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Temporal range: Devonian
Ahlberg et al., 2000
Four legs developed in water, not on land, to better escape waterliving predatory creatures like Hyneria. There were very lush forests, and particularly swamps, where four limbs became very useful to avoid predators.
- Ahlberg, P.E., E. Luksevics and E. Mark-Kurik. 2000. "A near-tetrapod from the Baltic Middle Devonian". Paleontology 43(3): 533-548.
- Livoniana multidentata at Devonian Times
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