|Chambal River, Uttar Pradesh, India|
|Distribution of Crocodylus palustris|
The mugger crocodile (Crocodylus palustris = "crocodile of the marsh"), also called the Indian, Indus, Persian, Sindhu, marsh crocodile or simply mugger, is found throughout the Indian subcontinent. It is one of the three crocodilians found in India, the others being the gharial and the saltwater crocodile. It is a medium-sized crocodile that mostly inhabits freshwater lakes, ponds, sluggish rivers, swamps and marshes. Males of the species are said to grow up to 4–5 m (13–16 ft) in length. As with other crocodilians, females are smaller. The mugger crocodile has the broadest snout of any extant crocodile, giving it an alligator-like appearance. It is a more heavily armored species with enlarged scutes around the neck. Adults are dark grey or brown, while hatchlings are tan colored.
The mugger crocodile is a skilled predator that preys on a variety of species. Like other crocodilians they are ambush hunters and wait for their prey to come close. They wait camouflaged in the murky waters to launch the attack in the suitable moment. They mostly prey on fish, reptiles, birds and mammals. Reproduction takes place in winter months. Females lay eggs in nests that are holes dug in the sand. Temperature during incubation is the determinant of sex in the young. The mugger crocodile possesses the size to be a serious threat to humans, but they are not as aggressive as some other species, such as the sympatric saltwater crocodiles. They are also observed to usually avoid areas with saltwater crocodiles. Muggers are a fairly social species and tolerate each other during basking and feeding.
Taxonomy and etymology
The name "mugger" is a borrowing of magar, the Hindi word for "crocodile". This is in turn derived from makara, a Sanskrit name for a mythical water monster that is sometimes depicted as a crocodile.
Mugger crocodiles have 19 upper teeth on each side; a snout that is 1⅓ to 1½ as long as broad at the base; a rough head but without any ridges; mandibular symphysis extending to the level of the fourth or fifth tooth; pre-maxillo-maxillary suture, on the palate, transverse, nearly straight, or curved forwards; and nasal bones separating the pnemaxillaries above. Four large nuchals forming a square, with a smaller one on each side; two pairs of smaller nuchals on a transverse series behind the occiput. Dorsal shield well separated from the nuchal, the scutes usually in 4, rarely in 6, longitudinal series, those of the two median usually considerably broader than long; 16 or 17 transverse series. Scales on limbs keeled. Fingers webbed at the base; outer toes extensively webbed. A serrated fringe on the outer edge of the leg. Adult blackish olive above: young pale olive, dotted and spotted with black.
The mugger crocodiles is considered a medium-sized species. Although much smaller than a saltwater crocodile on average, the mugger crocodile has the broadest snout of any extant true crocodile and may appear bulkier than many crocodiles of around the same length. Sexual maturity is obtained at around 1.7–2 m (5.6–6.6 ft) and 2.6 m (8 ft 6 in) for females and males respectively. The largest specimen in the British Museum measures 3.7 m (12 ft 2 in). However, some mugger crocodiles are said to grow up to 4–5 m (13 ft 1 in–16 ft 5 in) long in exceptional cases. The average size of adult mugger crocodiles in Manghopir, Pakistan was cited as 2.89 m (9 ft 6 in), with an estimated average weight of around 100 kg (220 lb). However, some muggers of around 3 m (9 ft 10 in) may weigh 195 kg (430 lb), as was one male caught by Pakistani Forestry, Wildlife and Fisheries department. An 5 m (16 ft 5 in) outsized mugger crocodile may scale up to 700 kg (1,500 lb).
Distribution and habitat
Mugger crocodiles are a freshwater species found in lakes, rivers and marshes. Muggers prefer slow-moving, shallower bodies of water rather than fast-flowing, deep areas. They are also known to thrive in man-made reservoirs and irrigation canals. Although they prefer freshwater, they have some tolerance to saltwater therefore are occasionally found in saltwater lagoons. The species is sympatric with the gharial (Gavialis gangeticus) in some areas of India and Nepal and with the saltwater crocodile (Crocodylus porosus) in other areas, but separated by habitat most of the time.
Muggers are better adapted to terrestrial life than most other crocodilians, like their cousin the Cuban crocodile, but they are ecologically most similar to the African Nile crocodile. Muggers are known to be more mobile on land, can migrate considerable distances over land in search of a more suitable habitat, and can chase prey on land for short distances. They are also known to dig burrows as shelters during the dry seasons.
The mugger crocodile can be found in India, Bangladesh, Sri Lanka, Pakistan, Nepal, and the southern tip of Iran. They are probably also found in Indo-China. This crocodile is the most common and widespread of the three species of crocodiles in India, far outnumbering the much larger saltwater crocodile within the country (and most likely within neighboring countries). The mugger is the only crocodilian found in Iran and Pakistan.
In the 1980s, the largest population of wild crocodiles in Tamil Nadu, South India lived in the Amaravathi Reservoir, and in the Chinnar, Thennar and Pambar rivers that drain into it. Their total population was estimated to be 60 adults and 37 sub-adults.
The Amaravati Sagar Crocodile Farm, established in 1975, is the largest crocodile nursery in India. Eggs are collected from wild nests along the perimeter of the reservoir to be hatched and reared at the farm. There were up to 430 animals maintained in captivity at one time. Hundreds of adult crocodiles have been reintroduced from there into the wild. The estimated population in Pakistan is between 400 and 450 animals found in the coastal areas and rivers of Sindh and Baluchistan provinces. They are indigenous to the Sarbaz River, Bahu Kalat River, Kaju and Pishin river basins. The mugger crocodile is the national reptile of Pakistan.
On the Iranian Makran coast, above Chabahar, there is a population of around 200 mugger corocodiles. Due to human activity and a long drought in the late 1990s and early 2000s, the mugger had been pushed to the brink of extinction over the past few years. Following several tropical cyclones, such as the Cyclone Gonu and Cyclone Yamyin in 2007, and Cyclone Phet in 2010, much of the habitat of the Iranian mugger crocodiles has been restored as the dry lakes and hamuns have flooded once again. The animal is known as gando in the local Iranian vernaculars.
Biology and behavior
Mugger crocodiles can achieve speed of around 8 mph over a short distance in pursuit of prey. They can swim much faster than they can run—achieving speeds of 10 to 12 mph in short bursts—and can cruise at about 1 to 2 mph.
Hunting and diet
Being a large carnivorous reptile, the mugger crocodile eats fish, snakes, turtles, birds and mammals. The mammalian prey are usually small to medium-sized, such as monkeys, squirrels, chital and otters. In fact, most vertebrates that approach to drink are potential prey, and may suffer being seized and dragged into the water to be drowned and devoured when the opportunity arises. Large adults will sometimes prey on larger mammals, such as large deer, like sambar, and large bovines, like cattle and domestic water buffalo, but such large prey are not common prey as in the case of sympatric saltwater crocodiles. At night, they sometimes hunt on land, lying in ambush near forest trails. During these nocturnal hunts, they often steal kills from other predators, such as leopards, sometimes preying on the predators as well. This species is generally considered to be occasionally dangerous to humans.
Mugger crocodiles have been documented using lures to hunt prey such as birds. This means they are among the first reptiles recorded to use tools. By balancing sticks and branches on their heads, Mugger crocodiles are able to lure birds looking for suitable nesting material. This strategy, which is shared by the American alligator, is particularly effective during the nesting season, in which birds are more likely to gather appropriate nesting materials.
- The Asiatic lion, which inhabits Gir Forest National Park in Gujarat.
- The Bengal tiger, which was documented to have preyed on the crocodile in Ranthambore National Park in Rajasthan. However, after a tiger was killed by a crocodile in 2016, Bivash Pandav of the Wildlife Institute of India's Endangered Species Department said "this is the first time we have heard of an attack by crocodiles on a tiger," adding that it was common for the opposite to happen.
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