|L. p. phlaeus|
Lycaena phlaeas, the small copper, American copper, or common copper, is a butterfly of the Lycaenids or gossamer-winged butterfly family. According to Guppy and Shepard (2001), its specific name phlaeas is said to be derived either from the Greek φλέγω (phlégo), "to burn up", or from the Latin floreo, "to flourish".
The upperside forewings are a bright orange with a dark outside edge border and with eight or nine black spots. The hindwings are dark with an orange border. Some females also have a row of blue spots inside the orange border and are known as form caeruleopunctata. The undersides are patterned in a similar way but are paler. The black spots on the forewings are outlined in yellow and the dark colouring is replaced by a pale brownish grey. The hindwings are the same brown/grey colour with small black dots and a narrow orange border. The caterpillars (larvae) are usually green, but some have a purple stripe down the middle of the back and along each side.
L. p. eleus
male L. p. eleus ab. caeruleopunctata
female L. p. eleus
L. p. phlaeas male
In bright sun it is a very active little butterfly with the males setting up small territories which they will defend vigorously against rival males or indeed any unlucky passing insect. Even the shadow of a large bird passing overhead is enough to elicit a response. Females are pursued and mating usually occurs in vegetation.
The eggs are laid singly and conspicuously on the upperside of food plant leaves and the young caterpillar feeds on the underside of the leaf creating "windows" by leaving the upper epidermis of the leaf untouched. Pupation takes place in the leaf litter and the pupa is thought to be tended by ants. There are between two and three broods a year, fewer further north. In exceptionally good years, a fourth brood sometimes occurs in the south and adults can still be seen flying into November. The species overwinters as a caterpillar.
Lycaena phlaeas belongs to the subgenus Lycaena and is the type-species of genus Lycaena. The species is in turn divided into several subspecies, although many probably lack reason to be named, and others may merit an elevation of rank to be considered autonomous species:
- L. p. hibernica Goodson, 1948 — Republic of Ireland, Northern Ireland
- L. p. eleus Fabricius, 1798 — England, Wales, Scotland
- L. p. phlaeas Linnaeus, 1761 — Europe, western Siberia, Caucasus, South Caucasus (type locality = Sweden)
- L. p. polaris Courvoisier, 1911 — northern Ural Mountains, northern Siberia, Chukotka Peninsula (Russia)
- L. p. kamtschatica Gorbunov, 1994 — Kamchatka Peninsula (Russia)
- L. p. ganalica Gorbunov, 1995 — Kamchatka Peninsula (Russia)
- L. p. daimio (Seitz, ) — south Kurile Islands, Sakhalin (Russian islands)
- L. p. gonpaensis (Yoshino, ) — North Yunnan, China
- L. p. hypophlaeas (Boisduval, 1852) — N California Sierra Nevada Mts. (Type location) and White Mts., Altai Mountains, southern Siberia, Amur (Russia–NW China border region), northern Ussuri (adjacent to Amur)
- L. p. chinensis (Felder, 1862) — southern Ussuri
- L. p. oxiana (Grum-Grshimailo, 1890) — Kopet Dag mountains, Alai Mountains, Ghissar-Darvaz, Tian Shan mountains
- L. p. comedarum (Grum-Grshimailo, 1890) — eastern Pamir Mountains
- L. p. stygiana Butler, 1880 — western Pamir Mountains
- L. p. shima Gabriel, 1954 — Arabia
- Lycanea phlaeas pseudophlaeas (Lucas, 1866) — Ethiopia, Uganda
- L. p. ethiopica (Poulton, 1922) — Ruwenzori Mountains (SW Uganda)
- L. p. abbottii (Holland, 1892) — Kenya, Tanzania, Malawi
- L. p. flavens (Ford, 1924) — Tibet
- L. p. coccinea (Ford, 1924)
- L. p. americana (Morris, 1862) — Nova Scotia and W. to Minnesota, and S. to: Virginia, Montane N Georgia, Missouri, Kansas, N Dakota.
- L. p. arethusa (Dod, 1907) — Alberta
- L. p. arctodon Ferris, 1974 — Montana
- L. p. feildeni (McLachlan, 1878) — Ellesmere
- L. p. alpestris Emmel & Pratt, 1998 — California
Subspecies daimio, Osaka, Japan
In Tokyo, Japan
- List of butterflies of India
- List of butterflies of India (Lycaenidae)
- List of butterflies of Great Britain
- "NatureServe Explorer 2.0 Lycaena phlaeas Little Copper". explorer.natureserve.org. Retrieved 3 October 2020.
- Boisduval. Annales de la Societe Entomologique de France. 1852:2(10),291, no. 23
- FERRIS, C. D. 1974. Distribution of arctic-alpine Lycaena phlaeas L. (Lycaenidae) in North America with designation of a new subspecies. Bull. Allyn Mus. 18:1-13
- Ballmer, G.R., Pratt, G.F., Journal of the Lepidopterists' Society 43(1), 1989, 59- 65
- Morris, John G (1862). Synopsis of the described Lepidoptera of North America Part 1. - Diurnal and Crepuscular Lepidoptera. Compiled for the Smithsonian Institution by John G. Morris, Smithsonian Miscellaneous Collections, Washington D.C., 358 pp. 1862.
- Asher, Jim, Martin Warren, Richard Fox, Paul Harding, Gaile Jeffcoate & Stephen Jeffcoate (Eds) (2001) The Millennium Atlas of Butterflies in Britain and Ireland. Oxford University Press.
- Darby, Gene (1958). What is a Butterfly. Chicago: Benefic Press. p. 34.
- Dempster, J.P. & A.M. Emmet (1990) Lycaena phlaeas (Linnaeus). Pp. 134–139 in A. Maitland Emmet, John Heath et al. The Butterflies of Great Britain and Ireland. The Moths and Butterflies of Great Britain and Ireland vol. 7, part 1. Harley Books, Colchester, UK.
- Evans, W.H. (1932). The Identification of Indian Butterflies (2nd ed.). Mumbai, India: Bombay Natural History Society.
- Gaonkar, Harish (1996). Butterflies of the Western Ghats, India (including Sri Lanka) - A Biodiversity Assessment of a Threatened Mountain System. Bangalore, India: Centre for Ecological Sciences.
- Gay, Thomas; Kehimkar, Isaac David; Punetha, Jagdish Chandra (1992). Common Butterflies of India. Nature Guides. Bombay, India: World Wide Fund for Nature-India by Oxford University Press. ISBN 978-0195631647.
- Guppy, C.S. and Shepard, J. (2001) Butterflies of British Columbia British Columbia Museum, Canada.
- Haribal, Meena (1992). The Butterflies of Sikkim Himalaya and Their Natural History. Gangtok, Sikkim, India: Sikkim Nature Conservation Foundation.
- Kunte, Krushnamegh (2000). Butterflies of Peninsular India. India, A Lifescape. Hyderabad, India: Universities Press. ISBN 978-8173713545.
- Tomlinson, David & Rob Still (2002) Britain's Butterflies. WildGuides, Old Basing, UK.
- Wynter-Blyth, Mark Alexander (1957). Butterflies of the Indian Region. Bombay, India: Bombay Natural History Society. ISBN 978-8170192329.
- Yoshino, K. (2019). "Description of a new subspecies of Lycaena phlaeas from northern Yunnan, China". Butterflies. Japan. 80: 16–20.
- Lycaena at Markku Savela's website on Lepidoptera