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|Full name||Jonathan Santana Gehre|
|Date of birth||19 October 1981|
|Place of birth||Buenos Aires, Argentina|
|Height||1.81 m (5 ft 11 in)|
|2009||→ San Lorenzo (loan)||10||(2)|
|2013||Belgrano de Córdoba||2||(0)|
|* Senior club appearances and goals counted for the domestic league only and correct as of 16 March 2017.
‡ National team caps and goals correct as of 27 December 2015
Santana was a key part of Paraguay's national team whilst under management of Tata Martino, featuring in three major competitions since his called up to the squad in 2007, the 2007 Copa América, the 2010 FIFA World Cup and the 2011 Copa América. Santana missed the final of the latter due to a red card in the semi-final against Venezuela.
Santana started his career at San Telmo of the Argentine 3rd Division. He went on to play for a number of Argentine Primera clubs, including San Lorenzo and River Plate. In August 2006 he signed a five-year contract with German VfL Wolfsburg. In the winter of 2009, he was loaned out to San Lorenzo.
On 26 July 2010, the Turkish club Kayserispor announced the hiring of the Paraguayan midfielder for three seasons.
In June 2007, Santana received Paraguayan citizenship (his mother is a Paraguayan citizen born in San Ignacio, Misiones Department and Argentina did not recognize dual citizenship). He played for Paraguay in the 2007 Copa América. He was also named in Paraguay's final 23-men squad for the 2010 FIFA World Cup. He entered the field during Paraguay's first game, a 1–1 draw with Italy, but suffered an injury during the match. However, as Paraguay's coach Gerardo Martino had already made the three available substitutions, Santana had to stay on the field and play injured for the remaining time.
- San Lorenzo
- VfL Wolfsburg
- Refuerzo de experiencia: En Sarmiento firmó Jonathan Santana, laverdadonline.com, 5 August 2016
- "12 de Octubre vs. Cerro Porteño 1 - 3". Soccerway. 16 February 2014. Retrieved 20 February 2014.
- "Jonathan Santana se nacionalizó paraguayo". adnmundo.com (in Spanish). 4 June 2007. Retrieved 2 May 2010.
- "Este Santana suena mal" (in Spanish). Olé. 16 June 2010. Retrieved 16 June 2010.