|Reconstructed Paraceratherium skull.|
Indricotheriinae is a group of long-limbed, hornless rhinoceroses that evolved in the Eocene epoch and continued through to the early Miocene. Early paraceratheres were only about the size of dogs, but by the late Eocene and early Oligocene, they grew to much bigger sizes. They flourished in the rainforests of a coastal region that became Kazakhstan, India, and southwest China, and lived further inland throughout central Asia as well.
The paraceratheres reached the peak of their evolution from the middle Oligocene through to the early Miocene, where they became very large herbivorous mammals. Most genera were about the size of modern draft horses, which some growing significantly larger. The largest genus was Paraceratherium, which was more than twice as heavy as an African elephant was one of the largest land mammals that ever lived. However, they remained confined to central Asia, which at the time was part of a large, lush lowland region. The collision with the Indian subcontinent and the Himalayan uplift led to global cooling, desertification, and the disappearance of forest habitats, which resulted in the extinction of these giant ungulates.
Although considered a subfamily of Hyracodontidae by some authors, recent authors treat paraceratheres as a distinct family, Paraceratheriidae (Wang et al. 2016 recover hyracodonts as more basal than paraceratheres).
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