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Temporal range: Middle Permian, 270–260 Ma
Life restoration of Titanophoneus
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Order: Therapsida
Suborder: Dinocephalia
Clade: Anteosauria
Family: Anteosauridae
Boonstra, 1954

Anteosauridae is a family of large carnivorous dinocephalian therapsids that are known from the Middle Permian of Asia, Africa, and South America.These animals were by far the largest predators of the Permian period, with skulls reaching 80 cm in length in adult individuals, far larger than the biggest gorgonopsian.


Life restoration of Archaeosyodon

Anteosaurids are characterized by very large pointed incisors and canines, bulbous spatulate (spoon shaped) postcanines, a very strongly upturned margin of the premaxilla, so the front of mouth curves strongly upwards, and a long, very robust lower jaw (Boonstra 1963).

Anteosaurids distinguished from Brithopus and other non-anteosaurid anteosaurians by the presence of a large thickened region or "boss" on the side of the angle of the lower jaw; this was probably used in intraspecific behaviour. In Doliosauriscus and Anteosaurus, not only was this boss very prominent, but the bones were very thick and rugose. The same situation is found in the herbivorous Tapinocephalidae, and it has been proposed that these animals engaged in head-butting behaviour. (Hopson & Barghusen 1986)


The subfamily Anteosaurinae has also been used in the past to include these therapsids. The Russian genera were previously included by Efremov in the family Brithopodidae. Anteosauridae is part of a larger group of dinocephalians called Anteosauria. Several recent phylogenetic studies of anteosaurians find support for two subfamilies within Anteosauridae: Anteosaurinae and Syodontinae.[1] Below is a cladogram showing syodontine relationships from a 2012 phylogenetic study of anteosaurians:[2]


Biarmosuchus tener


Estemmenosuchus uralensis

Ulemosaurus svijagensis

Tapinocaninus pamelae


Archaeosyodon praeventor


Sinophoneus yumenensis

Titanophoneus adamanteus

Titanophoneus potens

Anteosaurus magnificus


Pampaphoneus biccai

Notosyodon gusevi

Syodon biarmicum

Australosyodon nyaphuli


  1. ^ Kammerer, C.F. (2011). "Systematics of the Anteosauria (Therapsida: Dinocephalia)". Journal of Systematic Palaeontology. 9 (2): 261–304. doi:10.1080/14772019.2010.492645. 
  2. ^ Cisneros, J.C.; Abdala, F.; Atayman-Güven, S.; Rubidge, B.S.; Şengör, A.M.C.; Schultz, C.L. (2012). "Carnivorous dinocephalian from the Middle Permian of Brazil and tetrapod dispersal in Pangaea". Proceedings of the National Academy of Sciences of the United States of America. 109 (5): 1584–1588. doi:10.1073/pnas.1115975109. PMC 3277192Freely accessible. PMID 22307615. 

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