Holdfasts vary in shape and form depending on both the species and the substrate type. The holdfasts of organisms that live in muddy substrates often have complex tangles of root-like growths. Those of organisms that live in sandy substrates are bulb-like and very flexible, such as the holdfast of sea pens, thus permitting the organism to pull the entire body into the substrate when the holdfast is contracted. The holdfasts of organisms that live on smooth surfaces (such as the surface of a boulder) have the flattened base of the holdfast literally glued to the surface. The organism derives no nutrition from this intimate contact with the substrate, as the process of liberating nutrients from the substrate requires enzymatically eroding the substrate away, thereby increasing the risk of organism falling off the substrate.
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