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In botany, a seed is described as showing hypogeal germination when the cotyledons of the germinating seed remain non-photosynthetic, inside the seed shell, and below ground. The converse, where the cotyledons expand, throw off the seed shell and become photosynthetic above the ground, is epigeal germination.
In water purification works, the hypogeal (or Schmutzdecke) layer is a biological film just below the surface of slow sand filters. It contains microorganisms that remove bacteria and trap contaminant particles.
- Adrian D. Bell; Alan Bryan (2008). Plant Form: An Illustrated Guide to Flowering Plant Morphology. Timber Press. p. 200. ISBN 978-0-88192-850-1.