Benthosuchus

From Wikipedia, the free encyclopedia
  (Redirected from Benthosuchidae)
Jump to: navigation, search
Benthosuchus
Temporal range: Early Triassic
Benthosuchus.jpg
Skull of Benthosuchus sushkini
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Order: Temnospondyli
Suborder: Stereospondyli
Family: Benthosuchidae
Efremov, 1937
Genus: Benthosuchus
Efremov, 1937
Species
  • B. sushkini (Efremov, 1929 [originally Benthosaurus sushkini]) (type)
  • B. korobkovi Ivachnenko, 1972
  • B. bashkiricus Otschev, 1972
  • B. gusevae Novikov, 2012
Life restoration of Benthosuchus sushkini
Skull

Benthosuchus is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. The genus was named in 1937, and the type species, Benthosuchus sushkini, was named in 1940. Benthosuchus has traditionally been considered a member of the temnospondyl superfamily Trematosauroidea, and the family Benthosuchidae was established in 1940 to include Benthosuchus and the related trematosauroid Thoosuchus. Some recent phylogenetic studies have removed Benthosuchus from Trematosauroidea entirely, placing it as a closer relative of Mastodonsauroidea, another group of Triassic temnospondyls. Other studies retain Benthosuchus within Trematosauroidea, but since Benthosuchus and Thoosuchus have successively basal positions on these trees they form a paraphyletic grouping, not a valid clade.[1] In either case, Benthosuchidae is a monotypic family containing only Benthosuchus.

Description[edit]

Benthosuchus skull is about 0.75 m long (it is comparatively short in young individuals but lengthens with age), and the overall body length is about 2.5 m. It existed in widely overflowing rivers of the East Russian Depression.[2]

Phylogeny[edit]

Benthosuchus is traditionally considered a close relative of a group of temnospondyls called capitosaurids. In particular, it bears a close resemblance to the capitosaurid Wetlugasaurus. Many of the early large-scale phylogenetic analyses of temnospondyls place Benthosuchus within a clade called Capitosauria. However, more recent analyses such as that of Fortuny et al. (2011) recover Benthosuchus within another clade called Trematosauria. Occasionally it is grouped near the trematosaurid family Trematosauridae as a basal member of the group Trematosauroidea.[3] Fortuny et al. found that Benthosuchus positioned outside Trematosauroidea as a more basal trematosaurian. Below is a cladogram from Fortuny et al. (2011) showing the phylogenetic placement of Benthosuchus:[4]

Stereospondyli

Lydekkerina huxleyi


Rhinesuchidae

Rhineceps nyasaensis



Uranocentrodon senekalensis




Capitosauria

Wetlugasaurus angustifrons





Odenwaldia heidelbergensis



Vladlenosaurus alexeyevi







Edingerella madagascariensis



Watsonisuchus spp.





Xenotosuchus africanus




Cherninia denwai




Paracyclotosaurus crookshanki





Stanocephalosaurus pronus



Stanocephalosaurus birdi





Procyclotosaurus stantonensis




Eocyclotosaurus spp.



Quasicyclotosaurus campi











Parotosuchus orenburgensis




Calmasuchus acri





Cyclotosaurus robustus



Tatrasuchus wildi





Eryosuchus garjainovi



Mastodonsaurus giganteus









Trematosauria

Benthosuchus sushkini


Trematosauroidea

Thoosuchus yakovlevi




Angusaurus spp.



Trematosaurus brauni







References[edit]

  1. ^ Damiani, R.J.; Yates, A.M. (2003). "The Triassic amphibian Thoosuchus yakovlevi and the relationships of the Trematosauroidea (Temnospondyli: Stereospondyli)" (PDF). Records of the Australian Museum 55: 331–342. doi:10.3853/j.0067-1975.55.2003.1388. 
  2. ^ http://encyclopedia2.thefreedictionary.com/Benthosuchus
  3. ^ Shishkin, M.A.; Sulej, T. (2009). "Early Triassic temnospondyls of the Czatkowice 1 tetrapod assemblage" (PDF). Palaeontologica Polonica 65: 31–77. 
  4. ^ Fortuny, J.; Galobart, À.; Santisteban, C. D. (2011). "A New Capitosaur from the Middle Triassic of Spain and the Relationships within the Capitosauria". Acta Palaeontologica Polonica 56 (3): 553. doi:10.4202/app.2010.0025.  edit