|In Joshua Tree National Park, California|
Yucca brevifolia is a plant species belonging to the genus Yucca. It is tree-like in habit, which is reflected in its common names: Joshua tree, yucca palm, tree yucca, and palm tree yucca.
This monocotyledonous tree is native to the arid southwestern United States, specifically California, Arizona, Utah, and Nevada, where it is confined mostly to the Mojave Desert between 400 and 1,800 m (1,300 and 5,900 ft) elevation. It thrives in the open grasslands of Queen Valley and Lost Horse Valley in Joshua Tree National Park. A dense Joshua tree forest also exists on the Cima Dome (Cima, California) in the Mojave National Preserve; northeast of Kingman, Arizona, in Mohave County, Arizona; as well as along U.S. 93 between the towns of Wickenburg and Wikieup, and designated as the Joshua Tree Parkway of Arizona.
The Joshua tree is also called izote de desierto (Spanish, "desert dagger"). It was first formally described in the botanical literature as Yucca brevifolia by George Engelmann in 1871 as part of the Geological Exploration of the 100th meridian (or "Wheeler Survey").
The name Joshua tree is commonly said to have been given by a group of Mormon settlers crossing the Mojave Desert in the mid-19th century: The tree's role in guiding them through the desert combined with its unique shape reminded them of a biblical story in which Joshua keeps his hands reached out for an extended period of time in order to guide the Israelites in their conquest of Canaan (Joshua 8:18–26). There is, however, no direct or contemporary attestation of this origin, and the name Joshua tree is not recorded until after Mormon contact; moreover, the physical appearance of the Joshua tree more closely resembles a similar story told of Moses.
In addition to the autonymic subspecies Yucca brevifolia subsp. brevifolia, two other subspecies have been described: Yucca brevifolia subsp. jaegeriana (the Jaeger Joshua tree or Jaeger's Joshua tree or pygmae yucca) and Yucca brevifolia subsp. herbertii (Webber's yucca or Herbert Joshua tree), though both are sometimes treated as varieties or forms.
Growth and development
Joshua trees are fast growers for the desert; new seedlings may grow at an average rate of 7.6 cm (3.0 in) per year in their first ten years, then only about 3.8 cm (1.5 in) per year. The trunk consists of thousands of small fibers and lacks annual growth rings, making it difficult to determine the tree's age. This tree has a top-heavy branch system, but also what has been described as a "deep and extensive" root system, with roots reaching up to 11 m (36 ft). If it survives the rigors of the desert, it can live for hundreds of years; some specimens survive a thousand years. The tallest trees reach about 15 m (49 ft). New plants can grow from seed, but in some populations, new stems grow from underground rhizomes that spread out around the parent tree.
The evergreen leaves are dark green, linear, bayonet-shaped, 15–35 cm long and 7–15 mm broad at the base, tapering to a sharp point; they are borne in a dense spiral arrangement at the apex of the stems. The leaf margins are white and serrate.
Flowers typically appear from February to late April, in panicles 30–55 cm tall and 30–38 cm broad, the individual flowers erect, 4–7 cm tall, with six creamy white to green tepals. The tepals are lanceolate and are fused to the middle. The fused pistils are 3 cm tall and the stigma cavity is surrounded by lobes. The semi-fleshy fruit that is produced is green-brown, elliptical, and contains many flat seeds. Joshua trees usually do not branch until after they bloom (though branching may also occur if the growing tip is destroyed by the yucca-boring weevil), and they do not bloom every year. Like most desert plants, their blooming depends on rainfall at the proper time. They also need a winter freeze before they bloom.
Once they bloom, the trees are pollinated by the yucca moth, which spreads pollen while laying her eggs inside the flower. The moth larvae feed on the seeds, but enough seeds remain to reproduce. The Joshua tree is also able to actively abort ovaries in which too many eggs have been laid.
Distribution and habitat
Yucca brevifolia is endemic to the Southwestern United States with populations in western Arizona, southeastern California, southern Nevada, and southwestern Utah. This range mostly coincides with the geographical reach of the Mojave Desert, where it is considered one of the major indicator species for the desert. It occurs at altitudes between 400 and 1,800 m (1,300 and 5,900 ft).
There is a large area of Joshua trees in the desert between Zacatecas and Saltillo, Mexico.
In a 2001 paper published in the journal Ecosystems, Joshua trees are one of the species predicted to have their range reduced and shifted by climate change. There is concern that they will be eliminated from Joshua Tree National Park, with ecological research suggesting a high probability that their populations will be reduced by 90% of their current range by the end of the 21st century, thus fundamentally transforming the ecosystem of the park. There is also concern about the ability of the species to migrate to favorable climates due to the extinction of the giant Shasta ground sloth (Nothrotheriops shastensis) 13,000 years ago; ground sloth dung has been found to contain Joshua tree leaves, fruits, and seeds, suggesting that the sloths might have been key to the tree's dispersal.
Uses and cultivation
Different forms of the species are cultivated, including smaller plants native from the eastern part of the species range. These smaller plants grow 2.5 meters tall and branch when about a meter tall.
Cahuilla Native Americans, who have lived in the southwestern United States for generations, identify this plant as a valuable resource and call it "hunuvat chiy’a" or "humwichawa". Their ancestors used the leaves of Y. brevifolia to weave sandals and baskets, in addition to harvesting the seeds and flower buds for meals.
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An application to Mr. Frederick V. Coville, botanist of the Department of Agriculture, elicited the following response: 'The statement is often made that this name Joshua-tree was applied to Cleistoyucca brevifolia because it was this tree which led the Mormons through the desert. I have no means of knowing, however, whether this explanation is authentic or whether it was invented as an explanation of the name. It seems to me more likely that Joshua-tree is a garbled Indian name' […] I asked Professor Marcus E. Jones, whose knowledge of the desert flora is unsurpassed, and who has had a long acquaintance with members of the Mormon church. In reply he kindly wrote as follows: 'The Mormon church officials do not know exactly the origin of the term, but assume that it is from the wide-spreading arms (branches) that in the night remind of the time when in battle Joshua had his arms held up in order to win a battle. This I got from one of the twelve apostles of the Mormon church.' Plausible as this explanation is, its value is more or less shaken when one finds, as I did after looking up the family Bible, that it was Moses, not Joshua, who had his arms held up during the battle, while Joshua conducted the fighting (Ex. 17:8–13). There is, however, another account of a fight, which may be what the Mormon apostle had in mind. It is told in the book of Joshua 7:18–26 [sic].
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- Cole, Kenneth L.; Ironside, Kirsten; Eischeid, Jon; Garfin, Gregg; Duffy, Phillip B.; Toney, Chris (2011). "Past and ongoing shifts in Joshua tree distribution support future modeled range contraction" (PDF). Ecological Applications. 21 (1): 137–149. doi:10.1890/09-1800.1. PMID 21516893.
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