American avocet

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American avocet
American Avocet1.jpg
Breeding plumage
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Charadriiformes
Family: Recurvirostridae
Genus: Recurvirostra
Species:
R. americana
Binomial name
Recurvirostra americana
Gmelin, 1789
Recurvirostra americana map.svg
American avocet adult with chicks, Great Sand Dunes National Park

The American avocet (Recurvirostra americana) is a large wader in the avocet and stilt family, Recurvirostridae. It spends much of its time foraging in shallow water or on mud flats, often sweeping its bill from side to side in water as it seeks its crustacean and insect prey.[2]

Taxonomy and systematics[edit]

The American avocet is a bird in the order Charadriiformes, which includes shorebirds, gulls, and alcids. Its family—Recurvirostridae—includes stilts and avocets. The American avocet is one of four avocet species; the Andean avocet, the pied avocet, and the red-necked avocet are the remaining three species in the genus.[3]

Extant Species[edit]

Common and binomial names Image Range
Black-winged stilt
Himantopus himantopus
Black-winged stilt (Himantopus himantopus).jpg widespread
Hawaiian stilt
Himantopus knudseni
Black-necked Stilt.jpg Hawaii
Pied stilt
Himantopus (himantopus) leucocephalus
1 (83) Pied Stilt (Himantopus leucocephalus).JPG Malesia, Australia
White-backed stilt
Himantopus (himantopus/mexicanus) melanurus
Himantopus melanurus 4.jpg South America
Black-necked stilt
Himantopus mexicanus
Black-necked Stilt (Himantopus mexicanus), Corte Madera.jpg Americas
Black stilt
Himantopus novaezelandiae
Himantopus-novaezelandiae.jpg South Island (New Zealand)
American avocet
Recurvirostra americana
Recurvirostra americana -Palo Alto Baylands-8.jpg North America
Andean avocet
Recurvirostra andina
Recurvirostra andina.jpg Andes
Pied avocet
Recurvirostra avosetta
Pied Avocet Recurvirostra avosetta.jpg widespread
Red-necked avocet
Recurvirostra novaehollandiae
Recurvirostra novaehollandiae in flight - Lake Joondalup.jpg Australia

Description[edit]

The American avocet measures 40–51 cm (16–20 in) in length, has a wingspan of 68–76 cm (27–30 in) and weighs 275–420 g (9.7–14.8 oz)[4][5] The bill is black, pointed, and curved slightly upwards towards the tip. It is long, surpassing twice the length of the avocet's small, rounded head. Like many waders, the avocet has long, slender legs and slightly webbed feet.[6] The legs are a pastel grey-blue, giving it its colloquial name, blue shanks. The plumage is black and white on the back, with white on the underbelly. During the breeding season, the plumage is brassy orange on the head and neck, continuing somewhat down to the breast. After the breeding season, these bright feathers are swapped out for white and grey ones.[7] The avocet commonly preens its feathers - this is considered to be a comfort movement.[2]

Distribution and habitat[edit]

American avocets were previously found across most of the United States until extirpated from the East Coast. The breeding habitat consists of marshes, beaches, prairie ponds, and shallow lakes in the mid-west, as far north as southern Canada.[4] These breeding grounds are largely in areas just east of the rocky mountains including parts of Alberta, Saskatchewan, Montana, Idaho, Washington, Utah, North and South Dakota, Nebraska, Colorado, and even down to parts of New Mexico, Oklahoma and Texas.[7] Their migration route lands them in almost every state in the western United States. The avocet's wintering grounds are mainly coastal. Along the Atlantic Ocean, they are found in North and South Carolina, Georgia and Florida. There are also wintering grounds along the Gulf of Mexico in Florida, Texas, and Mexico, and along the Pacific Ocean in California and Mexico. There are resident populations in the Mexican States of Zacatecas, San Luis Potosí, Guanajuato, Hidalgo, Mexico City and Puebla, and in Central California.[7]

Behavior and ecology[edit]

Vocalizations[edit]

The American avocet call has been described as both a shrill and melodic alarm bweet, which rises in inflection over time.[6][7][8] Avocets use three distinct calls: common call, excited call, and broken wing call.[9] The common call is a loud repeated wheep. The excited call has a similar wheep sound, but it speeds up rather than having an even rhythm. Lastly, the broken wing call is noticeably different from the other two calls. It is a distressed screech sound and is alarming rather than melodic.[9]

Migration and breeding[edit]

Flocks ranging from 50 to 300 avocets migrate together to their breeding sites together during peak season, which is typically in the spring.[10] The American avocet breeds in anything from freshwater to hypersaline wetlands in the western and mid-west United States.[7][11] After reaching their breeding grounds, the avocets quickly establish territory in pairs.[10] Throughout the breeding season, these pairs engage in a series of copulatory displays, which is initiated by either sex and often involves preening.[7][10] After breeding, the birds gather in large flocks, sometimes amounting to hundreds of birds. Nesting takes place near water, often alongside black-necked stilts, usually on small islands or mucky shorelines. The avocets nest on the ground in places with little to no surrounding vegetation. They make shallow indentations in the ground for their nest and line it with grass, feathers, pebbles or other small objects. Some avocets do not line their nests at all.[4] During the breeding season, avocets lay around 3–5 eggs, with 4 being the most common number.[9] One case study of the 1968 and 1969 breeding seasons suggests that this number allows for the highest rate of hatching success.[10] They incubate them over a period of 23–25 days. Both parents are involved in the incubation process, taking turns during the day (except at night, where only the female incubates the eggs).[10][9]

Growth and development[edit]

Both parents take care of the newborn American avocets after they hatch. Male parents are more present in incubation during the first 8 days, whereas female parents dominate incubation throughout the remaining 16 days.[12] Time spent caring for newborn chicks equates to that spent incubating. Since newborns are able to walk, they are capable of feeding themselves and do not require extensive care from their parents.[9][12] The young are expected to begin flying 4 to 5 weeks after hatching.[10]

Anti-Predator Behavior[edit]

Each pair defends their own territory against predators and other avocets. Nests are built in areas that predators would struggle to gain access to. Predators that do try to approach the colony find themselves surrounded by several adult avocets running around, trying to distract them. While incubating however, the avocets tend only to their nest and defense is considered a lower priority, unless the eggs are in immediate danger, in which case the avocets may take more aggressive measures, calling loudly and flying at the threat.[10][9]

Food and feeding[edit]

The American avocet tends to prefer habitats with fine sediments for foraging.[13] In the winter, it feeds extensively on brine shrimp. Usually, this entails the avocet pecking while walking or wading on the shore, but it can also swim to expand foraging area. During the breeding season, avocets continue to eat brine shrimp but switch mainly to consuming brine flies. Brine flies and brine fly larvae are so abundant in avocet breeding grounds that they often blacken the surface of any exposed mud. Brine flies sustain the avocet during its breeding season.[2] The avocet employs both visual and tactile methods of feeding.[7] The primary visual feeding method is pecking at flies. However the avocet will also feed by plunging (wherein the head and neck of the bird are submerged), snatching (wherein the bird catches a flying insect), and bill pursuit (wherein the bird repeatedly opens and closes its bill while moving its head along the surface of the water). Methods of tactile feeding include filtering, scraping, and scythe feeding.[2]

Threats and conservation status[edit]

Shooting and trapping of American avocets led to population decline until the 1900s. During this time, the species was extirpated from most of the East Coast of the United States.[7] By 1918, Avocets became protected under the Migratory Bird Treaty Act (16 U.S.C. 703-712). Since that time, other threats have emerged. Contaminants and toxins such as DDT, selenium, and methylmercury, have increased in American Avocet habitats in recent years. Although the effects are highly contested, these increased concentrations have potential negative effects on American avocet breeding success, especially because Avocet chicks are more susceptible to environmental disruptions than adults.[7][11] The avocet also faces habitat loss. While agricultural and industrial environments have become alternate habitat for the avocet, natural wetlands are decreasing rapidly.[7] Additionally, increases in the salinity levels of wetlands have had adverse effects on the breeding and development of avocets. Young avocets in saline or hypersaline environments often combat weight loss and dehydration.[11] Although American avocets face numerous environmental challenges, numbers have stayed steady, and according to the National Audubon Society, numbers have been increasing steadily in the eastern United States for the past few years.[9]

John James Audubon's depiction of the American avocet in breeding plumage.

In culture[edit]

Art[edit]

In his famous The Birds of America, John James Audubon describes a day of stalking and spying on the avocet. He judiciously noted their foraging, nesting, defensive, and flight behaviors.[6]

Protected status[edit]

The American avocet is protected under the Migratory Bird Treaty Act of 1918.[14]

Gallery[edit]

References[edit]

  1. ^ BirdLife International (2016). "Recurvirostra americana". IUCN Red List of Threatened Species. 2016: e.T22693717A93418724. doi:10.2305/IUCN.UK.2016-3.RLTS.T22693717A93418724.en. Downloaded on 17 February 2018.
  2. ^ a b c d Hamilton, Robert Bruce (1975). "Comparative Behavior of the American Avocet and the Black-Necked Stilt (Recurvirostridae)" (PDF). Ornithological Monographs: iii-98. JSTOR 40166701.
  3. ^ "Recurvirostra Linnaeus, 1758". the Integrated Taxonomic Information System on-line database. Retrieved 2017-10-17.
  4. ^ a b c "American Avocet, Identification". allaboutbirds.org. The Cornell Lab of Ornithology, Cornell University.
  5. ^ "American Avocet". seaworld.org. SeaWorld. Archived from the original on 2013-07-31.
  6. ^ a b c Audubon, John James (1861). The birds of America : from drawings made in the United States and their territories. 6. New York. pp. 24–30.
  7. ^ a b c d e f g h i j Ackerman, Joshua; Hartman, Alex; Herzog, Mark; Takekawa, John; Robinson, Julie; Oring, Lewis; Skorupa, Joseph; Boettcher, Ruth (2013). "American Avocet (Recurvirostra americana)". The Birds of North America. Ithaca, New York, USA: Cornell Lab of Ornithology. doi:10.2173/bna.275. Retrieved 2017-10-17.
  8. ^ "Plovers, Oystercatchers, and Stilts". Larkwire. 2017. Retrieved 2017-10-17.
  9. ^ a b c d e f g "American Avocet". Audubon. 2014-11-13. Retrieved 2021-03-14.
  10. ^ a b c d e f g Gibson, Flash (1971). "The Breeding Biology of the American Avocet (Recurvirostra americana) in Central Oregon". The Condor. 73 (4): 444–454. doi:10.2307/1366666. ISSN 0010-5422.
  11. ^ a b c Hannam, Kristina M.; Oring, Lewis W.; Herzog, Mark P. (2003). "Impacts of Salinity on Growth and Behavior of American Avocet Chicks". Waterbirds: The International Journal of Waterbird Biology. 26: 119–125. doi:10.1675/1524-4695(2003)026[0119:iosoga]2.0.co;2. JSTOR 1522475.
  12. ^ a b Gibson, Flash (1978). "Ecological Aspects of the Time Budget of the American Avocet". The American Midland Naturalist. 99 (1): 65–82. doi:10.2307/2424934. ISSN 0003-0031.
  13. ^ Danufsky, Tamar; Colwell, Mark A. (2003). "Winter Shorebird Communities and Tidal Flat Characteristics at Humboldt Bay, California". The Condor. 105: 117–129. doi:10.1650/0010-5422(2003)105[117:wscatf]2.0.co;2. JSTOR 1370610.
  14. ^ Migratory Bird Treaty
  • O'Brien, Michael, et al. (2006). The Shorebird Guide. New York: Houghton Mifflin. ISBN 0-618-43294-9

External links[edit]