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Temporal range: Carboniferous
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Coleoidea
(unranked): Belemnoidea
Order: Hematitida
Doguzhaeva et al., 2002[1]
Family and Genera


  • Hematites
  • Bactritimimus
  • Paleoconus

Hematitida is a group of coleoid cephalopods known from the early Carboniferous Period. They are the oldest definite coleoids,[2] although there are controversial claims for even older coleoids from the Devonian.[3] Fossil hematitidans have so far been found only in Arkansas and Utah of the United States.[2] The only family described so far is Hematitidae.


Some features shared by hematitidans include


The classification for this group comes from Doguzhaeva et al. 2003[4]

Order Hematitida Doguzhaeva, Mapes, & Mutvei, 2002

  • Family Hematitidae Gustomesov 1976
    • Hematites Flower & Gordon 1959 - type genus
    • Bactritimimus Flower & Gordon, 1959
    • Paleoconus Flower & Gordon, 1959


  1. ^ a b Doguzhaeva, Larisa A.; Mapes, Royal H.; Mutvei, Harry (2002). "Shell morphology and ultrastructure of the Early Carboniferous coleoid Hematites Flower & Gordon, 1959 (Hematitida ord. nov.) from Midcontinent (USA)". Abhandlungen der Geologischen Bundesanstalt. 57: 299–320'. ISBN 3-85316-014-X. ISSN 0016-7800. 
  2. ^ a b Nishiguchi, Michelle; Mapes, Royal K. (2008), "Cephalopoda", in Ponder, Winston F., & Lindberg, David R., Phylogeny and evolution of the Mollusca, Dordrecht, The Netherlands: Springer, pp. 163–199, ISBN 978-0-520-25092-5 
  3. ^ Bandel, Klaus; Reitner, Joachim; Sturmer, Wilhelm (1983). "Coleoidea from the Devonian Black Slate ("Hunsruck-Schiefer")" (PDF). Neues Jahrbuch fur Geologie und Paläontology, Abhandlungen. Stuttgart. 165 (3): 397–417. 
  4. ^ Doguzhaeva, Larisa A.; Mapes, Royal H.; Mutvei, Harry (2003). "The shell and ink sac morphology and ultrastructure of the Late Pennsylvanian cephalopod Donovaniconus and its phylogenetic significance". Berliner Paläobiologische Abhandlungen. 3: 61–78.