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Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Superfamily: Scarabaeoidea
Family: Bolboceratidae
Mulsant, 1842


Bolboceratidae, the dor beetles, is a family of beetle.[1] It was historically treated as a subfamily of the earth-boring dung beetles (family Geotrupidae), but has been considered a separate family since 1995.[2] Some recent classifications have persisted in treating bolboceratids as a subfamily (e.g.[3][4]) but these classifications are contradicted by recent phylogenetic studies of relationships indicating that bolboceratids are not closely related to geotrupids (e.g., that bolboceratids are more closely related to Pleocomidae and Passalidae[5]).



  1. ^ Gomes Gonçalves, Marcos Paulo (December 2017). "Relationship Between Meteorological Conditions and Beetles in Mata de Cocal". Revista Brasileira de Meteorologia. 32 (4): 543–554. doi:10.1590/0102-7786324003. ISSN 0102-7786.
  2. ^ Bolboceratidae at TOL
  3. ^ Smith, A.B.T. (2006) A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification. The Coleopterists Bulletin, 60:144-204.
  4. ^ Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Ślipiński, A.; Smith, A. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys. 88 (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
  5. ^ MCKENNA, D. D., WILD, A. L., KANDA, K., BELLAMY, C. L., BEUTEL, R. G., CATERINO, M. S., FARNUM, C. W., HAWKS, D. C., IVIE, M. A., JAMESON, M. L., LESCHEN, R. A. B., MARVALDI, A. E., MCHUGH, J. V., NEWTON, A. F., ROBERTSON, J. A., THAYER, M. K., WHITING, M. F., LAWRENCE, J. F., ŚLIPIŃSKI, A., MADDISON, D. R. and FARRELL, B. D. (2015), The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Syst Entomol, 40: 835–880. doi:10.1111/syen.12132

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