|In Joshua Tree National Park, California|
|Natural range in the United States|
Yucca brevifolia is a plant species belonging to the genus Yucca. It is tree-like in habit, which is reflected in its common names: Joshua tree, yucca palm, tree yucca, and palm tree yucca.
This monocotyledonous tree is native to the arid Southwestern United States, specifically California, Arizona, Utah, and Nevada, and to northwestern Mexico (Baja California, Baja California Sur, Sinaloa, and Sonora). It is confined mostly to the Mojave Desert between 400 and 1,800 m (1,300 and 5,900 ft) elevation. It thrives in the open grasslands of Queen Valley and Lost Horse Valley in Joshua Tree National Park. A dense Joshua tree forest also existed on the Cima Dome (Cima, California) in the Mojave National Preserve until the Dome fire of August 2020; northeast of Kingman, Arizona, in Mohave County; and along U.S. 93 between the towns of Wickenburg and Wikieup, and designated as the "Joshua Tree Parkway of Arizona". The Joshua Tree has barely any leaves, just a few balls on the edges.
The Joshua tree is also called izote de desierto (Spanish, "desert dagger"). It was first formally described in the botanical literature as Yucca brevifolia by George Engelmann in 1871 as part of the Geological Exploration of the 100th meridian (or "Wheeler Survey").
The name "Joshua tree" is commonly said to have been given by a group of Mormon settlers crossing the Mojave Desert in the mid-19th century: The tree's role in guiding them through the desert combined with its unique shape reminded them of a biblical story in which Joshua keeps his hands reached out for an extended period of time to guide the Israelites in their conquest of Canaan (Joshua 8:18–26). Further, the shaggy leaves may have provided the appearance of a beard. However, no direct or contemporaneous attestation of this origin exists, and the name Joshua tree is not recorded until after Mormon contact; moreover, the physical appearance of the Joshua tree more closely resembles a similar story told of Moses.
Ranchers and miners who were contemporary with the Mormon immigrants used the trunks and branches as fencing and for fuel for ore-processing steam engines. They referred to these fallen or collapsed Joshua trees as tevis stumps.[failed verification]
In addition to the autonymic subspecies Y. b. subsp. brevifolia, two other subspecies have been described: Y. b. subsp. jaegeriana (the Jaeger Joshua tree or Jaeger's Joshua tree or pygmae yucca) and Y. b. subsp. herbertii (Webber's yucca or Herbert Joshua tree), though both are sometimes treated as varieties or forms.
Growth and development
Joshua trees are fast growers for the desert; new seedlings may grow at an average rate of 7.6 cm (3.0 in) per year in their first 10 years, then only about 3.8 cm (1.5 in) per year. The trunk consists of thousands of small fibers and lacks annual growth rings, making determining the tree's age difficult. This tree has a top-heavy branch system, but also what has been described as a "deep and extensive" root system, with roots reaching down to 11 m (36 ft). If it survives the rigors of the desert, it can live for hundreds of years; some specimens survive a thousand years. The tallest trees reach about 15 m (49 ft). New plants can grow from seed, but in some populations, new stems grow from underground rhizomes that spread out around the parent tree.
The evergreen leaves are dark green, linear, bayonet-shaped, 15–35 cm long, and 7–15 mm broad at the base, tapering to a sharp point; they are borne in a dense spiral arrangement at the apex of the stems. The leaf margins are white and serrated.
Flowers typically appear from February to late April, in panicles 30–55 cm tall and 30–38 cm broad, the individual flowers erect, 4–7 cm tall, with six creamy white to green tepals. The tepals are lanceolate and are fused to the middle. The fused pistils are 3 cm tall and the stigma cavity is surrounded by lobes. The semifleshy fruit that is produced is green-brown, elliptical, and contains many flat seeds. Joshua trees usually do not branch until after they bloom (though branching may also occur if the growing tip is destroyed by the yucca-boring weevil), and they do not bloom every year. Like most desert plants, their blooming depends on rainfall at the proper time. They also need a winter freeze before they bloom.
Once they bloom, the flowers are pollinated by the yucca moth (Tegeticula synthetica), which spreads pollen while laying eggs inside the flower. The larvae feed on the seeds, but enough seeds remain to reproduce. The Joshua tree is also able to actively abort ovaries in which too many eggs have been produced.
Distribution and habitat
Y. brevifolia is native to the Southwestern United States with populations in western Arizona, southeastern California, southern Nevada, and southwestern Utah, and to northwestern Mexico. This range mostly coincides with the geographical reach of the Mojave Desert, where it is considered one of the major indicator species for the desert. It occurs at elevations between 400 and 1,800 m (1,300 and 5,900 ft).
Joshua trees are one of the species predicted to have their range reduced and shifted by climate change. Concern remains that they will be eliminated from Joshua Tree National Park, with ecological research suggesting a high probability that their populations will be reduced by 90% of their current range by the end of the 21st century, thus fundamentally transforming the ecosystem of the park. Also, concern exists about the ability of the species to migrate to favorable climates due to the extinction of the giant Shasta ground sloth (Nothrotheriops shastensis) 13,000 years ago; ground sloth dung has been found to contain Joshua tree leaves, fruits, and seeds, suggesting that the sloths might have been key to the trees' dispersal.
Uses and cultivation
Different forms of the species are cultivated, including smaller plants native from the eastern part of the species range. These smaller plants grow 2.5 m tall and branch when about 1 m tall. Red-shafted flickers make nests in the branches, which are later used by other birds.
Cahuilla Native Americans, who have lived in the Southwestern United States for generations, identify this plant as a valuable resource and call it hunuvat chiy’a or humwichawa. Their ancestors used the leaves of Y. brevifolia to weave sandals and baskets, in addition to harvesting the seeds and flower buds for meals. Native Americans also used the reddish roots to make dye. Yucca tree roots have saponin glycosides.
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An application to Mr. Frederick V. Coville, botanist of the Department of Agriculture, elicited the following response: 'The statement is often made that this name Joshua-tree was applied to Cleistoyucca brevifolia because it was this tree which led the Mormons through the desert. I have no means of knowing, however, whether this explanation is authentic or whether it was invented as an explanation of the name. It seems to me more likely that Joshua tree is a garbled Indian name' […] I asked Professor Marcus E. Jones, whose knowledge of the desert flora is unsurpassed, and who has had a long acquaintance with members of the Mormon church. In reply, he kindly wrote as follows: 'The Mormon church officials do not know exactly the origin of the term, but assume that it is from the wide-spreading arms (branches) that in the night remind of the time when in battle Joshua had his arms held up in order to win a battle. This I got from one of the twelve apostles of the Mormon church.' Plausible as this explanation is, its value is more or less shaken when one finds, as I did after looking up the family Bible, that it was Moses, not Joshua, who had his arms held up during the battle, while Joshua conducted the fighting (Ex. 17:8–13). There is, however, another account of a fight, which may be what the Mormon apostle had in mind. It is told in the book of Joshua 7:18–26 [sic].
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- Cole, Kenneth L.; Ironside, Kirsten; Eischeid, Jon; Garfin, Gregg; Duffy, Phillip B.; Toney, Chris (2011). "Past and ongoing shifts in Joshua tree distribution support future modeled range contraction" (PDF). Ecological Applications. 21 (1): 137–149. doi:10.1890/09-1800.1. PMID 21516893.
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