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Temporal range: Tremadocian to Pragian488.3–409.1 Ma
Calymene clavicula dorsal.jpg
Calymene clavicula 32mm, Henryhouse Formation, Oklahoma, Cayugan
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita
Order: Phacopida
Family: Calymenidae
Genus: Calymene
Münster, 1840
Type species
Calymene blumenbachii
Brongniart in Desmarest, 1817 [1]

Calymene (meaning beautiful crescent as a reference to the glabella) is a genus of trilobites in the order Phacopida that are found throughout North America, North Africa, and Europe in primarily Silurian outcrops.[2] Calymene is closely related to Flexicalymene, and both genera are frequently found inrolled.[3] Calymene trilobites are small, typically 2 cm in length. Their cephalon is the widest part of the animal, and the thorax is usually in 13 segments.[4]

Known species and locations[edit]

Reassigned species[edit]

Since the genus Calymene was established early on in paleontology, a number of species previously assigned to it have since been transferred to other genera:[7]


  1. ^ a b Derek J. Siveter (1985). "The type species of Calymene (Trilobita) from the Silurian of Dudley, England" (PDF). Palaeontology. 28 (4): 783–792. 
  2. ^ "†Calymene Muenster 1840 (trilobite)". The Paleontology Database. Retrieved April 29, 2012. 
  3. ^ John P. Rafferty, ed. (2010). The Paleozoic Era: Diversification of Plant and Animal Life. Geologic History of Earth. Britannica Educational Publishing. ISBN 978-1-61530-196-6. 
  4. ^ Milson, C.; Rigby, S. (2004). Fossils at a Glance. Oxford: Blackwell Science Ltd. 
  5. ^ "Calymene celebra Raymond 1916 (trilobite)". Paleobiology Database. Retrieved 20 August 2011. 
  6. ^ Alex J. Chestnut. "Using morphometrics, phylogenetic systematics and parsimony analysis to gain insight into the evolutionary affinities of the Calymenidae Trilobita". OhioLINK ETD Center. Retrieved August 21, 2011. 
  7. ^ Moore, R.C. (1959). Arthropoda I - Arthropoda General Features, Proarthropoda, Euarthropoda General Features, Trilobitomorpha. Treatise on Invertebrate Paleontology. Part O. Boulder, Colorado/Lawrence, Kansas: Geological Society of America/University of Kansas Press. pp. 1–560. ISBN 0-8137-3015-5. 
  8. ^ a b Esteve, Jorge (2015). "Systematic revision of the genus Solenopleura Angelin, 1854, Ptychopariida, Trilobita, Cambrian Series 3". Annales de Paléontologie. 101: 185–192. doi:10.1016/j.annpal.2015.05.001. 
  9. ^ Pärnaste, Helje (2004). "Revision of the Ordovician cheirurid trilobite genus Reraspis with the description of the earliest representative" (PDF). Proc. Estonian Acad. Sci. Geol. 53 (2): 125–138. 
  10. ^ Ramskjöld, L.; Werdelin, L. (1991). "The phylogeny and evolution of some phacopid trilobites". Cladistics. 7: 29–74. doi:10.1111/j.1096-0031.1991.tb00021.x. 

External links[edit]