Temporal range: middle Ordovician
However, a more recent study has argued that the shell of Bactroceras has important differences from those of true bactritids. For instance, the first shell chamber of Bactroceras resembles that of other Ordovician orthocerids, such as Archigeisonoceras and Hedstroemoceras: it is about 10 mm in diameter and is short, forming a spherical cap. True bactritids more strongly resemble late Silurian and Devonian orthocerids, whose first chamber is only about 5 mm across. Moreover, there is a large stratigraphic gap of nearly 50 million years between Bactroceras and the next orthocones with a spherical apex and ventral siphuncle.
Bactroceras had a thin, cone shaped shell. Its siphuncle, the tube connecting its chambers to the animal's body, was ventral, which means that it ran next to the shell wall on the underside of the shell. The first chamber of the shell was spherical. The soft anatomy of the animal was not known, but it lacked mineral deposits in its shell, it may have floated with the point of its shell upward, but somewhat inclined. Bactroceras differs from the later cephalopod Bactrites in being more circular in cross section and having deeper V-shaped ventral lobes.
Fossils were found in Middle Ordovician strata dating from the Whiterockian/Llanvirnian age. Locations were varied and stretched from New South Wales, Australia to El Puente, Bolivia to Antelope Valley, Nevada.
- Nautiloidea – Ellesmerocerida by W.M Furnish and Brian F. Glensister; Treatise on Invertebrate Paleontology Part K R.C. Moore (ed) Univ.Kans. press
- D. H. Evans. 2005. The Lower and Middle Ordovician cephalopod faunas of England and Wales. Monograph of the Palaeontographical Society 623:1-81
- Holland, Charles H. (2003). "Some observations on bactritid cephalopods". Bulletin of Geosciences. Czech Geological Survey. 78 (4): 369–372. ISSN 1214-1119.
- Kröger, Björn & Mapes, Royal H. (2007). "On the origin of bactritoids (Cephalopoda)". Paläontologische Zeitschrift. 81 (3): 316–327. doi:10.1007/BF02990181.