SP used these locomotives in the Sierra Nevada for about 20 years, retiring them in the mid-1930s. They were stored in the railroad's Sacramento, California, shops for a couple years before being rebuilt with 4B Worthington feedwater heaters and uniform cylinders ("simpling" them) measuring 22 in × 28 in (559 mm × 711 mm), diameter × stroke. The rebuilds increased the class weight to 424,200 lb (192,400 kg) with 356,900 lb (161,900 kg) on the drivers, 210 psi (1.4 MPa) boiler pressure and 76,800 lbf (342 kN) tractive effort.
The rebuilt locomotives were renumbered into the 3900 series then used on SP's Portland Division in Oregon until they were again retired in the late 1940s. The locomotives were all scrapped soon after retirement with the last, 3907 (originally 4207), on September 23, 1948.