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NASA Aldabra Atoll.jpg
NASA photo of Aldabra Atoll
Location Indian Ocean
Archipelago Seychelles
Area 153.8 km2 (59.4 sq mi)
Population none
Aldabra Atoll is located in Indian Ocean
Aldabra Atoll
Aldabra Atoll
Location of Aldabra Atoll in the Indian Ocean
UNESCO World Heritage Site
Aldabra Atoll
Name as inscribed on the World Heritage List
Aldabra Tortoise Image 002.jpg
Aldabra giant tortoise
Type Natural
Criteria vii, ix, x
Reference 185
UNESCO region Africa
Coordinates 9°25′0.05″S 46°24′59.94″E / 9.4166806°S 46.4166500°E / -9.4166806; 46.4166500 (Aldabra Atoll)Coordinates: 9°25′0.05″S 46°24′59.94″E / 9.4166806°S 46.4166500°E / -9.4166806; 46.4166500 (Aldabra Atoll)
Inscription history
Inscription 1982 (6th Session)

Aldabra is the world's second-largest coral atoll. It is situated in the Aldabra Group of islands in the Indian Ocean that are part of the Outer Islands of the Seychelles.

Uninhabited and extremely isolated, Aldabra is virtually untouched by humans. It has distinctive island fauna including the Aldabra giant tortoise (Aldabrachelys gigantea). It consists of four islands around a large shallow lagoon,[1] encircled by fringing coral reef.[2] The atoll reflects both fossil and geomorphological features, the former is the source of the biodiversity seen today.[3] The atoll has the largest population of giant tortoises in the world (about 100,000 animals).[4] Sir David Attenborough called Aldabra "One of the wonders of the world", and it is also known as one of "crown jewels" of the Indian Ocean.[5][6] Aside from its vast population of tortoises, it is also the largest raised coral reef in the world with an elevation of 26 feet (7.9 m); and the second largest atoll in the world after Kiritimati Atoll. Aldabra has a large population of the world's largest terrestrial arthropod, the coconut crab; and hosts the white-throated rail, the only surviving flightless rail species in the Indian Ocean.[7]

Designated by UNESCO as a World Heritage Site in 1982, it is one of the two UNESCO World Heritage Sites in the Seychelles ;[8] both are administered by the Seychelles Islands Foundation (SIF). BirdLife International declared Aldabra as an Important Endemic Bird Area in 2001 on account of its large seabird colonies. Aldabra became a Ramsar Wetland Site of International Importance in 2010.[5] In 2014 Aldabra was designated as a site under the Indian Ocean South East Asia (IOSEA) turtle network.


Historically, the name Aldabra was Al-Hadra or Al-Khadra (with several variants), given by Arab seafarers[9] for "the atoll’s harsh, sun-baked environment"; this name was included in the Portuguese maps of the 16th century.[10][11] They had named the Indian Ocean as Bahr-el zanj.[12] It was visited by Portuguese navigators in 1511. The islands were already known to the Persians and Arabs, from whom they got their name. In the middle of the 18th century, they became dependencies of the French colony of Réunion, from where expeditions were made for the capture of the giant tortoises.[2] As there are no surface freshwater sources on Aldabra, the interests of the explorers (no proof of any explorer’s visit prior to 1742) was only to exploit the species of tortoise, turtle and fish, and not to inhabit the atoll.[10]

Admiral W.J.L. Wharton of the British navy landed in Aldabra in 1878 to conduct hydrographic surveys of the islands. A decade later the first settlement was established after the Concession was granted by the Seychelles authorities in 1888.[10][13] A small settlement was established on Ile Picard facing west near the beach. The intention was to exploit and export the natural resources of the islands. As part of the settlements, in the middle of the badamier trees, a chapel built with timber and steel was also established as an essential addition to the plantation houses and office buildings. As Aldabra had no water resources, large rectangular-shaped water storage structures were built adjoining each of the houses. A two-roomed jail was also part of this establishment, a remnant of which is still seen at Aldabra. The exploitation of tortoises for commercial purposes at that time is borne out by the remnants of a crushing mill near Picard, which was used to crush bones of tortoises, which were also brought from other islands in the atoll. Efforts made to grow plantation crops of coconuts, cotton, and sisal failed due to inadequate water sources on the atoll; relics of these plantations are still found on some of the islands.[13]

Around the rim of the lagoon are the larger islands of Ile Picard (West Island), Polymnie, Malabar Island (Middle Island) and Grand Terre (South Island). Goats were introduced in the late 19th century as a food source for the few inhabitants (about 200) living there. Ship rats were introduced and recorded before 1870, and house geckos were noted from the 1970s.[10]

In 1810, with Mauritius, Réunion, the Seychelles and other islands, Aldabra passed into the possession of Great Britain. Réunion was later returned to France, and Mauritius gained possession of Aldabra as well as the rest of the Seychelles. The previous inhabitants were emigrants from the Seychelles. Sailors landed on the atoll in the 19th century and attempted to raid the island for tortoises as food; in 1842, two ships were reported to have taken 1200 of them. By 1900, the tortoises were nearly extinct, and a crew would often have to hunt for three days to find one.[14]

In the early 1800s, concessions given to individuals almost destroyed the forests and tortoise habitats in many islands in Seychelles; on Aldabra Atoll, in view of its remoteness and rugged topography, only small areas of forests were cleared for agricultural operations (mostly coconut plantations) but the tortoises were intensely captured for meat and trade.[15] However, James Spurs, who had the concession of the atoll, was responsible initially for saving the tortoises on the atoll when he banned killing them in 1891.[16]

Following World War II, exploitation of Aldabra for commercial use came to an end and restrictions were even imposed on the number of people who could stay on the islands; this number was fixed at 200 at a time. Introduction of invasive species was banned, faunal species were protected under law, and active research on the ecology and biodiversity of the atoll was undertaken by the Royal Society of London from the middle of the 1970s.[13]

Aldabra, along with Desroches and Farquhar, was part of the British Indian Ocean Territory from 1965 until Seychelles' independence in 1976. In the 1960s, as a part of their 'Ocean Island Policy', and to support East of Suez commitments, the British government considered establishing a RAF base on the island and invited the United States to help fund the project in return for shared use of the facility and a settlement of 11 million dollars. Simultaneously (mid-1960s), the British Broadcasting Corporation became interested in Aldabra as a possible site to locate transmitters with which to rebroadcast the BBC Overseas Service (BBC) into the African mainland. The BBC mounted a fact-finding expedition (Expedition Turtle) to assess its suitability for this purpose. The BBC were dependent upon the RAF for developing the atoll as without this their own ambitions would not have been feasible. After an international protest by scientists (known as 'the Aldabra Affair'), however, the military plans were abandoned and the atoll instead received full protection.[14] The "Environmental lobbyists" under the leadership of Julian Huxley, with the support of the MP Tam Dalyell, got the British venture torpedoed.[17][18] In 1966, the Minister of Defence Dennis Healy of the British Government had observed that: "As I understand it, the island of Aldabra is inhabited - like Her Majesty's Opposition Front bench - by giant turtles, frigate birds and boobies."[5]

Subsequent to the thwarting of plans to establish a military station at Aldabra(which instead focussed on Diego Garcia in the Chagos Islands), the Royal Society of London resumed their scientific study of the flora and fauna of the atoll with Professor David Stoddart as the leader. The Royal Society bought the lease of the atoll in 1970 and their research station became functional from 1970. After completion of their assigned work, the Royal Society left and the Seychelles Island Foundation (SIF), a public trust of Seychelles, took over the management and protection of the atoll in 1979.[13][5] SIF functions under the patronage of the President of Seychelles and Aldabra was declared a Special Nature Reserve in 1981, and a year later it became a UNESCO World Heritage Site on 19 November 1982. A brass plaque inscribed with the citation "Aldabra, wonder of nature given to humanity by the people of the Republic of Seychelles" is stationed on the atoll. This appreciation befits the atoll which is truly one of the greatest ecologically undisturbed raised coral atolls in the world.[13]


Aldabra and Assumption Island
Aerial view of Aldabra

Aldabra is a raised coral atoll, [19] located at 9°24′S 46°22′E / 9.400°S 46.367°E / -9.400; 46.367 and belongs to the Aldabra Group, one of the island groups of the Outer Islands of the Seychelles, which includes the island of Assumption and the atolls of Astove and Cosmoledo.[20] At more than 700 miles (1,100 km) from Mahé, the principal island of the Seychelles, it is closer to the coast of Africa, and is in the most southwesterly part of the Seychelles. It is 265 miles (426 km) northwest of Madagascar and a similar distance northeast from the Comoro Islands. The atoll is made of reef limestone (with irregular formations called "champignon"[21]) and this extends over an average width of 2 kilometres (1.2 mi) rising to a height of 8 metres (26 ft) above sea level, and forming the rim line of the shallow central lagoon.[5][22]

Map of Aldabra Atoll

The atoll is the second largest in the world by land area, after Kiritimati. It is 34 kilometres (21 mi) long (in east-west direction) and 14.5 kilometres (9.0 mi) wide. It is up to 8 metres (26 ft) above sea level, and has a land area of 155.4 square kilometres (60.0 sq mi).[23]. The lagoon measures 196 square kilometres (76 sq mi) in area, of which roughly two thirds is dry during low tide. The atoll consists of a ring of four larger islands: counter-clockwise they are South Island (Grand Terre, 116.1 square kilometres (44.8 sq mi)), Malabar or Middle Island (26.8 square kilometres (10.3 sq mi)), Polymnie Island (4.75 square kilometres (1.83 sq mi)) and Picard or West Island (9.4 square kilometres (3.6 sq mi)).[5]

The outside rim of the atoll has three passages which connect to the lagoon, which is 6–10 kilometres (3.7–6.2 mi) in width as it opens to the sea. The depth of water in the lagoon averages about 5 metres (16 ft); however, the passages that open to the sea are up to 20 metres (66 ft) deep and strongly affected by tidal currents. The coral formations are of Pleistocene age and rise to a height of 7.5 metres (25 ft) and are made up of two layers of varying stages of crystallization. They form low cliffs with "deep notches, preceded by jagged pinnacles".[3] Aldabra is formed of cays, beaches, sand dunes, reef limestone islets, and also the remnants of a fringed coral reef.[23] Additionally, there are some 40 smaller islands and rocks, all inside the lagoon, as well as a few very small islets at the West Channels between South and Polymnie Islands, the largest of those being Îlot Magnan. The largest islands inside the lagoon are Île Michel (Michael Island), in the east (0.34 square kilometres (0.13 sq mi)) and Île Esprit (Spirit Island), in the west (0.41 square kilometres (0.16 sq mi)). Geologically the limestone beds have been subjected to striation, sink holes and pits with prominent and continuous limestone bed on the eastern side above the sediment deposits. The coastline has undercut limestone cliffs above a perched beach; it is in two clear terraces of 8 metres (26 ft) and 6 metres (20 ft) height above the sea level.[2][24] Sand dunes dominate the windward south coast.[2]

While the terrestrial topography (spread over an elevation range of 0–8 metres (0–26 ft)[21]) is rugged and dictated by the geomorphic conditions, the land surface comprises limestone of about 125,000 years age, which has uplifted many times above the sea level. The surface conditions are criss-crossed and riddled with pot holes and pits. In the eastern zone of the lagoon, though the surface is continuous, sediment beds are also seen. The windward southern coast is made up of sand dunes.[2] The undercut limestone is of mushroom shape known as champignon.[24]

A small scientific research station of the SIF is based on Picard Island on Aldabra. A small team of staff conduct research to study its biodiversity.

Aldabra is a remote and not easily accessible atoll. No airstrips, helipads or landing jetties have been permitted to be built on the atoll. The nearest airstrip on Assumption Island is about 40 kilometres (25 mi) south-east of Aldabra and cruises are operated by several companies.[25][26] Cruise ships or dive boats may visit the atoll on expedition tours. Supply ships operating from Mahe provide food and other essentials once every few months to the scientists and staff at the research station.[15] Visits to the island by people other than the scientists and staff of the SIF are strictly controlled and only guided tours are provided with prior permission.[25]

Within the atoll, walking paths exist from the settlement of La Gigi which leads to a premonitory from where scenic views of the large lagoon (during low tides) and the mangrove species are seen.[25]

List of smaller islands in the lagoon[edit]

Climate and Tides[edit]

Aldabra is situated in the dry zone of the south-west Indian Ocean. The northwest monsoon is from November to March and brings the heaviest rainfall. In the remaining months, the south-easterly trade winds are recorded. Aldabra receives, on average, 960 millimetres (38 in) rainfall per year.[27] Cyclones are rare in the Seychelles on account of its nearness to the Equator.[27] Tides in the coastal zone rise to more than 3 metres (9.8 ft) height causing channel currents,[2] and a huge influx of water; the main channel drains 60% flow into the lagoon.[5] The reported monthly mean maximum temperature recorded in December is 31 °C (88 °F). The mean minimum temperature recorded in August is 22 °C (72 °F).


Left: coral "mushrooms". Right: nesting male great frigatebird.

The earliest study of the flora and fauna, and also the geomorphological structure is dated to 1910. There are 307 species of animals and plants on Aldabra.[27] Reptiles are the prominent terrestrial fauna.[2]


The higher areas of Aldabra are covered in pemphis, a thick coastal shrub, while the lower areas which are home to the giant tortoises, are a mixture of trees, shrubs, herbs and grasses. There has been recorded 273 species of flowering plants, shrubs, and ferns on the atoll. There are dense thickets of Pemphis acidula, and a mixture of grasses and herbs called "tortoise turf" in many areas.[2][21] This flora includes 19 endemic species and 22 species that are only common to neighbouring islands, and several of these species are on the IUCN Red List. The tropicbird orchid (Angraecum seychellarum) is the national flower of Seychelles and is found in the dry craggy limestone champignon of Aldabra.[28] Other endemic plants such as Pandanus aldabrensis, the Aldabra lily (Lomatophyllum aldebrense) and a sub-species of tropicbird orchid, Angraecum eburneum.[29] The lagoon is bordered by mangrove forests, and has large inland seagrass meadows as well as areas of coral reef and sand flats.[2] The mangroves, which thrive in tidal mudflat areas and saline conditions, are seen on the shores of the lagoon and are integral to the coastal ecosystem. There are seven species of mangrove occurring on Aldabra, three of which are rarely occurring species. These include 'Mangliye blan' or white mangrove (Avicennia marina) which grows to 12 metres (39 ft), 'Mangliye lat' or black mangrove (Bruguiera gymnorrhiza) which grows to 18 metres (59 ft) in a conical shape, 'Mangliye zonn' (Ceriops tagal) which grows to 7 metres (23 ft) with a buttressed trunk, and 'Mangliye rouz' or red mangrove (Rhizophora mucronata) which is the tallest species up to 20 metres (66 ft) in height.[30]


Aldabra tortoises in captivity
Souimanga sunbird (Cinnyris souimanga)

The atoll is home to the world's largest population of giant tortoises, the Aldabra giant tortoise (Aldabrachelys gigantea), numbering some 100,000 individuals.[31][32] Tortoise size varies substantially across the atoll but adult tortoises typically have a carapace length of about 105 centimetres (41 in) and can weigh up to 350 kilograms (770 lb). They are herbivores and feed on a variety of plants, trees and even algae that grows in the freshwater pools. The tortoises mate between February and May, the females then lay their eggs from June to September in areas with suitable soil layers. They lay eggs (the size of golf balls [16]) in a clutch of 3-5 eggs every few years in high density areas and 14-16 eggs in low density areas. The females can lay several clutches in a year and [33] the incubation period is 73–160 days. The small vulnerable juveniles have to survive the predation by coconut crabs, land crabs, rats and birds.[16] In the past giant tortoises have been relocated to many other islands in Seychelles and also to Victoria Botanical Gardens in Mahé.[16]

The atoll is home to the coconut crab (Birgus latro), the world's largest land arthropod, and sharks, manta rays, barracuda and is a breeding ground for the hawksbill sea turtle (Eretmochelys imbricata) and green sea turtle (Chelonia mydas).[6] Aldabra has one of the largest populations of nesting green turtles in the Western Indian Ocean.

During the Pleistocene the dominant land predator was the crocodilian Aldabrachampsus but this became extinct. Three extant species of lizards occur, the skink Cryptoblepharus boutonii and the geckos Phelsuma abbotti and Hemidactylus mercatorius. Pleistocene fossils also indicate the former presence of an Oplurus iguana and other skink and gecko species.[34] There are three endemic species of bat from Aldabra: Paratriaenops pauliani,[35] Chaerephon pusilla and the Aldabra flying fox (Pteropus aldabrensis), as well as the more widely distributed Mauritian tomb bat (Taphozous mauritianus).[36] There are 1,000 species of insects, many of them endemic.[2] Many species of butterflies also flutter around Aldabra.[37]

Endemic birds include the Aldabra drongo (Dicrurus aldabranus), the Aldabran subspecies of the white-throated rail (Dryolimnas cuvieri aldabranus), the last surviving flightless bird of the Indian Ocean region, and the endemic Aldabra fody (Foudia aldabrana),[21] The islands are important breeding grounds for thousands of seabirds, including several species of tern, red-tailed tropicbirds, white-tailed tropicbirds, red-footed boobies, and the world’s second largest breeding populations of great and lesser frigate birds.[2][6][38] The bird fauna is most similar to Madagascar or Comoros and other birds found here include greater flamingos, the Madagascar pond heron, Comoros blue pigeon, Madagascar kestrel, Madagascar coucal, Madagascar nightjar, Madagascar bulbul and souimanga sunbird.[39][40]


Conservationists feared a major threat to the atoll's biodiversity in the 1960s when, as part of the British Indian Ocean Territory, the British made plans to set up a military establishment on the atoll. Due to national and international opposition this plan was cancelled in 1967. This incident became known as the 'Aldabra Affair' in England. Invasive alien species such as rats, cats and goats that were introduced in the past threaten the native biodiversity of the atoll. Goats were eradicated from the atoll in 2012 after a long-term eradication programme. Cats have been removed from all of the islands except Grande Terre, which allowed for the reintroduction of the Aldabra rail to Picard Island. Research into a feasibility study to eradicate rats from the atoll has been undertaken. Aldabra was until recently free of introduced birds but unfortunately the introduced Foudia madagascariensis that was introduced to Assumption Island, now occurs on Aldabra. an eradication programme for this bird on both Assumption and Aldabra is underway. Due to the limited space of its habitat, extreme weather conditions, epidemic and limited range could also pose serious threats to the entire ecology of the atoll.[33]


Aldabra, listed as "Aldabra Atoll", was designated a UNESCO World Heritage Site on 19 November 1982, and is managed and protected by the Seychelles Islands Foundation.[41] The marine protected area extends 1 kilometre (0.62 mi) into the sea to ensure preservation of its marine fauna.[2][21] Eco tourism is controlled and introduction of invasive species is restricted.[2] Based on the evaluation process, UNESCO inscribed the site, a legally protected special reserve of 35,000 hectares (86,000 acres), on the list of World Heritage Sites under three criteria: Criterion (vii) Aldabra Atoll encompasses a large expanse of relatively untouched natural beauty where a number of important animal species and some plant species thrive, along with remarkable land formations, and its process provides a unique spectacle of natural phenomena;[2] Criterion (ix): The atoll is a superlative example of an oceanic island ecosystem in which evolutionary processes are active within a rich biota. The size and morphological diversity of the atoll has permitted the development of a variety of discrete insular communities with a high incidence of endemicity among the constituent species that are typical of island ecosystems. The natural processes take place with minimal human interference and can be clearly demonstrated in their full complexity;[2] and Criterion (x): Aldabra provides a natural laboratory for the study of the process of evolutionary ecology and is a platform for key scientific discovery. The atoll constitutes a refuge harbouring viable populations of a range of rare and endangered species of plants and animals. These include the last giant tortoise and flightless bird populations of the Western Indian Ocean, a substantial marine turtle breeding population, and large seabird colonies which number in the tens of thousands. The substantial tortoise population is self-sustaining and all the elements of its inter-relationship with the terrestrial environment are evident. .[2][42]

BirdLife International declared Aldabra as an Important Bird Area (IBA) in 2001 on account of its large seabird colonies under categories A1, A2, A4i, A4ii and A4iii, covering an area of 33,180 hectares (82,000 acres) overlapping with the special reserve area of 35,000 hectares (86,000 acres) of Aldabra Atoll.[21][43]

Covering 25,100 ha (over half the area of the whole atoll) the wetland ecosystem of Aldabra was listed as a Wetland of International Importance under the Ramsar Convention in 2010. These wetlands at Aldabra include the extensive shallow lagoon inside the atoll, which is carpeted with lush seagrass beds and patchy coral reefs, the intertidal mud flats, the coral reefs outside the lagoon (see next section), freshwater pools, beaches, and 2000 ha of mangrove stands. These wetlands support several endangered species including the increasing number of turtles at the atoll, dugongs and many other bird, fish and invertebrate species.

See also[edit]


  1. ^ Coe 1998, p. 3.
  2. ^ a b c d e f g h i j k l m n o p "Aldabra Atoll". Unesco. Retrieved 23 March 2013. 
  3. ^ a b Dev Raj Khanna; P. R. Yadav (1 January 2005). Biology Of Coelenterata. Discovery Publishing House. pp. 343–. ISBN 978-81-8356-021-4. Retrieved 27 March 2013. 
  4. ^ MairBeckley 2012, p. 189.
  5. ^ a b c d e f g "Aldabra". Official web site of Aldebra Foundation Organization. Retrieved 24 March 2013.  Cite error: Invalid <ref> tag; name "add" defined multiple times with different content (see the help page).
  6. ^ a b c "Aldabra Atoll, Seychelles". Ocean Portal by The Smithsonian Institution. Retrieved 25 March 2013. 
  7. ^ Matthew Murrie (21 September 2010). The First Book of Seconds: 220 of the Most Random, Remarkable, Respectable (and Regrettable) Runners-Up and Their Almost Claim to Fame. Adams Media. pp. 180–. ISBN 978-1-4405-1068-7. Retrieved 10 March 2013. 
  8. ^ MairBeckley 2012, p. 27, 48.
  9. ^ Dodd, Jan (2004). Mauritius, R¿̐ưeunion and Seychelles. Lonely Planet. pp. 268–. ISBN 978-1-74059-301-4. Retrieved 10 April 2013. 
  10. ^ a b c d Anthony Cheke. "Timing of arrival of humans and their commercial animals on the western Indian Ocean oceanic islands" (pdf). Retrieved 27 March 2013. 
  11. ^ "Aldabra Atoll: Aldabra Group". Official website of Tourism department of Sechelles. Retrieved 25 March 2013. 
  12. ^ MairBeckley 2012, p. 11.
  13. ^ a b c d e MairBeckley 2012, p. 186.
  14. ^ a b Carpin, Sarah,(1998) Seychelles, Odyssey Guides, p.162, The Guidebook Company Limited, Retrieved on June 22, 2008
  15. ^ a b MairBeckley 2012, p. 181.
  16. ^ a b c d MairBeckley 2012, p. 35.
  17. ^ MairBeckley 2012, p. 19.
  18. ^ Tam Dalyell (2011). The Importance of Being Awkward: The Autobiography of Tam Dalyell. Birlinn. pp. 124–131. ISBN 9780857900753. Retrieved 25 January 2015. 
  19. ^ SwinglandKlemens 1989, p. 106.
  20. ^ Carpin, Sarah,(1998) Seychelles, Odyssey Guides, p. 161, The Guidebook Company Limited, Retrieved on June 22, 2008
  21. ^ a b c d e f "Aldabra atoll". Bird Life Organization. Retrieved 25 March 2013. 
  22. ^ MairBeckley 2012, pp. 181–.
  23. ^ a b Coe 1998, p. 11.
  24. ^ a b MairBeckley 2012, pp. 6–7.
  25. ^ a b c MairBeckley 2012, p. 187.
  26. ^ "Lonely Planet review for Aldabra Atoll". Retrieved 25 March 2013. 
  27. ^ a b c Coe 1998.
  28. ^ MairBeckley 2012, p. 30.
  29. ^ MairBeckley 2012, p. 31.
  30. ^ MairBeckley 2012, p. 188.
  31. ^ MairBeckley 2012, p. 35, 189.
  32. ^ Payne, Roger (2004-04-05). "Losing Aldabra". Voyage of the Odyssey. PBS. Retrieved 2008-10-21. 
  33. ^ a b Brian Groombridge (1982). Iucn Amphibia-Reptilia Red Data Book. IUCN. pp. 79–. ISBN 978-2-88032-601-2. Retrieved 27 March 2013. 
  34. ^ Arnold, E.N. (1976). "Fossil reptiles from Aldabra Atoll, Indian Ocean". Bull. British Mus. (Nat. Hist.) Zoology 29 (2): 83–116. 
  35. ^ Goodman, S. M.; Ranivo, J. (2008). "A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles)". Zoosystema 30 (3): 681–693. Retrieved 2014-09-13. 
  36. ^ Hutson, A.M. (2004). "The bats of Aldabra atoll, Seychelles". Phelsuma 12: 126–132. 
  37. ^ MairBeckley 2012, p. 33.
  38. ^ MairBeckley 2012, p. 28.
  39. ^ MairBeckley 2012, p. 39.
  40. ^ Seychelles Bird Records Committee, Retrieved on June 26, 2014
  41. ^ Seychelles Islands Foundation, Retrieved on June 22, 2008
  42. ^ "List of World Heritage Sites". Unesco. Retrieved 23 March 2013. 
  43. ^ "Aldabra Island xeric scrub". World Retrieved 25 April 2013. 


External links[edit]