Frank Ruddle

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Frank Ruddle
Francis Hugh Ruddle

(1929-08-09)August 9, 1929
DiedMarch 10, 2013(2013-03-10) (aged 83)
Alma materUniversity of California, Berkeley (PhD)
AwardsDickson Prize in Medicine (1982)
William Allan Award (1984)[2]
Scientific career
InstitutionsYale University
ThesisChromosome variation in cell populations (1960)
Doctoral studentsAnne Ferguson-Smith[3]

Francis Hugh Ruddle (1929–2013) was an American cell and developmental biologist and Sterling Professor at Yale University.[4] Ruddle was an early pioneer of the Human Genome Project and created the first genetically modified mouse.[4]


  1. ^ In memoriam: Francis (Frank) Ruddle
  2. ^ Ruddle, Frank H. (1984). "The William Allan Memorial Award address: Reverse genetics and beyond". American Journal of Human Genetics. 36 (5): 944–953. ISSN 0002-9297. PMC 1684509. PMID 6594045.
  3. ^ Ferguson-Smith, Anne Carla (1989). A genomic analysis of the human homeobox gene loci HOX 1 and HOX 2 (PhD thesis). hdl:10079/bibid/9839056. OCLC 702620572.
  4. ^ a b Kucherlapati, Raju; Leinwand, Leslie A. (2013). "Frank Ruddle (1929–2013)". American Journal of Human Genetics. 92 (6): 839–840. doi:10.1016/j.ajhg.2013.05.012. PMC 3675234. PMID 24242788.