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Octoglena sierra (Family Polyzoniidae) (3405605943).jpg
Octoglena sierra (Polyzoniida)
Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Class: Diplopoda
Infraclass: Helminthomorpha
Clade: Colobognatha
Brandt, 1834


Colobognatha is a clade (formally considered a subterclass) of helminthomorph millipedes containing four orders: Platydesmida, Polyzoniida, Siphonocryptida, and Siphonophorida.[1]


The Colobognatha are united by several shared traits (synapomorphies). Males have two pairs of simple, leg-like gonopods, consisting of the rear leg pair of body segment 7 and the foreword-most leg pair of segment 8. They possess tubular defensive glands that open on the sides of the body, and lack a palp-like structure on their gnathochilaria. Other traits found in colobognathans, but not exclusively, include a distinctly narrow head, lack of Tömösváry organs, and no more than two pairs of ocelli.[2][3]


  1. ^ Shear, W. (2011). "Class Diplopoda de Blainville in Gervais, 1844. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness" (PDF). Zootaxa. 3148: 159–164. 
  2. ^ "Millipede Body Organization". Milli-PEET: The class Diplopoda. The Field Museum, Chicago. Retrieved 18 March 2014. 
  3. ^ "Putative apomorphies of millipede clades" (PDF). Milli-PEET: Millipede Systematics. The Field Museum, Chicago, IL. 26 September 2006.