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Scientific classification
Domain: Eukaryota
(unranked): Sar
Superphylum: Alveolata
Phylum: Apicomplexa
Levine 1970
Class: Conoidasida
Levine 1988
Subclass: Gregarinasina
Dufour 1828
Order: Neogregarinorida
Grassé & Schrével 1953
Suborder: Schizogregarinina
Family: Caulleryellidae
Keilin 1914


The Caulleryellidae are a family of parasitic protozoa in the phylum Apicomplexa. Species in this family mostly infect dipteran larvae.


This family was created by the parasitologist David Keilin in 1914.[1]


Two genera are placed in this family - Caulleryella with five species and Tipulocystis with one. The type species is Caulleryellida aphiochaetae.


Species in the family are generally spread by the oral-faecal route. Replication occurs in the gut. Development is extracellular. The parasites develop attached to the host cell by an epimerite. Merogony occurs when nuclei are peripheral within the cell. The merozoites form a bouquet structure around the parent cell.

The gamonts are elongated with an anterior neck. The gametes have similar morphology. The gametocysts give rise to one to eight oocysts.


  1. ^ Keilin D (1914) CR Soc Biol Paris 76 (1) 768