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Anodonthyla boulengerii.jpg
Anodonthyla boulengerii
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Amphibia
Order: Anura
Family: Microhylidae
Subfamily: Cophylinae
Cope, 1889
Type genus
Boettger, 1880

8 genera; see article.

Cophylinae is a subfamily of microhylid frogs endemic to Madagascar. It has over 70 species in eight genera.[1][2]


As of February 2017, the following genera are recognized:[1][2]


Cophylines are characterized by a derived mode of larval development: whereas most microhylids have a specialized filter-feeding tadpole, cophylines have non-feeding tadpoles that develop either in tree holes, terrestrial foam nests, or terrestrial jelly nests.[3] Most cophylines have very simple advertisement calls, consisting of single melodious notes that are repeated after regular intervals and for long periods of time, usually lasting several minutes. Correlated to the reproductive mode of the various cophyline lineages is their arboreal versus terrestrial or fossorial ecology, and apparently, multiple evolutionary shifts between arboreal and terrestrial habits have occurred in this subfamily.[4]


  1. ^ a b Frost, Darrel R. (2016). "Cophylinae Cope, 1889". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 10 May 2016. 
  2. ^ a b Scherz, Mark D.; Vences, Miguel; Rakotoarison, Andolalao; Andreone, Franco; Köhler, Jörn; Glaw, Frank; Crottini, Angelica (2016). "Reconciling molecular phylogeny, morphological divergence and classification of Madagascan narrow-mouthed frogs (Amphibia: Microhylidae)". Molecular Phylogenetics and Evolution. 100: 372–381. PMID 27085671. doi:10.1016/j.ympev.2016.04.019. 
  3. ^ Blommers-Schlösser RMA (1975). "Observations on the larval development of some Malagasy frogs, with notes on their ecology and biology (Anura: Dyscophinae, Scaphiophryninae and Cophylinae)". Beaufortia. 24: 7–26. 
  4. ^ Andreone F; Vences M; Vieites DR; Glaw F; Meyer M (2005). "Recurrent ecological adaptations revealed through a molecular analysis of the secretive cophyline frogs of Madagascar". Molecular Phylogenetics and Evolution. 34: 315–322. PMID 15619444. doi:10.1016/j.ympev.2004.10.013. 


  • Vences M., Glaw F., Köhler J. & Wollenberg K. Molecular phylogeny, morphology and bioacoustics reveal five additional species of arboreal microhylid frogs of the genus Anodonthyla from Madagascar. Contributions to Zoology 2010; 79(1): 1-32. full text released under license CC-by