USS S-29 was transferred to the Royal Navy at New London on 5 June 1942. She arrived at Gibraltar, via Bermuda, in August 1942 to refit, nominally as part of the 8th Flotilla. She moved to Plymouth in February 1943, for an anti-submarine training role. In November 1943, she moved to Portsmouth for service in the same role.
HMS P556 was damaged by a battery explosion on 27 January 1944, and was decommissioned into the reserve in April. She was returned to the United States Navy on 26 January 1945. She was sold for scrap to Messrs Pounds, Portsmouth on 24 January 1947.