Harma theobene

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Harma theobene
Naturkundliche Sammlung Übermaxx Überseemuseum Bremen 0044.JPG
Specimen from Cameroon
Angular glider (Cymothoe theobene) female.jpg
Female, Kibale National Park, Uganda
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Nymphalidae
Subfamily: Limenitidinae
Genus: Harma
Doubleday, [1848]
Species: H. theobene
Binomial name
Harma theobene
Doubleday, [1848][1]


  • Amphidema Felder, 1861


  • Cymothoe theobene
  • Cymothoe theobene f. umbrina Joicey and Talbot, 1921
  • Cymothoe blassi Weymer, 1892
  • Cymothoe theobene nebetheo Suffert, 1904
  • Cymothoe theobene superna Fox, 1968
  • Cymothoe theobene ab. sordida Schultze, 1916
  • Cymothoe theobene ab. dualana Strand, 1914
  • Cymothoe theobene f. lutescens Poulton, 1922
  • Cymothoe theobene f. nigrolutescens Poulton, 1922
  • Cymothoe theobene f. nigrescens Poulton, 1922
  • Cymothoe (Harma) theobene f. jacksoni van Someren, 1939

Harma is a monotypic butterfly genus in the family Nymphalidae. Its one species is Harma theobene, the angular glider. It is found in Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, from Cameroon to Angola and in the Democratic Republic of the Congo, Uganda, Kenya, Tanzania, Malawi, Zambia and Mozambique.[2] The habitat consists of forests and heavy woodland.

The larvae possibly feed on Caloncoba gilgiana, Buchnerodendron, Rinorea, Lindackeria (including L. schweinfurthii) and Dovyalis species.


  • Harma theobene theobene – Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria
  • Harma theobene blassi (Weymer, 1892) – coast of Kenya, eastern Tanzania
  • Harma theobene superna (Fox, 1968) – Nigeria: Cross River loop, Cameroon, equatorial Africa, Angola, Democratic Republic of the Congo, Uganda, western Kenya, western Tanzania, Malawi, north-eastern Zambia
Harma theobene superna from Gabela forest, Angola


  1. ^ "Harma Doubleday, [1848]" at Markku Savela's Lepidoptera and Some Other Life Forms
  2. ^ Afrotropical Butterflies: Nymphalidae - Tribe Limenitidini