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Undetermined Harpalinae species
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: Adephaga
Family: Carabidae
Subfamily: Harpalinae
Bonelli, 1810

Harpalinae is a huge subfamily of ground beetles that contains 20,000 species[citation needed]. A rarely used common name for the subfamily is the harp beetles[citation needed]. The Harpalinae contain the most apomorphic ground beetles, displaying a wide range of forms and behaviors. Some are, rare among ground beetles, omnivores or even herbivores.

Many closely related subfamilies have been treated as subordinate taxa of the Harpalinae by various authors. Among these are the Dryptinae, Lebiinae (including Cyclosominae, Mormolycinae, Odacanthinae, Perigoninae), Licininae (including Chlaeniinae, Oodinae), Orthogoniinae, Panagaeinae, Platyninae, Pseudomorphinae, Pterostichinae (including Zabrinae). Here, they are considered independent families within the harpaline (sensu lato) assemblage, and this is also tentatively assumed for the enigmatic monotypic genus Ginema.


At least 4 large and several smaller supertribes can be distinguished. The former "Pterostichitae" on the other hand seem to be, as presently delimited, a paraphyletic assemblage, but probably also contains a number of tribes closer to each other than to other Harpalinae. The clade around the type genus Pterostichus is thus recognized as subfamily Pterostichinae here.[1]


The Harpalinae include the following genera:[2][verification needed]


  1. ^ CWD (2008)
  2. ^ "Harpalinae Bonelli, 1810". Carabidae of the World. 2012. Retrieved 11 Nov 2012.