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Starr 080117-1775 Arachniodes simplicior.jpg
Arachniodes simplicior
Scientific classification e
Kingdom: Plantae
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Polypodiales
Suborder: Polypodiineae
Family: Dryopteridaceae
Subfamily: Dryopteridoideae
Genus: Arachniodes
  • Byrsopteris C.V.Morton
  • Leptorumohra (H.Ito) H.Ito
  • Lithostegia Ching
  • Phanerophlebiopsis Ching

Arachniodes is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I).[1] A number of species in this genus are known as "holly ferns".[2]

The genus Arachniodes was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species Arachniodes aspidioides.[3] The genus was not widely recognized until Mary Douglas Tindale transferred the two species (Byrsopteris amabilis and Byrsopteris aristata) into it in 1961.[4]


The Pteridophyte Phylogeny Group classification of 2016 (PPG I) considers Arachniodes to have 60 species.[1] As of January 2020, the Checklist of Ferns and Lycophytes of the World accepted 78 species:[5]

Some hybrids were also accepted:[5]


  1. ^ a b c PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229.
  2. ^ Arachniodes Flora of North America 26 Jan 2012
  3. ^ Blume, Carl (Karl) Ludwig von; Enumeratio Plantarum Javae fasc. 2: 241. 1828.
  4. ^ Tindale, Mary Douglas; Contributions from the New South Wales National Herbarium 3(1): 90. 1961.
  5. ^ a b Hassler, Michael & Schmitt, Bernd (January 2020). "Arachniodes". Checklist of Ferns and Lycophytes of the World. 8.20. Retrieved 2020-01-17. CS1 maint: discouraged parameter (link)

External links[edit]