornithomimosauria is published by Allain et al. (2014).[200]
  • A study of flight ability in some non-avian paravian theropods is published by Sorkin (2014).[201]
  • "Saurornitholestes" robustus, initially thought to be a dromaeosaurid, is reinterpreted as a troodontid by Evans et al. (2014).[202]
  • A well-preserved specimen of Microraptor zhaoianus is described by Pei et al. (2014).[203]
  • A study of anatomy and phylogenetic relationships of Antetonitrus ingenipes is published by McPhee et al. (2014).[204]
  • Fragmentary partial skeleton of a small sauropod belonging to the genus Haplocanthosaurus collected from the Rocky Mountains of central Colorado is described by Foster & Wedel (2014).[205]
  • The purported size of the holotype vertebra of Amphicoelias fragillimus is reevaluated by Woodruff and Foster (2014).[206]
  • A study of phylogenetic relationships of Lourinhasaurus alenquerensis is published by Mocho, Royo-Torres and Ortega (2014).[207]
  • A study of anatomy and phylogenetic relationships of Aragosaurus ischiaticus is published by Royo-Torres et al. (2014).[208]
  • A study of titanosaur osteoderms from the Upper Cretaceous Lo Hueco site in Cuenca, Spain is published by Vidal, Ortega and Sanz (2014).[209]
  • A study of species richness of South American titanosaur assemblages during the Late Cretaceous is published by Vieira et al. (2014).[210]
  • A study of the effect of intervertebral cartilage on neck posture of sauropod dinosaurs is published by Taylor (2014).[211]
  • A study of the dentition of Manidens condorensis is published by Becerra et al. (2014).[212]
  • A study of the postcranial anatomy of Heterodontosaurus tucki is published by Galton (2014).[213]
  • A study of the impact of osteoderm placement on the centre of mass of stegosaurs is published by Mallison (2014).[214]
  • A study of Early Cretaceous Spanish iguanodont ornithopod diversity and a description of new remains referrable to Delapparentia is published by Gasca, Canudo and Moreno-Azanza (2014).[215]
  • A specimen of Edmontosaurus regalis with remains of a soft-tissue cranial crest is described by Bell et al. (2014).[216]
  • A juvenile specimen of Edmontosaurus annectens is described by Prieto-Márquez (2014).[217]
  • An assemblage of Psittacosaurus juveniles associated with a larger specimen from the Lujiatun beds of the Yixian Formation in Liaoning, China is described by Hedrick et al. (2014).[218]
  • An aggregation of four juveniles of Protoceratops andrewsi from the Tugrikin Shire locality of the Djadokhta Formation in Central Gobi region, Mongolia and two associated subadults of the same species from the same locality are described by Hone et al. (2014).[219]
  • A study of ontogenetic changes in the craniofacial skeleton of Centrosaurus apertus is published by Frederickson and Tumarkin-Deratzian (2014).[220]
  • A new specimen attributable to Arrhinoceratops brachyops is described by Mallon et al. (2014).[221]
  • A new specimen of Spinosaurus is described by Ibrahim et al., with a controversial reconstruction of Spinosaurus as a quadrupedal semi-aquatic genus.[222]
  • Two new specimens of the previous enigma Deinocheirus are described and analysed by Lee et al. (2014).[223]
  • A tiny theropod was found in the South Korea. (2014)

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Adelolophus[224]

Gen. et sp. nov

Valid

Gates et al.

Late Cretaceous (Campanian)

Wahweap Formation

 United States

A lambeosaurine hadrosaurid. The type species is Adelolophus hutchisoni.

Allosaurus lucasi[225]

Sp. nov

Valid

Dalman

Late Jurassic (Tithonian)

Morrison Formation

 United States

An allosauroid theropod, a species of Allosaurus.

Amargastegos[226]

Gen. et sp. nov

Disputed

Ulansky

Early Cretaceous (Barremian-early Aptian)

La Amarga Formation

 Argentina

A stegosaur. The type species is Amargastegos brevicollus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Andhrasaurus[226]

Gen. et. sp. nov

Disputed

Ulansky

Early Jurassic (Pliensbachian-Toarcian)

Kota Formation

 India

A basal thyreophoran. The type species is Andhrasaurus indicus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Anzu[228]

Gen. et sp. nov

Valid

Lamanna et al.

Late Cretaceous (late Maastrichtian)

Hell Creek Formation

 United States

A caenagnathid theropod. The type species is Anzu wyliei.

Aquilops[229]

Gen. et sp. nov

Valid

Farke et al.

Early Cretaceous (Albian)

Cloverly Formation

 United States

A basal member of Neoceratopsia. The type species is Aquilops americanus.

Arcovenator[230]

Gen. et sp. nov

Valid

Tortosa et al.

Late Cretaceous (late Campanian)

 France

An abelisaurid theropod. The type species is Arcovenator escotae.

Camarillasaurus[231]

Gen. et sp. nov

Valid

Sánchez-Hernández & Benton

Early Cretaceous (early Barremian)

Camarillas Formation

 Spain

A basal ceratosaurian. The type species is Camarillasaurus cirugedae.

Changyuraptor[232]

Gen. et sp. nov

Valid

Han et al.

Early Cretaceous

Yixian Formation

 China

A microraptorine dromaeosaurid theropod. The type species is Changyuraptor yangi.

Chuanqilong[233]

Gen. et sp. nov

Valid

Han et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

An ankylosaurid. The type species is Chuanqilong chaoyangensis.

Chungkingosaurus giganticus[226]

Sp. nov

Disputed

Ulansky

Late Jurassic (Oxfordian)

Shangshaximiao Formation

 China

A huayangosaurid, a species of Chungkingosaurus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Chungkingosaurus magnus[226]

Sp. nov

Disputed

Ulansky

Late Jurassic (Oxfordian)

Shangshaximiao Formation

 China

A huayangosaurid, a species of Chungkingosaurus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Datanglong[234]

Gen. et sp. nov

Valid

Mo et al.

Early Cretaceous

Xinlong Formation

 China

A carcharodontosaur theropod. The type species is Datanglong guangxiensis.

Dreadnoughtus[235]

Gen. et sp. nov

Valid

Lacovara et al.

Late Cretaceous (Campanian or Maastrichtian)

Cerro Fortaleza Formation

 Argentina

A titanosaurian sauropod. The type species is Dreadnoughtus schrani.

Eoplophysis[226]

Gen. et comb. nov

Disputed

Ulansky

Middle Jurassic (Bathonian)

Cornbrash Formation

 England

A stegosaur; a new genus for "Omosaurus" vetustus von Huene (1910). According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Eousdryosaurus[236]

Gen. et sp. nov

Valid

Escaso et al.

Late Jurassic

Alcobaça Formation

 Portugal

A dryosaurid ornithopod. The type species is Eousdryosaurus nanohallucis.

Ferganastegos[226]

Gen. et sp. nov

Disputed

Ulansky

Middle Jurassic (Callovian)

Balabansai Formation

 Kyrgyzstan

A stegosaur. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Fosterovenator[237]

Gen. et sp. nov

Valid

Dalman

Late Jurassic (Kimmeridgian/Tithonian)

Morrison Formation

 United States

A theropod dinosaur. The type species is Fosterovenator churei.

Gobivenator[238]

Gen. et sp. nov

Valid

Tsuihiji et al.

Late Cretaceous (Campanian)

Djadochta Formation

 Mongolia

A troodontid theropod. The type species is Gobivenator mongoliensis.

Gongpoquansaurus[239]

Gen. et comb. nov

Valid

You, Li & Dodson

Early Cretaceous (Albian)

Zhonggou Formation

 China

A non-hadrosaurid hadrosauroid; a new genus for "Probactrosaurus" mazongshanensis Lü (1997).

Huangshanlong[240]

Gen. et sp. nov

Valid

Huang et al.

Middle Jurassic

Hongqin Formation

 China

A mamenchisaurid sauropod. The type species is Huangshanlong anhuiensis.

Kulindadromeus[241]

Gen. et sp. nov

Valid

Godefroit et al.

Middle or Late Jurassic (Bajocian to Tithonian)

Ukureyskaya Formation

 Russia

A non-cerapodan neornithischian. The type species is Kulindadromeus zabaikalicus.

Laquintasaura[242]

Gen. et sp. nov

Valid

Barrett et al.

Early Jurassic

La Quinta Formation

 Venezuela

A basal ornithischian. The type species is Laquintasaura venezuelae.

Leinkupal[243]

Gen. et sp. nov

Valid

Gallina et al.

Early Cretaceous (late Berriasian to Valanginian)

Bajada Colorada Formation

 Argentina

A diplodocine diplodocid sauropod. The type species is Leinkupal laticauda.

Mercuriceratops[244]

Gen. et sp. nov

Valid

Ryan et al.

Late Cretaceous (Campanian)

Dinosaur Park Formation
Judith River Formation

 Canada
 United States

A chasmosaurine ceratopsid. The type species is Mercuriceratops gemini.

Nanuqsaurus[245]

Gen. et sp. nov

Valid

Fiorillo & Tykoski

Late Cretaceous (Maastrichtian)

Prince Creek Formation

 United States

A tyrannosaurid theropod. The type species is Nanuqsaurus hoglundi.

Natronasaurus[226]

Gen. et comb. nov

Disputed

Ulansky

Late Jurassic (Kimmeridgian-Tithonian)

Morrison Formation

 United States

A stegosaurid; a new genus for "Stegosaurus" longispinus Gilmore (1914). According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature; Galton and Carpenter made "S." longispinus the type species of a new genus Alcovasaurus.[227]

Panguraptor[246]

Gen. et sp. nov

Valid

You et al.

Early Jurassic

Lufeng Formation

 China

A coelophysoid theropod. The type species is Panguraptor lufengensis.

Pentaceratops aquilonius[247]

Sp. nov

Valid

Longrich

Late Cretaceous (late Campanian)

Dinosaur Park Formation

 Canada

A ceratopsid, a species of Pentaceratops.

Plesiohadros[248]

Gen. et sp. nov

Valid

Tsogtbaatar et al.

Late Cretaceous (probably late Campanian)

Djadochta Formation

 Mongolia

A non-hadrosaurid hadrosauroid. The type species is Plesiohadros djadokhtaensis.

Qianzhousaurus[249]

Gen. et sp. nov

Valid

et al.

Late Cretaceous (Maastrichtian)

 China

A tyrannosaurid theropod. The type species is Qianzhousaurus sinensis.

Quetecsaurus[250]

Gen. et sp. nov

Valid

González Riga & Ortiz David

Late Cretaceous (Turonian)

Cerro Lisandro Formation

 Argentina

A titanosaur sauropod related to Mendozasaurus and Futalognkosaurus. The type species is Quetecsaurus rusconii.

Rhinorex[251]

Gen. et sp. nov

Valid[252]

Gates & Scheetz

Late Cretaceous (Campanian)

Neslen Formation

 United States

A saurolophine hadrosaurid. The type species is Rhinorex condrupus.

Rukwatitan[253]

Gen. et sp. nov

Valid

Gorscak et al.

Cretaceous (Aptian to Cenomanian)

Galula Formation

 Tanzania

A titanosaur sauropod. The type species is Rukwatitan bisepultus.

Saldamosaurus[226]

Gen. et sp. nov

Disputed

Ulansky

Late Jurassic

Saldam Formation

 Russia

A stegosaur. The type species is Saldamosaurus tuvensis. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Scelidosaurus arizonensis[226]

Sp. nov

Disputed

Ulansky

Early Jurassic (?Sinemurian-Pliensbachian)

Kayenta Formation

 United States

A basal thyreophoran, a species of Scelidosaurus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Siamodracon[226]

Gen. et sp. nov

Disputed

Ulansky

Late Jurassic

Phu Kradung Formation

 Thailand

A stegosaur. The type species is Siamodracon altispinus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Sinopeltosaurus[226]

Gen. et sp. nov

Disputed

Ulansky

Early Jurassic (Hettangian-Pliensbachian)

Lufeng Formation

 China

A basal thyreophoran. The type species is Sinopeltosaurus minimus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Tachiraptor[254]

Gen. et sp. nov

Valid

Langer et al.

Earliest Jurassic

La Quinta Formation

 Venezuela

A stem-averostran theropod. The type species is Tachiraptor admirabilis.

Tambatitanis[255]

Gen. et sp. nov

Valid

Saegusa and Ikeda

Early Cretaceous (probably early Albian)

Sasayama Group

 Japan

A titanosauriform sauropod, a member of Somphospondyli. The type species is Tambatitanis amicitiae.

Torvosaurus gurneyi[256]

Sp. nov

Valid

Hendrickx & Mateus

Late Jurassic (late Kimmeridgian)

Lourinhã Formation

 Portugal

A megalosauroid theropod, a species of Torvosaurus.

Vahiny[257]

Gen. et sp. nov

Valid

Curry Rogers & Wilson

Late Cretaceous

Maevarano Formation

 Madagascar

A titanosaur sauropod. The type species is Vahiny depereti.

Wuerhosaurus mongoliensis[226]

Sp. nov.

Disputed

Ulansky

Early Cretaceous (Aptian-Albian)

Khukhtyk Formation

 Mongolia

A stegosaur, a species of Wuerhosaurus. According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.[227]

Yongjinglong[258]

Gen. et sp. nov

Valid

Li et al.

Early Cretaceous

Hekou Group

 China

A titanosaur sauropod. The type species is Yongjinglong datangi.

Zaraapelta[259]

Gen. et sp. nov

Valid

Arbour, Currie & Badamgarav

Late Cretaceous

Barun Goyot Formation

 Mongolia

A member of Ankylosauridae. The type species is Zaraapelta nomadis.

Zby[260]

Gen. et sp. nov

Valid

Mateus, Mannion & Upchurch

Late Jurassic (late Kimmeridgian)

Lourinhã Formation

 Portugal

A turiasaurian sauropod. The type species is Zby atlanticus.

Zhanghenglong[261]

Gen. et sp. nov

Valid

Xing et al.

Late Cretaceous (Santonian)

Majiacun Formation

 China

A non-hadrosaurid hadrosauroid ornithopod. The type species is Zhanghenglong yangchengensis.

Ziapelta[262]

Gen. et sp. nov

Valid

Arbour et al.

Late Cretaceous (late Campanian)

Kirtland Formation

 United States

An ankylosaurid. The type species is Ziapelta sanjuanensis.

Birds[edit]

Research[edit]

  • A study on the antiquity of birds is published by Lee et al. (2014).[263]
  • A new specimen of Archaeopteryx is described by Foth, Tischlinger and Rauhut (2014).[264]
  • Zhongornis haoae, initially thought to be a bird, is argued to be a non-avian maniraptoran by O'Connor and Sullivan (2014).[265]
  • A study of ecological disparity in Early Cretaceous birds is published by Mitchell and Makovicky (2014).[266]
  • A subadult specimen of Zhouornis hani is described by Zhang et al. (2014).[267]
  • A study of the species status of the moa from the genus Euryapteryx is published by Huynen and Lambert (2014).[268]
  • A study of growth in the moa from the genus Euryapteryx is published by Huynen et al. (2014).[269]
  • A study of tbx5 gene of the moa from the genus Dinornis is published by Huynen et al. (2014).[270]
  • Fossil remains of a relative of the hoatzin, possibly a species belonging to the genus Namibiavis, from the middle Miocene of Kenya, are described by Mayr (2014).[271]
  • A specimen of Pumiliornis tessellatus with preserved stomach contents including pollen grains is described by Mayr and Wilde (2014).[272]
  • A Late Pleistocene specimen of griffon vulture (Gyps fulvus) with exceptionally well preserved fossilized soft tissues is described from the Alban Hills volcanic region, Italy by Iurino et al. (2014).[273]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aegypius varswaterensis [274]

Sp nov.

Valid

Albrecht Manegold,

Marco Pavia,

Pippa Haarhoff

Early Pliocene

Varswater Formation

 South Africa

An Aegypius sp. related to the cinereous vulture

Anser djuktaiensis[275]

Sp nov.

Valid

Zelenkov & Kurochkin

Late Pleistocene

 Russia

A species of Anser.

Australornis lovei [276]

Gen. et Sp. nov.

Valid

Mayr & R. Paul Scofield

Early Paleocene

Waipara Greensand

 New Zealand

An incertae sedis Neognathae, Monotypic with A. lovei.

Bambolinetta lignitifila [277]

Gen. et Comb. nov.

Valid

Mayr & Pavia

Miocene

 Italy

An Anatinae genus, Comb. nov. for "Anas" lignitifila

Baselrallus intermedius [278]

Gen. et Sp. nov.

Valid

De Pietri,

Gerald Mayr

Early Miocene

 France

A member of the Rallidae. The type species of the new genus.

Caracara seymouri [279]

Sp. nov.

Valid

Suarez & Olson

Quaternary

 Ecuador
 Peru

A Caracara, Falconidae.

Cryptopsar ischyrhynchus [280]

Gen. et Sp. nov.

Valid

Hume

Holocene

 Mauritius

A member of the Sturnidae, it is the type species of the new genus.

Eopengornis martini [281]

Gen. et Sp. nov.

Valid

Wang, O'Connor, Zheng,
Wang, Hu, Zhou

Early Cretaceous,

Huajiying Formation

 China

A member of the Enantiornithes, it is the type species of the new genus.

Eudyptes calauina [282]

Sp. nov.

Valid

Hoffmeister, Carrillo Briceño, & Nielsen

Pliocene

Horcon Formation

 Chile

An extinct crested penguin.

Evgenavis nobilis [283]

Gen. et Sp. nov.

Valid

O’Connor, Averianov, & Zelenkov

Barremian-Aptian

Ilek Formation

 Russia

Aves incertae sedis. it is the type species of the new genus.

Falco hezhengensis [284]

Sp. nov.

Valid

Zhiheng, Zhonghe, Tao,
Qiang, & Clarke

Late Miocene

Linxia Basin

 China

A member of the Falconidae.

Fortunguavis xiaotaizicus [285]

Gen. nov. et Sp. nov.

Valid

Wang Min

Jingmai K. O’Connor

Zhou Zhonghe

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes Walker, 1981. This is the type species of the new genus.

Gansus zheni [286]

Sp. nov.

Valid

Di Liu

Luis M. Chiappe

Yuguang Zhang

Alyssa Bell

Qingjin Meng

Qiang Ji

Xuri Wang

Early Cretaceous

Jiufotang Formation

 China

An early member of Ornithuromorpha Chiappe, 2002, a species of Gansus.

Garganornis ballmanni [287]

Gen. nov. et Sp. nov.

Valid

Hanneke J. M. Meijer

Miocene

Fissure fillings

 Italy

A member of the Anseriformes, this is the type species of the new genus.

Genucrassum bransatensis [288]

Gen. nov. et Sp. nov.

Valid

Vanesa L. De Pietri
R. Paul Scofield

Late Oligocene

MP 30; 23.03 Mya

 France

A stone-curlew, Burhinidae, this is the type species of the new genus.

Grabauornis lingyuanensis[289]

Gen. nov. et Sp. nov.

Valid

Johan Dalsätt

Per G.P. Ericson

Zhou Zhonghe

Early Cretaceous

Yixian Formation

 China

A member of Enantiornithes Walker, 1981. This is the type species of the new genus.

?Ibidopodia chavrochensis [290]

Sp. nov.

Valid

Vanesa L. De Pietri

Gerald Mayr

Early Miocene

Chavroches

MN 2

 France

An Idiornithid, Idiornithidae Cariamiformes, possibly a species of Ibidopodia Milne-Edwards, 1868.

?Ibidopodia minuta [290]

Sp. nov.

Valid

Vanesa L. De Pietri

Gerald Mayr

Early Miocene

Saulcet

 France

An Idiornithid, Idiornithidae Cariamiformes, possibly a species of Ibidopodia Milne-Edwards, 1868.

Iteravis huchzermeyeri[291]

Gen. nov. et sp. nov.

Valid

Shuang Zhou

Jingmai K. O’Connor

Min Wang

Early Cretaceous

Yixian Formation

 China

A basal member of the Ornithuromorpha Chiappe, 2002. This is the type species of the new genus.

Jeholornis curvipes [292]

Sp. nov.

Valid

Ulysse Lefèvre

Dongyu Hu

François Escuillié

Gareth Dyke

Pascal Godefroit

Early Cretaceous

Yixian Formation

 China

A member of Avialae Gauthier, 1986, a species of Jeholornis Zhou et Zhang, 2002.

Limnofregata hutchisoni [293]

Sp. nov.

Valid[294]

Thomas A. Stidham

Early Eocene

Wasatch Formation

 United States:

 Wyoming

A member of Fregatidae, a species of Limnofregata.

Longusunguis kurochkini [295]

Gen. nov. et Sp. nov.

Valid

Wang Min

Zhou Zhonghe

Jingmai K. O'Connor

Nikita V. Zelenkov

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes Walker, 1981 related to Bohaiornis. This is the type species of new genus.

Mergellus mochanovi [275]

Sp. nov.

Valid

Nikita V. Zelenkov

Evgeny N. Kurochkin

Late Pleistocene

 Russia

A member of the Anatidae related to the smew, a species of Mergellus.

Mergus milleneri [296]

Sp. nov.

Valid

Murray Williams

Alan J. D. Tennyson

Holocene

Chatham Island

 New Zealand

A member of the Anatidae.

Nectornis africanus [297]

Sp. nov.

Valid

Gerald Mayr

Middle Miocene

MN 5

 Kenya

A cormorant, Phalacrocoracidae, a species of Nectornis Cheneval, 1984.

Nestor chathamensis[298]

Sp. nov.

Valid

Jamie R. Wood

Kieren J. Mitchell

R. Paul Scofield

Alan J. D. Tennyson in Wood et al.

Holocene

Chatham Islands

 New Zealand

A New Zealand parrot belonging to the family Nestoridae Bonaparte, 1849, a species of Nestor.

?Nupharanassa mabokoensis [297]

Sp. nov.

Valid

Gerald Mayr

Middle Miocene

MN 5

 Kenya

A jacana, Jacanidae, possibly a species of Nupharanassa Rasmussen, Olson et Simons, 1987.

Parabohaiornis martini [295]

Gen. nov. et Sp. nov.

Valid

Wang Min

Zhou Zhonghe

Jingmai K. O’Connor

Nikita V. Zelenkov

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes Walker, 1981 related to Bohaiornis. The type species of the new genus, placed in the family Bohaiornithidae Wang, Zhou, O’Connor et Zelenkov, 2014 with de genera Shenqiornis Wang, O’Connor, Zhao, Chiappe, Gao et Cheng, 2010, Sulcavis O’Connor, Zhang, Chiappe, Meng, Quanguo et Di, 2013, Zhouornis Zhang, Chiappe, Han et Chinsamy, 2013, Longusunguis Wang, Zhou, O’Connor et Zelenkov, 2014 and Bohaiornis Hu, Hou L. H. et Xu, 2011.

Parvavis chuxiongensis [299]

Gen. nov. et Sp. nov.

Valid

Wang Min

Zhou Zhonghe

Xu Guanghui

Late Cretaceous,

Turonian to Santonian

Jiangdihe Formation

 China

A member of Enantiornithes Walker, 1981. The type species of the new genus.

Pelagornis sandersi [300]

Sp. nov.

Valid

Daniel T. Ksepka

Late Oligocene

Chandler Bridge Formation,

Early Chattian

 USA:

 South Carolina

A member of the Pelagornithidae Fürbringer, 1888, a species of Pelagornis Lartet, 1857; the bird with the biggest wingspan known up to now.

Piscivoravis lii [301]

Gen. nov. et Sp. nov.

Valid

Zhou Shuang

Zhou Zhonghe

Jingmai K. O'Connor

Early Cretaceous

Jiufotang Formation

 China

A basal member of the Ornithuromorpha Chiappe, 2002. This is the type species the new genus.

Plotornis graculoides [277]

Comb. nov.

Valid

Gerald Mayr

Marco Pavia

Miocene

MN 1-2

 Italy

A member of the Diomedeidae, a new name for "Chenornis" graculoides Portis 1884, creating a Comb. nov.

Protoazin parisiensis [302]

Gen. nov. et Sp. nov.

Valid

Gerald Mayr

Vanesa L. De Pietri

Late Eocene

Priabonian,

MP 20

 France

A relative of the hoatzin, Opisthocomidae. This is the type species of the new genus.

Sobniogallus albinojamrozi [303]

Gen. et Sp. nov.

Valid

Tomek, Bocheński, Wertz, & Świdnicka

Rupelian

Krosno Formation

 Poland

Galliformes incertae sedis. This is the type species of the new genus.

Tianyuornis cheni [304]

Gen. et Sp. nov.

Valid

Xiao-Ting, O'Connor,
Xiao-Li, Xiao-Mei, & Yan

Early Cretaceous

Yixian Formation

 China

A member of the Hongshanornithidae, the type species of the new genus.

Pterosaurs[edit]

Research[edit]

  • A study of pneumaticity of pterosaur wing bones is published by Martin and Palmer (2014).[305]
  • A complete and fully articulated juvenile specimen of Scaphognathus crassirostris is described by Bennett (2014).[306]
  • Partial 3D egg of Pterodaustro guinazui is described by Grellet-Tinner et al. (2014).[307]
  • Cearadactylus atrox is redescribed by Vila Nova et al. (2014).[308]
  • The taxonomy and distribution of the family Azhdarchidae is reviewed by Averianov (2014).[309]
  • A reevaluation of the fossil material attributed to Bakonydraco galaczi, indicating that the fossils actually represent at least two pterosaur taxa, is published by Prondvai, Bodor and Ősi (2014).[310]
  • A study of medullary bone-like tissue in the mandibular symphyses of Bakonydraco galaczi is published by Prondvai and Stein (2014).[311]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Aerodactylus[312]

Gen. et comb. nov

Valid

Vidovic & Martill

Late Jurassic

Solnhofen Limestone

 Germany

A member of Pterodactyloidea related to Gladocephaloideus, Cycnorhamphus, Ardeadactylus and Aurorazhdarcho; a new genus for "Pterodactylus" scolopaciceps Meyer (1860).

Boreopterus giganticus[313]

Sp. nov

Valid

Jiang et al.

Early Cretaceous

 China

A species of Boreopterus.

Caiuajara[314]

Gen. et sp. nov

Valid

Manzig et al.

Late Cretaceous

Bauru Basin, Caiuá Group, Goio-Erê Formation

 Brazil

A tapejarid. The type species is Caiuajara dobruskii.

Hamipterus[315]

Gen. et sp. nov

Valid

Wang et al.

Early Cretaceous

Tugulu Group

 China

A member of Pteranodontoidea of uncertain phylogenetic placement. The type species is Hamipterus tianshanensis.

Ikrandraco[316]

Gen. et sp. nov

Valid

Wang et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A non-anhanguerian pteranodontoid. The type species is Ikrandraco avatar.

Kryptodrakon[317]

Gen. et sp. nov

Valid

Andres, Clark & Xu

Middle-Late Jurassic boundary

 China

An early member of Pterodactyloidea. The type species is Kryptodrakon progenitor.

Maaradactylus[318]

Gen. et sp. nov

Valid

Bantim et al.

Early Cretaceous (Aptian or Albian)

Romualdo Formation

 Brazil

A member of Anhangueridae. The type species is Maaradactylus kellneri.

Other reptiles[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Abyssomedon[319]

Gen. et sp. nov

Valid

Macdougall & Reisz

Early Permian

Garber Formation

 United States

A nyctiphruretid parareptile. The type species is Abyssomedon williamsi.

Atopodentatus[320]

Gen. et sp. nov

Valid

Cheng et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A marine reptile, probably a relative of sauropterygians. The type species is Atopodentatus unicus.

Delorhynchus cifellii[321]

Sp. nov

Valid

Reisz, Macdougall & Modesto

Early Permian

Garber Formation

 United States

A parareptile relative of lanthanosuchoids, a species of Delorhynchus.

Eohupehsuchus[322]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic

Jialingjiang Formation

 China

A hupehsuchian. The type species is Eohupehsuchus brevicollis.

Horaffia[323]

Gen. et sp. nov

Valid

Klein & Hagdorn

Middle Triassic (Ladinian)

Erfurt Formation

 Germany

A marine diapsid reptile of uncertain phylogenetic placement. The type species is Horaffia kugleri.

Largocephalosaurus qianensis[324]

Sp. nov

Valid

Li et al.

Triassic

Guanling Formation

 China

A member of Saurosphargidae, a species of Largocephalosaurus.

Parahupehsuchus[325]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic

Jialingjiang Formation

 China

A hupehsuchian. The type species is Parahupehsuchus longus.

Synapsids[edit]

Non-mammalian synapsids[edit]

Research[edit]

New taxa[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Abajudon[328]

Gen. et sp. nov

Valid

Angielczyk et al.

Permian

Ruhuhu Formation

 Tanzania

A dicynodont of uncertain phylogenetic placement. The type species is Abajudon kaayai.

Alierasaurus[329]

Gen. et sp. nov

Valid

Romano & Nicosia

Permian

Cala del Vino Formation

 Italy

A caseid. The type species is Alierasaurus ronchii.

Botucaraitherium[330]

Gen. et sp. nov

Valid

Soares, Martinelli & De Oliveira

Late Triassic (possibly early Norian)

Riograndia Assemblage Zone of the Candelária Sequence (Santa Maria Supersequence)

 Brazil

A prozostrodontian cynodont. The type species is Botucaraitherium belarminoi.

Eocasea[331]

Gen. et sp. nov

Valid

Reisz & Fröbisch

Carboniferous (late Pennsylvanian)

Calhoun Shale

 United States

A member of Caseidae . The type species is Eocasea martini.

Sungeodon[332]

Gen. et sp. nov

Valid

Maisch & Matzke

Early Triassic

Jiucaiyuan Formation

 China

A kannemeyeriiform dicynodont . The type species is Sungeodon kimkraemerae.

Mammals[edit]

Other animals[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Akmolina? tubulus[333]

Sp. nov

Valid

Percival et al.

Cambrian (Furongian)

Sluice Box Formation

 New Zealand

A brachiopod belonging to the group Acrotretida and the family Ephippelasmatidae.

Ascarites rufferi[334]

Sp. nov

Valid

Da Silva et al.

Middle Triassic (Ladinian)

 Brazil

An ascaridid nematode found in a cynodont coprolite, a species of Ascarites.

Ashetscolex[335]

Gen. et sp. nov

Valid

Muir et al.

Ordovician (earliest Floian)

Tonggao Formation

 China

A palaeoscolecid worm. The type species is Ashetscolex nuppus.

Athyris africana[336]

Sp. nov

Valid

Schemm-Gregory

Devonian (late Givetian)

Western Sahara

An athyridid brachiopod, a species of Athyris.

Atraktoprion podolicus[337]

Sp. nov

Valid

Szaniawski & Drygant

Early Devonian (middle Lochkovian to Pragian)

 Ukraine

An atraktoprionid polychaete, a species of Atraktoprion.

Beethovenia[338]

Gen. et 3 sp. et comb. nov

Valid

García-Alcalde

Devonian (Pragian and Emsian)

 Belgium
 France
 Spain
 Germany?

A strophodontid brachiopod. The type species is Beethovenia beethoveni; genus also includes new species B. bachi and B. brahmsi, as well as B. acutiplicata (Oehlert & Davoust, 1879) and possibly "Stropheodonta" steiningeri Drevermann (1907).

Cambroskiadeion[339]

Gen. et sp. nov

Valid

Moore, Porter & Li

Early Cambrian

Dengying Formation

 China

An animal of uncertain phylogenetic placement with a cap-shaped shell. The type species is Cambroskiadeion xiaowanense.

Cirpa alkayae[340]

Sp. nov

Valid

Vörös

Early Jurassic (Pliensbachian)

 Turkey

A wellerellid rhynchonellid brachiopod, a species of Cirpa.

Conchicolites crispisulcans[341]

Sp. nov

Valid

Vinn, Jarochowska & Munnecke

Silurian (Wenlock)

Halla Formation

 Sweden

A member of Cornulitida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Conchicolites.

Eklexibella[342]

Gen. et 2 sp. nov

Valid

Keupp, Gründel & Wiese

Early Jurassic (late Pliensbachian)

 Germany

A terebellid polychaete. Genus contains two species: Eklexibella buttenheimensis and Eklexibella johanni.

Enalikter[343]

Gen. et sp. nov

Valid

Siveter et al.

Silurian

Herefordshire Lagerstätte

 United Kingdom

An animal of uncertain phylogenetic placement; it might be a megacheiran arthropod[343][344] or an annelid.[345] The type species is Enalikter aphson.

Eopriapulites[346]

Gen. et sp. nov

Valid

Liu & Xiao in Liu et al.

Cambrian (Fortunian)

Kuanchuanpu Formation

 China

A priapulid-like scalidophoran. The type species is Eopriapulites sphinx.

Eximipriapulus[347]

Gen. et sp. nov

Valid

Ma et al.

Cambrian

Chengjiang Lagerstätte

 China

A member of Priapulida. The type species is Eximipriapulus globocaudatus.

Filihernodia[348]

Gen. et sp. nov

Valid

Taylor & Wilson in Frey et al.

Early Devonian (Pragian)

 Morocco

A reptariid hederellid. The type species is Filihernodia buccina.

Foliomena xinjiangensis[349]

Sp. nov

Valid

Zhan et al.

Ordovician (late Katian)

Yinpingshan Formation

 China

A brachiopod.

Haplistion toftanum[350]

Sp. nov

Valid

Rhebergen & Botting

Silurian (Telychian)

 Sweden

A haplistiid spirosclerophorid demosponge, a species of Haplistion.

Hyalonema vetteri[351]

Sp. nov

Valid

Janussen

Late Cretaceous (Coniacian)

Arnager Formation

 Denmark

A hexactinellid sponge, a species of Hyalonema.

Kassinella tarimensis[349]

Sp. nov

Valid

Zhan et al.

Ordovician (late Katian)

Yinpingshan Formation

 China

A brachiopod.

Leoniorthis rubeli[352]

Sp. nov

Valid

Hints

Ordovician

 Estonia
 Russia

A brachiopod belonging to the group Orthida and the family Orthidae.

Lindstroemispongia[350]

Gen. et sp. nov

Valid

Rhebergen & Botting

Silurian (Telychian)

 Sweden

An astylospongiid streptosclerophorid demosponge. The type species is Lindstroemispongia cylindrata.

Lyrarapax[353]

Gen. et sp. nov

Valid

Cong et al.

Early Cambrian

 China

A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods). The type species is Lyrarapax unguispinus.

Maruia[333]

Gen. et sp. nov

Valid

Percival et al.

Cambrian (Furongian)

Sluice Box Formation

 New Zealand

A brachiopod, possibly a member of Acrotretida. The type species is M. castellum.

Mirilingula antipodes[333]

Sp. nov

Valid

Percival et al.

Cambrian (Furongian)

Sluice Box Formation

 New Zealand

A brachiopod belonging to the group Linguloidea and the family Lingulellotretidae.

Multistella leipnitzae[350]

Sp. nov

Valid

Rhebergen & Botting

Silurian (Telychian)

 Sweden

An anthaspidellid orchocladine demosponge, a species of Multistella.

Nesonektris[354]

Gen. et sp. nov

Valid

García-Bellido et al.

Cambrian

Emu Bay Shale

 Australia

A member of Vetulicolida. The type species is Nesonektris aldridgei.

Nidelric[355]

Gen. et sp. nov

Valid

Hou et al.

Early Cambrian

Heilinpu Formation

 China

An animal of uncertain phylogenetic placement; a member of Chancelloriida or a chancelloriid-like animal. The type species is Nidelric pugio.

Nisusia granosa[356]

Sp. nov

Valid

Mao, Wang & Zhao

Cambrian

Kaili Formation

 China

A brachiopod, a species of Nisusia.

Norvegiograptus[357]

Gen. et comb. nov

Valid

Li et al.

Early Ordovician

 China
 Norway

A graptolite belonging to the family Dichograptidae. A new genus for "Didymograptus" liber Monsen (1937); genus also includes "Didymograptus" enshiensis Ni in Mu et al. (1979).

Oikozetetes mounti[358]

Sp. nov

Valid

Jacquet, Brock & Paterson

Early Cambrian

Mernmerna Formation

 Australia

A member of Halkieriidae (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Oikozetetes.

Opetionella incompta[350]

Sp. nov

Valid

Rhebergen & Botting

Silurian (Telychian)

 Sweden

A demosponge, possibly a member of the group Hadromerida; a species of Opetionella.

Paleoxyuris[359]

Gen. et sp. nov

Valid

Hugot et al.

Triassic

 Brazil

A heteroxynematid oxyurid nematode known from an egg found in a cynodont coprolite. The type species is Paleoxyuris cockburni.

Phragmodictya jinshaensis[360]

Sp. nov

Valid

Xinglian et al.

Cambrian

 China

A hexactinellid sponge, a species of Phragmodictya.

Pirabasoporella[361]

Gen. et 3 sp. nov

Valid

Zágoršek et al.

Miocene

Baitoa Formation
Chipola Formation
Pirabas Formation

 Brazil
 Dominican Republic
 United States

An ascophoran belonging to the family Jaculinidae. The type species is P. atalaiaensis; genus also includes P. baitoae and P. chipolae.

Plicostropheodonta crassicosta[338]

Sp. nov

Valid

García-Alcalde

Devonian (Emsian)

 Spain

A strophodontid brachiopod, a species of Plicostropheodonta.

Plicostropheodonta latronensis[338]

Sp. nov

Valid

García-Alcalde

Devonian (Emsian)

 Spain

A strophodontid brachiopod, a species of Plicostropheodonta.

Polychaetaspis kozlowskii[337]

Sp. nov

Valid

Szaniawski & Drygant

Early Devonian (early Lochkovian)

 Ukraine

A polychaetaspid polychaete, a species of Polychaetaspis.

Postperissocoelia[350]

Gen. et comb. et sp. nov

Valid

Rhebergen & Botting

Silurian

 Canada
 Sweden

A streptosolenid orchocladine demosponge. A new genus for "Perissocoelia" spinosa Rigby & Chatterton (1989); genus also contains new species Postperissocoelia gnisvardensis.

Pseudoperipatus[362]

Gen. et sp. nov

Valid

Haug & Haug

Cambrian

Burgess Shale

 Canada

A cycloneuralian worm. The type species is Pseudoperipatus hintelmannae.

Rogorthis[352]

Gen. et comb. nov

Valid

Hints

Ordovician (Darriwilian)

Kandle Formation
Pakri Formation

 Estonia

A brachiopod belonging to the group Orthida and the family Orthidae. The type species is "Cyrtonotella" pakriensis Rubel (1961).

Sanduscolex[335]

Gen. et sp. nov

Valid

Muir et al.

Ordovician (earliest Floian)

Tonggao Formation

 China

A palaeoscolecid worm. The type species is Sanduscolex regularis.

Sericoidea minuta[349]

Sp. nov

Valid

Zhan et al.

Ordovician (late Katian)

Yinpingshan Formation

 China

A brachiopod.

Spiriferellina fredericksi[363]

Sp. nov

Valid

Tazawa

Permian (Wordian)

 Japan
 Russia

A spiriferellinid spiriferinid brachiopod.

Urphaenomenospongia[350]

Gen. et sp. nov

Valid

Rhebergen & Botting

Silurian (Telychian)

 Sweden

A glass sponge, possibly a member of Lyssacinosida. The type species is Urphaenomenospongia euplectelloides.

Waflascolex[364]

Gen. et sp. nov

Valid

Wang et al.

Ordovician (Tremadocian)

 China

A palaeoscolecid worm. The type species is Waflascolex changdensis.

Wiwaxia foliosa[365]

Sp. nov

Valid

Yang et al.

Cambrian

Hongjingshao Formation

 China

An animal of uncertain phylogenetic placement, a species of Wiwaxia.

Yuganotheca[366]

Gen. et sp. nov

Valid

Zhang, Li & Holmer in Zhang et al.

Cambrian

Heilinpu Formation

 China

A member of Lophophorata of uncertain phylogenetic placement. The type species is Yuganotheca elegans.

Other organisms[edit]

Name Novelty Status Authors Age Unit Location Notes Images

Cyathus dominicanus[367]

Sp. nov

Valid

Poinar

Eocene or Miocene

Dominican amber

 Dominican Republic

A fungus, a species of Cyathus.

Nidula baltica[367]

Sp. nov

Valid

Poinar

Eocene

Baltic amber

 Russia
(Kaliningrad Oblast)

A fungus, a species of Nidula.

Nilpenia[368]

Gen. et sp. nov

Valid

Droser et al.

Precambrian

Ediacara Fossil Site at Nilpena, Flinders Ranges

 Australia

"An organism of unknown affinities with a growth pattern convergent on that of extant fungi, lichens and encrusting algae".[368] The type species is Nilpenia rossi.

Palaeogaster[369]

Gen. et sp. nov

Valid

Poinar, Alfredo & Baseia

Early Cretaceous (late Albian)

 Myanmar

A fungus belonging to the group Sclerodermatineae. The type species is Palaeogaster micromorpha.

Plexus[370]

Gen. et sp. nov

Valid

Joel, Droser & Gehling

Precambrian

 Australia

A tubular, serially divided organism with a bilateral morphology of uncertain phylogenetic placement. The type species is Plexus ricei.

Wutubus[371]

Gen. et sp. nov

Valid

Chen et al.

Precambrian

Dengying Formation

 China

An annulated tubular organism of uncertain phylogenetic placement. The type species is Wutubus annularis.

Xylaria antiqua[372]

Sp. nov

Valid

Poinar

Eocene or Miocene

 Dominican Republic

A fungus belonging to the family Xylariaceae found in Dominican amber, a species of Xylaria.

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